Dienerian (early triassic) ammonoids from the northern indian margin 1st edition david ware hugo bu

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Dienerian (Early Triassic) ammonoids from the Northern Indian Margin 1st

Edition David Ware Hugo Bucher

Thomas Brühwiler Elke

Schneebeli■Hermann Peter A. Hochuli†

Leopold Krystyn Ghazala Roohi Khalil

Ur■Rehman Amir Yaseen

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Dienerian (Early Triassic) ammonoids from the Northern Indian Margin

David Ware, Hugo Bucher, Thomas Brühwiler, Elke Schneebeli-Hermann, Peter A. Hochuli†, Leopold Krystyn, Ghazala Roohi, Khalil Ur-Rehman and Amir Yaseen

Acknowledgement

Financial support for the publication of this issue of Fossils and Strata was provided by the Lethaia Foundation

Foreword

Dienerian(EarlyTriassic)ammonoidsandtheEarlyTriassic bioticrecovery:areview

DAVIDWAREANDHUGOBUCHER

Ithasbeenestimatedthatabout90%ofallmarine speciesdisappearedduringtheend-Permianmass extinction(Raup&Sepkoski1982).Itisthebiggest knownbiodiversitycrisisinthehistoryofPhanerozoiclife,anditledtothereplacementoftypical Palaeozoicfaunasbytypicalmoderncommunities (Sepkoski1984).Therecoverywhichfollowedinthe EarlyTriassicisanintensivelystudiedtopic.This recoveryistraditionallyconsideredasdelayedin comparisonwithothermassextinctions(Erwin 1998,2006)asseveralmajormarinecladessuchas corals(Stanley2003),foraminifers(Tong&Shi 2000)orradiolarians(Racki1999)recoveredonlyin thelateSpathian(EarlyTriassic)orintheAnisian (MiddleTriassic),ca.5MyafterthePermian–Triassicboundary.Thisdelayisinterpretedastheconsequenceofpersistinganoxicconditions(Wignall& Twitchett2002)andunstableenvironmentalconditionsduringtheentireEarlyTriassic(Payne etal. 2004).However,severalrecentstudiessuggesta morecomplexscenario,withpulsesofrecovery interruptedbyperiodsofadditionalextinctions.For example,conodonts(Orchard2007;Goudemand etal. 2008)firstunderwentanimportantturnover attheGriesbachian–Dienerianboundary,followed byanexplosiveradiationintheearly–middle Smithian,adramaticextinctioninthelateSmithian, andanotherradiationduringtheearlySpathian. Ammonoidsalsorecoveredveryfastcomparedto othergroups,reachingpre-extinctionlevelsofdiversityalreadyduringtheSmithian(Fig.1;Brayard etal. 2009).Hofmann etal. (2014)showedthat benthicecosystemsstartedtorecoveralreadyinthe

Griesbachian,butthisrecoveryhasbeeninterrupted byareturntoharshenvironmentalconditions(e.g. anoxia,warmtemperatures)duringtheDienerian. Recoveryofthebenthosresumedduringthe Smithian.Basedonpalynologicalandcarbonisotopesanalysis,Hermann etal. (2011a,b,2012a,b) andSchneebeli-Hermann etal. (2012,2015)contradictedtheideaofpersistentwidespreadanoxiaand showedthatthisanoxiawasrestrictedtothemiddle–lateDienerianandlateSmithian.LatePermian andEarlyTriassicecologicalcrisesofterrestrial plantsalsoimmediatelypredateextinctioncrisesof marineorganisms,andtheDieneriandiversitylowis noexceptionasdocumentedbyHochuli etal. (2016).

Manystudiesaddressingtherecoveryarebased oninsufficientlyresolvedagecontrols.Theconstructionofadetailedtime-scalefortheEarlyTriassicis thecornerstoneonwhichanystudyaddressingthis bioticrecoverymustbebased.Ovtcharova etal. (2006)andGalfetti etal. (2007)establishedadurationofca.4.5MyrfortheEarlyTriassicandshowed thatthefourEarlyTriassicageswereofveryuneven duration,theSpathianrepresentingmorethanhalf ofthisinterval(Fig.2).Galfetti etal. (2007) obtainedamaximaldurationofca.1.4 0.4Myr fortheGriesbachian–Dieneriantimeinterval.No durationoftheDienerianaloneisavailable,butit canbereasonablyassumedthatitis <1Myr.Anew generationofhigh-resolutionU-PbagesforthePermian–Triassicboundary(Burgess etal. 2014)and fortheEarly–MiddleTriassicboundary(Ovtcharova etal. 2015)indicateadurationof4.83 0.19Myr © 2018LethaiaFoundation.PublishedbyJohnWiley&SonsLtd

Fig.1. Totalgenericrichness(blackboldline,allammonoids;greylines,majorammonoidgroups)andmeanChao2estimateofthe overallgenericrichnesswithits95%confidenceinterval(largecircleswithverticalbars).TheEarlyTriassicishighlightedindarkgrey. PTB,Permian–Triassicboundary;1,Kasimovian;2,Gzhelian;3,Asselian;4,Sakmarian;5,Artinskian;6,Kungurian;7,Roadian;8,Wordian;9,Capitanian;10,Wuchiapingian;unlabelledsuccessiveintervals,Changhsingian,Griesbachian,Dienerian,Smithian;15,Spathian; 16,EarlyAnisian;17,MiddleAnisian;18,LateAnisian;19,Ladinian;20,EarlyCarnian;21,LateCarnian;22,EarlyNorian;23,Middle Norian;24,LateNorian;25,Rhaetian.ModifiedafterBrayard etal. (2009).

± 0.56

et al., 2006

et al., 2007

et al., 2007

Fig.2. LowerTriassicstageandsubstagesubdivision(Ogg2012) calibratedwithrecentlypublishedradiometricagesfromSouth China.

fortheEarlyTriassic.However,therespectivedurationofeachofthefourLowerTriassicsubstages maynotbesignificantlychangedbecausethesenew U-Pbagesareconsistentlyyoungerthanthoseofthe previousgeneration.

Brayard&Bucher(2008)proposedanewdetailed biostratigraphicalschemebasedonammonoidsfor theSmithianofSouthChina.Bruhwiler etal. (2010a) constructedthemosthighlyresolvedbiostratigraphicalschemefortheSmithianbasedonammonoids fromtheNorthernIndianMargin.Thisworkshowed thatammonoidsunderwentanexplosiveradiation intheearlySmithian,withconstanthighdiversity associatedwithextremelyhighturnoverrates throughoutthemiddleSmithianandamajorextinctioninthelateSmithian.Thisextinctioncouldnot bedetectedbyBrayard etal. (2009)duetothecoarsertime-scaleofthisstudy.Conodontsalsosuffered fromadrasticextinctioninthelateSmithian.

ThestagesubdivisionsoftheLowerTriassicarea subjectofdebate(Fig.2).Inthelatestversionof TheGeologicTimeScale(Ogg2012),thetwofold subdivisionoftheLowerTriassicintroducedby Kiparisova&Popov(1956),withtheInduanand Olenekian,isendorsed.Thefourstagesdefinedby Tozer(1965)arethenconsideredassubstages,the InduanbeingsubdividedintoGriesbachianand Dienerian,andtheOlenekianintoSmithianand Spathian.Thistwofoldschemeishoweverstrongly criticized(e.g.Shevyrev2006),mainlyasitdoesnot reflecttheendSmithiancrisis,themostimportant extinctioneventknownforbothammonoidsand conodontswithintheentireTriassicineverylocality wherethisquestionhasbeenaddressed(e.g Stanley

E.T. M.Triassic Late Triassic
L.Perm. M.Perm.

2009;forammonoids:Brayard etal. 2006;Bruhwiler etal. 2010a;Brayard&Bucher2015;forconodonts: Orchard2007;Chen etal. 2013;Komatsu etal. 2016).Moreover,havingtheInduandefinedinthe TethyanrealmandtheOlenekianintheBoreal Realmmakesthecorrelationofthisstageboundary acrosssuchabroadpalaeolatitudinalrangeanarduoustask.Tozer(1965)providedabroaddefinition oftheDienerian–Smithianboundary(corresponding totheInduan–Olenekianboundary),explainingthat itwasprobablethatinCanada,theoldestknown Smithianfaunamaybeyoungerthantypical Smithianfaunasfromotherareas.Krystyn etal. (2007a,b)proposedtheMudsection(Spitivalley, India)asaGSSPcandidateforthisboundary.They basedtheirdefinitionoftheboundaryonthefirst occurrenceoftheconodont Novispathoduswaageni (Sweet1970) sensulato.However,Bruhwiler etal. (2010b)demonstratedthepresenceinthesamesectionofammonoidgeneratypicaloftheSmithian belowtheboundaryasdefinedbyKrystyn etal. (2007a,b)andthusproposedtousethefirstoccurrenceof Flemingitesbhargavai Bruhwiler etal. (2010b)astheindexfossilforthisboundary.

Tozer(1965,1994)originallysubdividedthe DienerianofCanadaintotwoparts(lowerand upper),eachcomposedofonezone(the Proptychites candidus Zoneandthe Vavilovitessverdrupi Zone), andwiththesecondonebeingfurthersubdivided intothreesubzones.However,thiszonationisbased onscatteredoccurrencesofthefaunas,oftenwithout superpositionalinformation(afactwhichcanbe checkedinthelistoflocalitiesprovidedinTozer 1994).Otherzonationshavebeenproposedfor NorthernSiberia(Dagys&Ermakova1996)andPrimorye,Russia(Shigeta&Zakharov2009),but uncertaintiesincorrelatingthesepersist.Basedon materialfromNepal,abiozonationhasbeenproposedbyWaterhouse(1994,1996)forthenorthern Gondwanamargin.Unfortunately,thisNepalese materialispoorlypreservedanddoesnotallowconstructingarobusttaxonomy(seeGaetani etal. 1995).Jenks etal. (2015)presentedareviewofthe biostratigraphyofTriassicammonoidsandmentionedthatDienerianammonoidfaunaswerestill poorlyknownandinneedofanextensiverevision. TheNorthernIndianMarginhaslongbeenrecognizedasakeyareaforthestudyofEarlyTriassic ammonoidsandtheestablishmentoftheEarlyTriassictime-scale.DuringtheEarlyTriassic,itwassituatedinsouthernTethys,atapalaeolatitudeofca. 40°S(Fig.3).TheveryfirstDienerianammonoids werediscoveredintheSaltRangebyAndrewFleminginthemid-19thcentury,andthismaterialwas describedbydeKoninck(1863).Waagen(1895)

Fig.3. SimplifiedpalaeogeographicalmapoftheEarlyTriassic withthepalaeopositionsofthestudiedlocalities(whitestars)and ofotherlocalitiesmentionedinthetext(blackstars).Modified afterBrayard etal. (2006).

conductedthemostimpressiveandexhaustivestudy onEarlyTriassicammonoidsfromtheSaltRange. Diener(1897)andvonKrafft&Diener(1909)publishedtwomonographsdescribingammonoidsfrom theEarlyTriassicoftheIndianHimalayas,manyof themfromtheSpitiValley.Theveryfirstammonoid biozonationoftheLowerTriassichasbeenpublishedbyMojsisovics etal. (1895)basedonthese tworegions.Theyrecognizedonlytwozonesinwhat weconsiderhereasDienerian,andthisbiozonation didnotchangeintheabsenceofanysubsequent detailedworkonDienerianammonoidsoftheSalt RangeandSpiti.FortheSaltRange,Noetling(1905) andSpath(1934)addedafewspeciesandproposed slightlydifferentclassifications.GriesbachianammonoidswerefirstdiscoveredbySchindewolf(1954)in theSaltRange.Kummelprovidedadetailedhistory ofthestratigraphicalandpalaeontologicalinvestigationsonthePermianandTriassicoftheSaltRange (Kummel1966;Kummel&Teichert1966,1970).He alsomentionedthathecollectednumerousDienerianammonoidsandthatheintendedtopublishthis materiallateron,ataskheunfortunatelynevercompleted.SinceKummel’swork,onlytwocontributionsonEarlyTriassicammonoidsfromtheSalt Rangewerepublished.TheworksbyGuex(1978) andbythePakistani-JapaneseResearchGroup (PJRG1985)onlyincludescarcematerialofGriesbachianandDienerianages.ConcerningammonoidsfromtheIndianHimalayas,onlyBando (1981)describedafewammonoidsfromKashmir, andKrystyn&Orchard(1996)andKrystyn etal. (2004,2007a,b)gavesomedetailsconcerning ammonoidbiostratigraphyofSpiti,butwithoutany descriptionofammonoids.Afewpoorlypreserved

ammonoidsfromSouthTibetwerealsodescribedby Wang&He(1976),andDienerianammonoidsfrom NepalweredescribedbyWaterhouse(1996).OutsidetheNorthernIndianMargin,well-preserved Dienerianammonoidfaunashavebeenstudiedfrom BritishColumbiaandArcticCanada(e.g.Tozer 1994),theVerkhoyanskbasin(Siberia,e.g.Dagys& Ermakova1996),Primorye(Russia,e.g.Shigeta& Zakharov2009),SouthChina(e.g.Bruhwiler etal. 2008)andNevada(Ware etal. 2011).

Presentwork

Thetwoarticlesincludedinthisvolumere-investigatethetaxonomyofDienerianammonoidsfrom theSaltRange(Pakistan)andSpiti(India).Thedata presentedinthetwocontributionsformthebasisof thebiochronologyanddiversitystudypublishedby Ware etal. (2015).

From2007to2010,theresearchgroupatthe UniversityofZurichcarriedoutintensivefieldwork intheSaltRangeandinSpiti.Bedrock-controlled high-resolutionsampling(i.e.samplingammonoids bed-by-bed,withthedrawingandmeasuringofeach section)ofseveralsectionsintheseregionswas performedtorevisetheDienerianammonoidtaxonomyandbuildanew,highlyresolvedbiostratigraphicalscheme.Wheneverpermittedbythesamplesize, greatcarewastakentointegrateontogeneticchanges andintraspecificvariationwithinthedefinitionof species,incontrasttothetraditionaltypological approach.Hence,thenumberofresultingvalidtaxa ismoreconservativeandmorerobustthanthatof previouswork.

Thefirstarticleofthisvolumemainlyaddresses DienerianammonoidsfromtheSaltRange.Asthe SaltRangeisthetypelocalityofmostDienerian ammonoidtaxa,itwastheregionwherethemost intensivefieldworkwasdone,andwherethemost abundantmaterialwasfound.Hence,thisarticle includesanin-depthrevisionofDienerianammonoidtaxonomy,includingemendeddiagnosesof families,generaandspecieswheneverpossible.It alsoincludesthedescriptionofthefewrareGriesbachianammonoidsfoundinthisregion.ThesecondarticleaddressesDienerianammonoidsfrom Spiti(HimachalPradesh,India)andtheircomparisonwiththeSaltRange.

Definitionsofstagesandsubstages

Inthepresentwork,thestageandsubstagesubdivisionoftheLowerTriassicfollowsthe

recommendationsofOgg(2012),loweringthe rankofthefourLowerTriassicstagesofTozer (1965)tothatofsubstage.TheInduanstageis thensubdividedintotheGriesbachianandDieneriansubstages,andtheOlenekianintothe SmithianandSpathian.Thereishoweverpresently noconsensusregardingthedefinitionsofthesubstagesboundariesrelevantforthepresentwork (theGriesbachian–DienerianandDienerian–Smithianboundaries).

TheproblemoftheDienerian–Smithianboundarywasalreadybrieflydiscussed.Here,thedefinition proposedbyBruhwiler etal. (2010b)isadopted,the baseoftheSmithianbeingthendefinedbythefirst occurrenceof F.bhargavai anditsco-occurringspeciescharacterizingUA-ZoneSM-1(Bruhwiler etal. 2010a).Thiszonehasbeenrecognizedineverysectiondescribedherein.

TheGriesbachian–Dienerianboundaryisproblematicandlesswelldocumented.Itwasoriginally definedbyTozer(1965)withthefirstoccurrenceof ‘Meekoceratidae’(i.e.Gyronitidae).InSpiti,Krystyn &Orchard(1996)placedthisboundarywithinthe LowerLimestoneMember,consideringthefauna fromtheupperpartofthisinterval(their Pleurogyronitesplanidorsatus Zone)astypicallyDienerian, basedonthepresenceofabundantGyronitidae,the absenceofthetypicallyGriesbachian Ophiceras and theoccurrenceoftheconodont Sweetospathodus kummeli (Sweet1970).InSpiti,thisdefinitionofthe boundarycoincidesgenerallywithaminorfacies change,withtheappearanceofthinshaleintervals andlessmassive,finergrainedlimestonesthaninthe lowerpartoftheLowerLimestoneMember.This definitionoftheGriesbachian–Dienerianboundary wassubsequentlyquestionedbyKrystyn etal. (2004),whoconsideredthesebedsaslateGriesbachianbutproposedtouseinsteadtheGangetian substageforthewholeLowerLimestoneMember. Krystyn etal. (2004)alsoreportedsomerare Gyronitidae(listedunderthegenusname‘ Pleurogyronites’)associatedwithtypicalGriesbachian Ophiceras inthebedjustbelowtheGriesbachian–DienerianboundaryasdefinedbyKrystyn&Orchard(1996).Thesameammonoidassociationishere documentedintheSaltRange(Ware etal. 2018) whereitisinterpretedascondensation.However, bothKrystyn&Orchard(1996)andKrystyn etal. (2004)reportedexclusivelyGriesbachianconodonts inthisbed,thusquestioningthecondensednature ofthisassociation.ThedefinitionoftheGriesbachian–Dienerianboundaryisthusstillproblematic,andmoreexpandedsectionsspanningthis boundaryarenecessarytosolvethisquestion.Here, wedecidedtoplacetheGriesbachian–Dienerian

boundaryinagreementwiththedefinitionofKrystyn&Orchard(1996).Thisdefinitioncaneasilybe appliedbothintheSaltRangeandinSpiti,thefirst Dienerianfaunalassociationcorrespondinginboth casestothe Gyronitesdubius RegionalZone(equivalentofUA-ZoneDI-1ofWare etal. 2015).

Biochronology:theUnitary AssociationMethodand terminology

FollowingtherecommendationsofMonnet etal. (2015),theconstructionofthebiochronological schemeisherebasedontheUnitaryAssociation MethodofGuex(1991)andGuex etal. (2016).As alreadymentioned,thetwopaperspresentedinthis volumeconstitutethebaseofthebiozonationestablishedbyWare etal. (2015).Asthepresentworkis onlythefirststepinbuildingthisbiozonation,we herepresenttheconstructionof‘RegionalZones’, whicharethenusedtoconstruct‘UnitaryAssociationZones’ofWare etal. (2015).RegionalZones correspondto‘UnitaryAssociationZones’builtfor onlyonebasin,withoutaddressingthelateralreproducibilityofthesezonesoutsideofthestudiedbasin. RegionalZonesarecustomarilytermed‘beds’by previousauthors(e.g.Brayard&Bucher2008; Bruhwiler etal. 2012).Here,theterm‘Regional Zone’ispreferredtoavoidanyconfusionwiththe term‘beds’asitwasusedinSpitibyprevious authorstodesignatesmalllithologicalsubdivisions ofthedifferentunitswithintheMikinFormation (e.g.the‘ Otoceras’and‘ Meekoceras’bedsofvon Krafft&Diener1909;‘ Gyronites’bedsofKrystyn& Orchard1996).Theterm‘LocalMaximalHorizon’ designatesmaximalassociationsofspeciesasdirectly observedwithineachsection.

Inthetwopaperspresentedhere,thelistofcharacteristicspeciesandpairsofspeciesisgivenforeach regionalzone.Aspeciesissaidtobecharacteristicif andonlyifitsrangeisequaltothezone.Apairof speciesissaidtobecharacteristicifandonlyifthe overlappingpartoftheirrangesisequaltothezone. Additionally,thenumberofspecimensofeachspecieswithineachzoneisindicatedinbrackets.This latterinformationgivesanideaoftherobustnessof thedifferentspeciesranges.Whenaspeciesis restrictedtooneregionalzonebutrepresentedonly byafewrarespecimens,itsrelevanceforcorrelations mustbe apriori takenwithcautionasitsstratigraphicalrangeislikelytobelonger.Ontheotherhand,a speciesrepresentedbymanyspecimensand restrictedtooneregionalzoneprovides apriori morerobustinformation.

Eachregionalzonepresentedhereisnamedafter themostabundantofthespecieswhoserangeis strictlyrestrictedtothecorrespondingzone.Formal ‘UnitaryAssociationZones’fortheDienerianofthe NorthernIndianMarginwereconstructedbyWare etal. (2015),andtheircorrelationwitheachregional zonesdescribedhereinissystematicallyprovided here.

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Griesbachian and Dienerian (Early Triassic) ammonoids from the Salt Range, Pakistan by

David Ware, Hugo Bucher, Thomas Brühwiler, Elke Schneebeli-Hermann, Peter A. Hochuli†, Ghazala Roohi, Khalil Ur-Rehman and Amir Yaseen

Acknowledgements

Financial support for the publication of this issue of Fossils and Strata was provided by the Lethaia Foundation

Introduction

Geological framework

Stratigraphy

Lithology and ammonoid preservation

Present work

Biostratigraphy

General subdivisions

Griesbachian ammonoid faunas

Hypophiceras cf. H. gracile Regional Zone

Ophiceras connectens Regional Zone

Ophiceras sakuntala Regional Zone

Early Dienerian ammonoid faunas.

Gyronites dubius Regional Zone.

Gyronites plicosus Regional Zone

Gyronites frequens Regional Zone

Middle Dienerian ammonoid faunas

Ambites atavus Regional Zone

Ambites radiatus Regional Zone……

Ambites discus Regional Zone

Ambites superior Regional Zone……

Ambites lilangensis Regional Zone…

Late Dienerian ammonoid faunas

Vavilovites cf. V. sverdrupi Regional Zone

Kingites davidsonianus Regional Zone

Koninckites vetustus Regional Zone.

Awanites awani Regional Zone

Conclusion

Systematic palaeontology by D. Ware and H. Bucher

Classification

The population approach: intraspecific variability and convergences

Suture lines

Ontogeny and growth allometry

Systematic descriptions

Superfamily Xenodiscaceae Frech, 1902

Family Xenodiscidae Frech, 1902

Genus Hypophiceras Trümpy, 1969

Hypophiceras aff. H. gracile (Spath, 1930)

Superfamily Meekocerataceae Waagen, 1895

Family Ophiceratidae Arthaber, 1911

Genus Ophiceras Griesbach, 1880

Ophiceras connectens Schindewolf, 1954

Ophiceras sakuntala Diener, 1897

Genus Kyoktites n. gen.

Kyoktites hebeiseni n. sp.

Kyoktites cf. H. hebeiseni n. sp.

Genus Ghazalaites n. gen.

Ghazalaites roohii n. sp.

Ophiceratidae? n. gen. A n. sp. A

Ophiceratidae? gen. et sp. indet..

Family Gyronitidae Waagen, 1895

Genus Gyronites Waagen, 1895

Gyronites frequens Waagen, 1895

Gyronites dubius (von Krafft, 1909)

Gyronites rigidus (Diener, 1897)

Gyronites plicosus Waagen, 1895

Gyronites sitala (Diener, 1897)

Gyronites schwanderi n. sp.

Genus Ambites Waagen, 1895

Ambites discus Waagen, 1895

Ambites atavus (Waagen, 1895)

Ambites tenuis n. sp.

Ambites radiatus (Brühwiler, Brayard, Bucher & Guodun, 2008)

Ambites bojeseni n. sp.

Ambites subradiatus n. sp.

Ambites? sp. indet.

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Ambites superior (Waagen, 1895) . .

Ambites lilangensis (von Krafft, 1909)

Ambites bjerageri n. sp.

Ambites cf. A. impressus (Waagen, 1895)

Family Paranoritidae Spath, 1930

Genus Vavilovites Tozer, 1971

Vavilovites cf. V. sverdrupi (Tozer, 1963)

Genus Koninckites Waagen, 1895

Koninckites vetustus Waagen, 1895

Koninckites khoorensis (Waagen, 1895)

Genus Radioceras Waterhouse, 1996

Radioceras truncatum (Spath, 1934)

Genus Pashtunites n. gen.

Pashtunites kraffti (Spath, 1934)

Genus Awanites n. gen.

Awanites awani n. gen. et n. sp.

Genus Koiloceras Brühwiler & Bucher, 2012

Koiloceras sahibi n. sp.

Family Flemingitidae Hyatt, 1900

Genus Xenodiscoides Spath, 1930

Xenodiscoides? sp. indet.

Genus Shamaraites Shigeta & Zakharov, 2009

Shamaraites? sp. indet.

Family Proptychitidae Waagen, 1895

Genus Bukkenites Tozer, 1994

Bukkenites sakesarensis n. sp.

Genus Proptychites Waagen, 1895

Proptychites lawrencianus (de Koninck, 1863) sensu Waagen, 1895

Proptychites oldhamianus Waagen, 1895

Proptychites wargalensis n. sp.

Proptychites ammonoides Waagen, 1895

Proptychites cf. P. pagei Ware, Jenks, Hautmann & Bucher, 2011

Family Mullericeratidae Ware, Jenks, Hautmann & Bucher, 2011

Genus Mullericeras Ware, Jenks, Hautmann & Bucher, 2011

Mullericeras spitiense (von Krafft, 1909)

Mullericeras shigetai n. sp.

Mullericeras indusense n. sp..

Mullericeras niazii n. sp.

Genus Ussuridiscus Shigeta & Zakharov, 2009

Ussuridiscus varaha (Diener, 1895)

Ussuridiscus ensanus (von Krafft, 1909)

Ussuridiscus ventriosus n. sp.

Ussuridiscus ornatus n. sp.

Ussuridiscus? sp. indet.

Family incertae sedis

Genus Kingites Waagen, 1895

Kingites davidsonianus (de Koninck, 1863)

Kingites korni Brühwiler, Ware, Bucher, Krystyn & Goudemand, 2010a

Superfamily Sagecerataceae Hyatt, 1884

Family Hedenstroemiidae Hyatt, 1884

Genus Clypites Waagen, 1895

Clypites typicus Waagen, 1895

Genus Pseudosageceras Diener, 1895

Pseudosageceras simplelobatum n. sp.

Superfamily incertae sedis

Family incertae sedis

Genus Subacerites n. gen.

Subacerites friski n. sp.

Gen. et sp. indet.

Acknowlegdements

References

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GriesbachianandDienerian(EarlyTriassic)ammonoids

fromtheSaltRange,Pakistan

DAVIDWARE,HUGOBUCHER,THOMASBRUHWILER,ELKESCHNEEBELI-HERMANN, PETERA.HOCHULI,† GHAZALAROOHI,KHALILUR-REHMANANDAMIRYASEEN

Ware,D.,Bucher,H.,Bruhwiler,T.,Schneebeli-Hermann,E.,Hochuli,P.A.,Roohi, G.,Ur-Rehman,K.&Yaseen,A.2018:GriesbachianandDienerian(EarlyTriassic) ammonoidsfromtheSaltRange,Pakistan.FossilsandStrata,No63,pp.13 175.doi: 10.1111/let.12273

Intensiveandbedrockcontrolledsamplingoffourareas(NammalNala,Chiddru, AmbandWargal)intheSaltRangeyieldedabundantwell-preservedGriesbachian andDienerian(EarlyTriassic)ammonoids.Thismaterialallowsestablishinganew, high-resolutionbiostratigraphicalframeandanextensiverevisionofthetaxonomy. TheGriesbachianisrepresentedby(inascendingorder)the Hypophiceras cf.H gracile RegionalZone,the Ophicerasconnectens RegionalZoneandthe Ophicerassakuntala RegionalZone.TheDieneriancomprises12distinctregionalzonesleadingtoathreefoldsubdivisionintolower,middleandupperDienerian.ThelowerDienerian,based ontheoccurrenceofthegenus Gyronites,canbedividedintothe Gyronitesdubius RegionalZone,the Gyronitesplicosus RegionalZoneandthe Gyronitesfrequens RegionalZone,inascendingorder.ThemiddleDienerian,basedontheoccurrenceofthe genus Ambites,canbedividedintofivezones:the Ambitesatavus RegionalZone,the Ambitesradiatus RegionalZone,the Ambitesdiscus RegionalZone,the Ambitessuperior RegionalZoneandthe Ambiteslilangensis RegionalZone.TheupperDienerian, whosebaseisdefinedbytheearliestrepresentativesofParanoritidae,canbedivided intofourzones:the Vavilovites cf.V.sverdrupi RegionalZone,the Kingitesdavidsonianus RegionalZone,the Koninckitesvetustus RegionalZoneandthe Awanitesawani RegionalZone.CorrelationswithbasinsoutsidetheNorthernIndianMarginaredifficultbecauseofthescarcityofsuchhighlyresolvedstudiesonDienerianammonoids. Emendeddiagnosesanddetailedsynonymylistsareprovidedformostpreviously knowntaxa.Inaddition,fivenewgenera( Kyoktites, Ghazalaites, Pashtunites, Awanites and Subacerites)and18newspecies( Kyoktiteshebeiseni, Ghazalaitesroohii, Gyronites schwanderi, Ambitestenuis, Ambitesbojeseni, Ambitessubradiatus, Ambitesbjerageri, Awanitesawani, Koilocerassahibi, Bukkenitessakesarensis, Proptychiteswargalensis, Mullericerasshigetai, Mullericerasindusense, Mullericerasniazii, Ussuridiscusventriosus, Ussuridiscusornatus, Pseudosagecerassimplelobatum and Subaceritesfriski)aredescribed. Ammonoidea,biostratigraphy,Dienerian,EarlyTriassic,SaltRange,Pakistan.

DavidWare[david.ware@mfn-berlin.de],MuseumfurNaturkunde,LeibnizInstitutefor EvolutionandBiodiversityScience,Invalidenstrasse4310115Berlin,Germany;Hugo Bucher[hugo.fr.bucher@pim.uzh.ch],ThomasBruhwiler[bruehwiler@pim.uzh.ch],Elke Schneebeli-Hermann[elke.schneebeli@pim.uzh.ch]andPeterA.Hochuli[peter.hochuli@ pim.uzh.ch],PalaontologischesInstitutundMuseumderUniversitatZurich,KarlSchmid-Strasse4CH-8006Zurich,Switzerland;GhazalaRoohi[roohighazala@yahoo.com], KhalilUr-Rehman[reman_geol@yahoo.com]andAmirYaseen[geologistgeologist@ yahoo.com],EarthScienceDivision,PakistanMuseumofNaturalHistory,GardenAvenueShakarparian,Islamabad44000,Pakistan;manuscriptreceivedon27/10/2015; manuscriptacceptedon11/08/2017.

Introduction

Thebioticrecoveryfollowingtheend-Permianmass extinctionisanintensivelystudiedtopic,forwhich highaccuracyandhighprecisiontimecontrolisof paramountimportance.Nekto-pelagiccladessuchas ammonoidsandconodontsrecoveredveryquickly comparedtoothermarineclades(e.g.Brayard etal. 2006,2009;Orchard2007)andplaytheleadingroles indatingofLowerTriassicmarinesedimentaryrocks. However,manystudiesaddressingtherecoveryare basedoninsufficientlyresolvedpalaeontologicalage

controls.ThisisparticularlythecasefortheDienerian,whereammonoidsandbiochronologyarestill poorlyunderstood(Jenks etal. 2015).Areviewofthe currentknowledgeofGriesbachianandDienerian ammonoidsfromtheSaltRangeisgivenintheforewordofthisvolume(Ware&Bucher2018)towhich thereaderisreferred.From2007to2010,ourresearch groupcarriedoutintensivefieldworkintheSalt RangeandtheSurgharRange.Palynologicalandcarbonisotoperecordshavebeenrecentlypublishedby Hermann etal. (2011a,b,2012a,b)andSchneebeliHermann etal. (2012),oxygenisotopesfrombiogenic

2018LethaiaFoundation.PublishedbyJohnWiley&SonsLtd

phosphatesbyRomano etal. (2013),Smithian ammonoidsbyBruhwiler etal. (2012)andbivalvesof SmithianandSpathianagesbyWasmer etal. (2012).

ThepresentworkfocusesonGriesbachianand Dienerianammonoidsfromfourdifferentareasin theSaltRange.Itisbasedonabundantandwell-preservedmaterialsampledbedbybed.Thisnewmaterialprovidesthebasisforacomprehensiverevision ofthetaxonomyandbiostratigraphyofGriesbachian andDienerianammonoidsintheSaltRange,where allrelevantsectionsarefound.Becauseofthe incomparablequalityoftheDienerianammonoid recordoftheSaltRange,thistaxonomicandbiostratigraphicalre-investigationisanessentialcontributiontotheLowerTriassicammonoidzonationof theNorthernIndianMarginandtotheunderstandingoftheEarlyTriassicbioticrecovery.

Geologicalframework

TheSaltRangeconstitutesalongandnarrowmountainrange,approximately150kmSSWofIslamabad,Pakistan(Fig.1B,C).Thesouthernlimitofthe SaltRangedefinestheHimalayanmainfrontal thrust,whichexposesanorthernGondwananrift marginsuccessionrangingfromtheCambrianto theCenozoic.Ittypicallyconsistsofastackoftectonicsliceswithasouthvergence,thusrepeatedly exposingTriassicrocksoftheMianwali,Tredian andKingrialiformations(Gee1980 1981).During theEarlyTriassic,theSaltRangewassituatedinthe southernTethysonthenorthernGondwanamargin, atapalaeolatitudeofca.30 S(Fig.1A).

Stratigraphy

IntheSaltRange,LowerTriassicsedimentaryrocks arereferredtotheMianwaliFormation(Kummel& Teichert1966).This120mthickformationiscomposedoflimestoneandsiliciclasticsedimentary rocks.ItunconformablyrestsontheChanghsingian (UpperPermian)ChiddruFormation.Intheeastern partoftheSaltRange,theMianwaliFormationis truncatedbypost-Cretaceouserosionanddirectly cappedbyPaleocenemarinesedimentaryrocks.In thewesternpart,itisoverlainbytheMiddleTriassic TredianFormation.

TheGriesbachianandDienerianarerepresented bythreeunitsoftheMianwaliFm:theKathwai Member(whichisfurthersubdividedintoadolomiticunitandalimestoneunit),theLowerCeratite LimestoneandthelowerpartoftheCeratiteMarls. Thethicknessofeachoftheseunitsishighlyvariable throughouttheSaltRange.Theirboundariesare

Fig.1. A. PalaeogeographicalmapoftheEarlyTriassicwiththe palaeopositionoftheSaltRange(modifiedfromBrayard etal. 2006). B. MapofPakistanwithpositionofthestudiedarea(black star). C. LocationmapofsampledsectionsintheSaltRange (modifiedafterBruhwiler etal. 2012).

heredemonstratedtobediachronousacrossthedifferenttectonicslices,thussuggestingthatthese thrustsmightcorrespondtoinvertedTriassicnormalfaults.Asatypicalexampleamongmanyothers, Bruhwiler etal. (2012)documentedthattheDienerian SmithianboundarycoincideswiththeLower CeratiteLimestone CeratiteMarlsboundaryin Chiddru,whereasitisfoundwithinthelowerthird (ca.7mabovethebase)oftheCeratiteMarlsin Nammal.Mostpreviousworks(e.g.Mojsisovics etal. 1895;Spath1934;Guex1978;PJRG1985)did notrecognizethisdiachronism,andassumedthat lithologicalboundariesaresynchronousthroughout theSaltRangeandtheSurgharRange.

Lithologyandammonoidpreservation

Despitedifferencesinthicknessesandagesinthedifferentsections,thethreeunitsstudiedhereshow remarkablyuniformfaciesthroughouttheSalt Range.Detailedlithologicaldescriptionsofthese unitswerealreadypublishedinKummel&Teichert (1970)andinHermann etal. (2011b).Therefore, onlyasummaryofthelithologicalsuccessionis givenhere,alongwithadditionalobservations

Mianwali
Kundian
Kamar Mashani Indus
Daud Khel

pertainingtothetaphonomyandpreservationof ammonoids.

KathwaiMember,dolomiticunit.– Thedolomitic unitoftheKathwaiMember(2 3.5mthick)consistsofafewmassivebedsofsandydolomite.Fossils areveryrareinthisunitandareusuallyonlyrepresentedbybrokenandunidentifiableshells.Onlyone ammonite( Hypophiceras aff. H. gracile)wasfound inthisunit,inNammalNala.Itoccurredinasmall lensrichinbivalveswithinamassivedolomiticbed.

KathwaiMember,limestoneunit.– Thelimestone unitoftheKathwaiMember(0.5 5mthick)consistsmostlyofcalcareousglauconiticsandstonebeds alternatingwiththinbedsofshale.Althoughfossils arenotrare,theyaregenerallyverypoorlypreserved. Somebedscontainaccumulationsofrhynchonellid brachiopodsandechinidspines.Ammonitesare rare,andusuallyrepresentedonlybyextremely poorlypreservedspecimens.Identification,evenat thegenuslevel,isimpossible.Therefore,theyhave notbeenincludedinthepresentstudy.Theonly exceptionisinChiddru,wherefew,betterpreserved specimensassignedto Ophicerasconnectens,were foundatthebaseofthisunit.

LowerCeratiteLimestone.– TheLowerCeratite Limestone(1 3mthick)consistsofthin,hard, coarse-grainedcoquinoidlimestonebeds.Glauconite andironoxidesarelocallyveryabundant.Although veryfrequent,thefossilsaremostlyfragmented,and generallyverydifficulttopreparemechanically.The coarsegrainedspariticmatrixoftencrossestheshell boundaries.Ammonoidsareveryunevenlydistributed,oftenimbricatedandaccumulatedinlenses withinthedifferentlayersoftheLowerCeratite Limestone.Thebodychambersofsmallspecimens aregenerallybroken,whilelargespecimens(Proptychitidae)arerepresentedbyincompletephragmocones,theuppersideofouterwhorlsbeingcorroded oreroded(seePl.21,figs38,39foragoodexample). Phragmoconesareoftencompletelyrecrystallized, hencesuturelinesareonlyoccasionallypreserved. Consideringtheabundanceofglauconiteandthe facies,theLowerCeratiteLimestonemaybeaffected bycondensation.However,intheabsenceofsimilar studiesinsectionswheretheLowerCeratiteLimestoneisexpanded,palaeontologicalcondensation cannotbedemonstrated.Onlythesecondbedofthe LowerCeratiteLimestoneinNammalisrecognized ascondensed.Itcontainsboth Ophicerassakuntala and Gyronitesdubius,anassociationofgenerawhich hasnotbeendocumentedinanyothersection. Moreover,thespecies Gyronitesdubius hasalsobeen

foundinAmb,withoutanyco-occurringrepresentativeof Ophiceras.InNammalNala,thetwouppermostbedsoftheLowerCeratiteLimestoneare different,beingcomposedoffinegrainedlimestone. Thepenultimatebedisonly1 2cmthick,locally absent,andcontainsnumerousandnearlycomplete ammonoids,somebivalvefragmentsandabundant fishscalesandteeth.Ammonoidsareaccumulatedat thetopofthebed,oftenencrustedbywormtubes andonlypartiallycoveredbyaverythinlimestone layer(Pl.3,figs1,2).Thelastbedisca.7cmthick andcomposedoffine-grainedlimestonewithlocally abundant,nearlycompletebutstronglyrecrystallized ammonoids.Itssurfaceisencrustedbycentimetric ironoxideconcretions,indicatingastopinsedimentation.

CeratiteMarls.– TheCeratiteMarlsarecomposed ofaca.30mthicksuccessionofmarlswithintercalatedlimestoneandsandstonebeds.Limestonebeds areabundantatthebase,whilesandstonebeds becomegraduallymoreabundantintheupperhalf oftheCeratiteMarls.Inthelowerthird,ammonoids areveryabundantandwellpreservedinnumerous limestonebedsandlenses.Thethicknessofthese bedsisvariable.Manyofthesebedsshowimbricationandsizesortingofshells,indicativeofbottom currents.Atypicalexampleofthisfaciesisshownin Figure2,withsmallimbricated,bothcompleteand brokenammonoidsaccumulatedatthebaseofthe bed.Thebodychambersaregenerallypartiallybroken.Largespecimens(allbelongingtoProptychitidae)oftenhavetheirventerabraded.Forexample, thespecimenillustratedonPlate25,figures10 14is acompletephragmocone,thebodychamberof whichismissing,andwhoseventerhasbeenabraded onalmosttheentirelastpreservedvolution.Inthe DienerianpartoftheCeratiteMarls,involuteshells oftenhavetheirnarrowumbilicusencrustedby bivalvesonbothsides,similartotheonesalready observedbyWare etal. (2011)inDienerianammonoidassemblagesfromNevada.Thesebivalvesmay occasionallyinduceanirregularcoilingofthe umbilicus,thusindicating invivo encrustingofepizoans.Thesebivalvesaresometimesalsopresenton theflanksofthewhorland,whereoverlappedbythe bodychamber,theyarelinedbyadorsalshelllayer. Usually,onlytheircementedvalveispreserved.The bivalvesvisibleontheammonoidfiguredonPlate25, Figure10,weredescribedindetailbyHautmann etal. (2017),togetherwithsimilarspecimensfrom theDienerianofSpiti(India)andtheGriesbachian ofGreenland.Theyidentifiedthemas Liostrea sp. ind.,belongingtothesub-familyGrypheinaeand representingtheoldestknownrepresentativeof

Fig.2. Polishedsectionofabedbelongingtothe Ambiteslilangensis RegionalZone(Nam100andequivalents)fromNammalNala, showingthetypicalfaciesoflimestonebedsatthebaseoftheCeratiteMarls,withanaccumulationofimbricatedshellsatitsbasewitha nautiloid(N),severalimbricatedcompleteandbrokenammonoids(A),andbioturbationincreasingtowardsthetopofthebed.Natural size,arrowindicatestopofbed.SpecimenPIMUZ30235.

Ostreidae.Besidesthisspecimen,theyarenotwell enoughpreservedtobeformerlyidentified,butsince noothercoevalbivalvetaxaareknowntoencrust ammonoids invivo,theycanreasonablybeassumed tobelongtothesametaxa.Itcannotbeexcluded thatsomeoftheintraspecificvariabilityofthe umbilicalwidthobservedontheseinvoluteshellsisa consequenceofthepresenceoftheseepizoans,but asnofaunawithouttheseencrustingbivalvescould befoundforcomparison,thishypothesiscannotbe tested.Thepresenceorabsenceofbivalvesinthe umbilicushasthereforenotbeentakenintoaccount inthetaxonomicalanalyses,exceptwhenthey inducedanobviouspathologicalcoilingofthe ammonoidshell.Such invivo encrustingbivalves havesofarnotbeenrecordedinolderoryounger ammonoidfaunasintheSaltRange,eveninearly Smithianfaunaswhicharefoundinsimilarfacies.It ispossiblethatthisunusualhighfrequencyofepizoanswascausedbythecoevaloxygen-poorwatersedimentinterface,butalternativehypothesesare alsoconceivable(seeHautmann etal. 2017,for details).

Hermann etal. (2011a,b,2012a,b)andSchneebeli-Hermann etal. (2012)proposedadetailed palaeoenvironmentalreconstructionbasedonseveralsections(includingtheonesstudiedhere) throughouttheSaltRangeandSurgharRange.These studiesdocumentedthatinNammal,themiddleto

lateDienerianrecordalocalpeakofoxygendepletion,unlikeprevious(GriesbachiantoearlyDienerian)andsubsequent(earlyandmiddleSmithian) timeintervals.Romano etal. (2013)showedthat temperaturealsopeakedduringthemiddleandlate Dienerian(phaseIbofRomano etal. 2013).

Presentwork

Theammonoidspresentedinthisstudywerecollectedinfourdifferentareas(Fig.1C):Nammal Nala,Chiddru,AmbandWargal,duringfourfield workseasonsfrom2007to2010.Severalsections weremeasuredandsampledwithinacoupleof squarekilometresineachoftheseareas.Following recommendationsofGuex(1991),acompositesectionforeachareawiththepositionofthesamples wasconstructedbasedonlithologicalcorrelations. Therefore,severalsamples(eachwithadistinctnumber)mayhavebeenobtainedfromthesamelayer.

NammalNala

NammalNalaisanarrowcanyonsituatedabout 5kmeastofthevillageofMusaKhel(ca.25km ENEofMianwali).Thisareawaspreviouslystudied byKummel(1966),Kummel&Teichert(1966, 1970),Guex(1978)andthePakistani-Japanese ResearchGroup(1985).TheLowerTriassic

Fig.3. SectionneartheentranceofNammalNala(N32 39 27.6 , E71 47 29.2 ).KM.d:KathwaiMember,dolomiticunit;KM.l: KathwaiMember,limestoneunit;LCL:LowerCeratiteLimestone; CM:CeratiteMarls.Notethepresenceofsmallnormalfaults (blacklines).

MianwaliFormationisbeautifullyexposedand repeatedbyfaulting.Alltheexposuresreportedhere arefromthenorthernsideofthecanyon,whichisin partillustratedinFigure3.Acompositesectionwith theammonoiddistributionisgiveninFigure4.The mostcompleteGriesbachianandDieneriansequence oftheSaltRangewasfoundinNammalNala,especiallyforthebaseoftheCeratiteMarls,where ammonoidsareveryabundantandwellpreserved. Thiscanyonisalsoveryeasilyaccessible,thusallowingintensivesampling.Thevastmajorityofthe ammonoids(about1200specimens)describedhere comefromNammalNala.

Chiddru

ThesectionsstudiedinChiddruaresituatedina valleyabout2.5kmeastofthevillage(ca.25km ESEofMianwali).Severalsectionsweresampledin 2008and2010,butthebestonesfortheDienerian werealreadydescribedbyKummel(1966)and Kummel&Teichert(1966,1970),especiallythe oneonthewestsideofthevalley(Fig.5).Acompositesectionshowingammonoiddistributionis giveninFigure6.Inthisarea,theKathwaiMemberandLowerCeratiteLimestonearemuch

thickerthaninotherareas,andDienerianfaunas arerestrictedtotheLowerCeratiteLimestone.The baseoftheCeratiteMarlsisalreadyearlySmithian inage(Bruhwiler etal. 2012).IntheLowerCeratiteLimestone,ammonoidsareabundantbutgenerallybrokenandpoorlypreserved.Specimenswell enoughpreservedforidentificationareratherrare. Despiteintensivesampling,onlyca.50specimens couldbeincludedinthisstudy.However,thisarea isveryimportantfortheDienerian,sinceitisthe typelocalityof Koninckitesvetustus,thetypespeciesof Koninckites.Thisareahasbeenknown sincethebeginningofthegeologicalinvestigations intheSaltRange,andhasbeenincludedinevery studydealingwiththeEarlyTriassicofthis region.

Amb Ambisasmallvillagesituatedca.35kmeastof Mianwali.Threedifferentoutcropsweresampled duringonefieldtrip(2010)inthevalleyjustsouth ofthevillage,thebestsectionbeingsituatedabout 1kmsouth-eastofthevillage(Fig.7).Acomposite sectionshowingtheammonoiddistributionisgiven inFigure8.TheKathwaiMemberandLowerCeratiteLimestonehavethicknessessimilartothoseof Nammal,buttheDienerianpartoftheCeratite Marlsisthinner,withfewerlimestonebeds. Althoughoftenbroken,ammonoidsfromtheLower CeratiteLimestoneareveryabundantandbetter preservedthaninNammal.IntheCeratiteMarls ammonoidsareratherrare,usuallystrongly recrystallizedanddifficulttoprepare.Atotalof about120specimensfromAmbcouldbeincluded inthisstudy.Thislocalityisthetypelocalityof thetypespeciesofthegenus Ambites ( Ambites discus).SincethepioneerworkofWaagen(1895), theEarlyTriassicammonoidfaunaofAmbhave notbeenstudied.

Wargal

ThesectionsnearWargalaresituatedinasyncline about2.5kmwestofthevillage,alongMuntaNala. AlthoughtheKathwaiMemberandLowerCeratite LimestonearethickerthaninAmb,andtheCeratite Marlsthinner,thetwosectionsareverysimilar.We spentonly3daysinthislocality,mostlytocollect specimensofthegenus Prionolobus,asitisthetype localityofitstypespecies Prionolobusatavus andof Prionolobusrotundatus.About100specimenswere collected,mostlyintheLowerCeratiteLimestone andattheverybaseoftheCeratiteMarls,easily accessibleinthesectionpreviouslydescribedby

RZ Nam60, Nam524 Nam339, Nam354 Nam76, Nam378 Nam320, Nam332 Nam377 Nam331, Nam335a, Nam393 Nam376, Nam390 Nam391 Nam361 Nam350 Nam543, Nam718 Nam540 Nam63, Nam70, Nam71, Nam83, Nam101c, Nam305, Nam312, Nam347, Nam349 Nam58, Nam59, Nam61, Nam67, Nam101d, Nam304, Nam313, Nam336, Nam346, Nam504 Nam62, Nam314 Nam55, Nam316, Nam318, Nam537, Nam724, Nam727 Nam92, Nam100, Nam301, Nam344, Nam501, Nam503 Nam53, Nam302, Nam308, Nam371 Nam50, Nam52, Nam364, Nam370, Nam380, Nam383, Nam527 Nam54, Nam303, Nam382, Nam522, Nam526 Nam521 Nam72, Nam81, Nam379, Nam381, Nam384, Nam520, Nam525 Nam345 Nam56, Nam315, Nam319, Nam337, Nam395, Nam499, Nam502, Nam721 Nam396

Hypophiceras aff. H. gracile

Ophiceras connectens

Ophiceras sakuntala

KM.DKathwaï MemberDolomitic unit

KM.LKathwaï MemberLimestone unit LCLLower Ceratite Limestone

OcOphiceras connectens GpGyronites plicosus GfGyronites frequens AaAmbites atavus ArAmbites radiatus AdAmbites discus AsAmbites superior AlAmbites lilangensis VsVavilovites cf . V. sverdrupi KdKingites davidsonianus KvKoninckites vetustus AwaAwanites awani FbFlemingites bharghavai OG Ophiceras / Gyronites (condensed horizon) HgHypophiceras cf . H. gracile

LMH-1...x : Local Maximal Horizon 1 to x RZ : Regional Zone

Gyronites dubius

Kyoktites hebeiseni n. gen. et sp.

Ussuridiscus? sp. indet.

Gyronites sitala

Ophiceratidae? gen. et sp. indet. Ophiceratidae? n. gen. A n. sp.

Gyronites plicosus

Proptychites oldhamianus

Ghazalaites roohii n. gen. et sp.

Ussuridiscus ensanus

Ussuridiscus ornatus n. sp.

Ussuridiscus ventriosus n. sp.

Gyronites frequens

Gyronites schwanderi n. sp. Ambites atavus

Ambites radiatus

Proptychites ammonoides

Ambites bojeseni n. sp.

Ambites discus

Mullericeras shigetai n. sp.

Ambites? sp. indet.

Ambites superior

Ambites subradiatus n. sp.

Mullericeras indusensis n. sp.

Ambites bjerageri n. sp.

Ambites cf. A impressus

Proptychites lawrencianus

Mullericeras niazii n. sp.

Ambites lilangensis

Proptychites cf P pagei

Vavilovites cf. V

Mullericeras spitiensis

Koninckites khoorensis

Kingites davidsonianus

Clypites typicus

Pseudosagecerasverdrupi s simplelobatum n. sp

Koninckites vetustus

Pashtunites kraffti n. gen. Kingites korni

Awanites awani n. gen. et sp. Radioceras truncatum

Fig.4. CompositesectionofNammalNala:lithostratigraphy,ammonoidoccurrencesandbiostratigraphy.FaunalcontentofNAM543, NAM718andNAM540:seeBruhwiler etal. (2012).

Kummel(1966)andKummel&Teichert(1966, 1970),andillustratedhereinFigure9.Onlythe upperpartofthelowerthirdoftheCeratiteMarls wassampledinanothernearbytributary,wherea

Koninckitesvetustus fauna(sampleWar104)was found.Acompositesectionshowingtheammonoid distributionisgiveninFigure10.AsinAmb, ammonoidsfromtheLowerCeratiteLimestoneare

Fig.5. SectionabovethevillageofChiddru(N32 32 59.7 , E71 47 55.9 )ontheWestsideofthegorge,previouslydescribed byKummel&Teichert(1966,1970)andKummel(1966).Hammerhighlightedforscale.ChiddruFm.,Chiddruformation;KM, KathwaiMember;LCL,LowerCeratiteLimestone;CM,Ceratite Marls.

abundantandratherwellpreserved,buttheyarealso abundantandwellpreservedintheCeratiteMarls. ThisareahasbeeninvestigatedfirstbyWaagen (1895)andlaterbyKummel(1966)andKummel& Teichert(1966,1970).

Biostratigraphy

Theextensivebedrockcontrolledsamplingonwhich thepresentworkisbasedallowstherecognitionofa totalof15regionalzones:threefromGriesbachian and12fromDienerian(Figs11,12).Theresulting zonationisnewwithimprovedresolution.Previously,onlythreezoneswererecognizedfromtheSalt Range(Guex1978;Pakistani-JapaneseResearch Group1985),andtwozonesandfoursubzoneswere establishedintheDieneriansuccessionofCanada (Tozer1965,1994).Apreliminaryversionofthis zonationwaspublishedinRomano etal. (2013). ThisregionalzonationservedasbasefortheconstructionoftheformalbiozonationfortheDienerianoftheNorthernIndianmarginpalaeoprovince ofWare etal. (2015).

Thedifferentregionalzonesaredescribedherein andtheircorrelationwithammonoidzonations fromotherbasinsisdiscussed.Syntheticrange chartsforGriesbachianandDienerianammonoid speciesandgenerafromtheSaltRangearegivenin Figures13and14,respectively.Technically,mostof theseregionalbiozonescorresponddirectlytolocal maximalhorizons,exceptforthe Gyronitesplicosus RegionalZone,the Gyronitesfrequens RegionalZone andthe Vavilovites cf. V. sverdrupi RegionalZone

(seebelow).Itshouldbenotedthatcorrelationof thisnewbiozonationwithpreviousones(e.g.Mojsisovics etal. 1895;Guex1978;Pakistani-Japanese ResearchGroup1985)cannotbeprovidedbecause theundetecteddiachronicityofthedifferentlithostratigraphicalunitsledtomixammonoidfaunasof different(lateGriesbachiantoearlySmithian)ages fortheLowerCeratiteLimestone.TheagedistributionoftheLowerCeratiteLimestonevariesgeographically,thisunitbeingupperGriesbachianto lowerDienerianinNammalNala,lowerDienerian tolowermiddleDienerianinAmbandWargal,and DieneriantolowermostSmithianinChiddru.These resultscontrastwiththepreviousageassignmentof theLowerCeratiteLimestonetoonesinglezone, namelythe Gyronitesfrequens Zone.

Foreachregionalzone,thelistofco-occurring speciesisgiven,andthecharacteristicspeciesand pairsofspeciesareindicatedinbraces.Additionally, thenumberofspecimensofeachspeciesineach zoneisgiveninbrackets.

Generalsubdivisions

Thestageandsub-stagesubdivisionoftheLower Triassicandtheirdefinitionswerealreadydiscussed intheforeword(Ware&Bucher2018;thisvolume). Tozer(1965)subdividedtheGriesbachianandthe Dienerianintotwoparts(lowerandupper)each,a schemewhichhasbeenfollowedbyeveryauthor since.BecauseofthepaucityofGriesbachianfaunas intheSaltRange,thistwofoldsubdivisioncannotbe assessedhere.However,fortheDienerian,themuch higherresolutionsubdivisionsobtainedinthisstudy leadstoathreefoldsubdivisionoftheDienerian (lower,middleandupper).ThelowerDienerianis characterizedbytheoccurrenceofthegenera Gyronites and Ussuridiscus,themiddleDienerianbythe co-occurrenceofthegenera Ambites and Mullericeras,andtheupperDienerianbytheappearance ofParanoritidaeandSagecerataceae.

Griesbachianammonoidfaunas

Griesbachianammonoidsareveryrareandgenerally poorlypreservedintheSaltRange.Moreover,Griesbachianammonoidsfromotherregionsofthe NorthernIndianMarginhavenotbeeninvestigated indetailsincetheworkofDiener(1897)andvon KrafftandDiener(1909).Anewbiozonationforthe GriesbachianofSpitiValley(IndianHimalayas)has beenattemptedbyKrystyn etal. (2004),butwithout explanationorillustrationofthetaxonomicdefinitions,thusmakingthisschemedifficulttoapplyor totestinotherareas.Asaconsequence,thethree

CH7A, Chi57, Chi56

Ophiceras connectens AmbitesatavusProptychitesamm onoides Mullericeras shigetai n. sp. Ambi tes dis cus Ambi tes subradi atus n. sp. Pse udosageceras simplelo batum n. sp. Konin ckites khoo rensis Konin ckites vet ustus Koiloce ras sa hibi n. sp. Radio ceras truncatum Kingites korni

Gyronites frequens Gyronites plicosus

LMH-1...x : Local Maximal Horizon 1 to x RZ : Regional Zone

SrShamaraites rursiradiatus

FbFlemingites bhargavai

KvKoninckites vetustus

AdAmbites discus

AaAmbites atavus

GfGyronites frequens

GpGyronites plicosus

OcOphiceras connectens

LCLLower Ceratite Limestone

KM.LKathwaï Member - Limestone unit

KM.DKathwaï Member - Dolomitic unit

Limestone Sandy limestone Shales

Dolomite

Concretion

Observed occurence

Virtual occurence

Observed occurence

Fig.6. CompositesectionofChiddru:lithostratigraphy,ammonoidoccurrencesandbiostratigraphy.FaunalcontentofCHI63,CHI52, CHI104andCHI53:seeBruhwiler etal. (2012).

Griesbachianregionalbiozonesdescribedherecannotbecorrelatedwithconfidence.

Hypophiceras cf. H.gracile RegionalZone Co-occurringspecies.–Hypophiceras cf. H.gracile ( n 1).

Occurrenceintheinvestigatedsections.– Thiszoneis herebasedonasinglespecimenfoundinthedolomiticunitoftheKathwaiMemberinNammalNala.

Correlation.– Althoughspeciesassignmentremains uncertain,identificationatthegenuslevelofthis singlespecimenisveryrobust.Thegenus Hypophiceras isknownintheArctic(Siberia,ArcticCanada andNEGreenland),whereitoccurstogetherwith lowerGriesbachiantaxa.However,Kummel(1970) reportedonespecimenfromtheKathwaiMemberof Kathwai,whichmaybeconspecificwithours,and correlatedthisbedonthebasisoflithologywiththe Ophicerasconnectens ZoneofChiddru(upperGriesbachian).Intheabsenceofadditionalmaterial,and Xeno discoi des? sp. indet

Sectionabout1kmsouth-eastofthevillageofAmb(N32 29 48.1 ,E71 56 20.6 ).Scaleindicatedbyageologist(circled)inthe middleoftheview.KM.d:KathwaiMember,dolomiticunit;KM.l:KathwaiMember,limestoneunit;LCL:LowerCeratiteLimestone; CM:CeratiteMarls.

consideringthatthiscorrelationisexclusivelybased onlithology,thiszoneisherekeptseparatefromthe subsequent Ophicerasconnectens regionalZone. AdditionalworkontheGriesbachianoftheTethys isnecessarytodecipherwhetherthegenus Hypophiceras isrestrictedtothelowerGriesbachianorifit rangesupintotheupperGriesbachian.Thiszone correspondstothehorizonMH-G1inRomano etal. (2013).

Ophicerasconnectens RegionalZone

Co-occurringspecies.–Ophicerasconnectens ( n 13).

Occurrenceintheinvestigatedsections.– Thiszone hasbeenidentifiedinNammalNala(baseofthe LowerCeratiteLimestone)andinChiddru(baseof limestoneunitoftheKathwaiMember)andonly yieldspoorlypreservedspecimensof Ophicerasconnectens

Correlation.– Thisspecieswasconsideredasa synonymof Ophicerastibeticum byWaterhouse (1994),whothustreatedthisfaunaasanequivalentoftheHimalayan Ophicerastibeticum zone oflateGriesbachianage.Acceptanceorrejection ofthistreatmentrequiresrevisionofthetaxonomyofOphiceratidaeandoftheHimalayan Griesbachianfaunalsuccession.ThiszonecorrespondstothehorizonMH-G2inRomano etal. (2013).

Ophicerassakuntala RegionalZone

Co-occurringspecies.–Ophicerassakuntala ( n 4).

Occurrenceintheinvestigatedsections.– Thisregionalzonehasonlybeenrecognizedinthecondensed layeratthebaseoftheLowerCeratiteLimestonein NammalNala.

Correlation.–Ophicerassakuntala alsooccursin ShalshalCliffinthecentralHimalayas,whereitwas originallydescribedbyDiener(1897).ThiscondensedlayercorrespondstothehorizonMH-G3in Romano etal. (2013).

Remarks.– BecausethislayeryieldsbothlateGriesbachianandearliestDieneriantaxa(LMH-3,Fig.4), thenaturalassociationofspeciesbelongingtothis regionalzoneisobscured.

EarlyDienerianammonoidfaunas

EarlyDienerianammonoidsfromtheSaltRangeare abundantandusuallyquitewellpreserved,butcan bedifficulttomechanicallyseparatefromthematrix. TheyareallfoundintheLowerCeratiteLimestone, anintervalcharacterizedbylowaccumulationand sedimentationrate.ThelowerDienerianbiostratigraphicalrecordmustbetreatedwithcautionwith respecttopotentialhiatusesandcondenseddeposition.TheSaltRangeneverthelessprovidesthemost

Fig.7.

Gyronites dubius Kyoktites cf. K. hebeiseni n. gen. et sp.

expandedanddiversefaunalsuccessionfortheearly Dienerianworldwide.Alikelycaseofcondensationis foundatthebaseoftheLowerCeratiteLimestonein NammalNala,where Gyronitesdubius ofearliest DienerianageoccurstogetherwiththelateGriesbachian Ophicerassakuntala (LMH-3,Fig.4).

ThelowerDienerianisbestcharacterizedbythe genus Gyronites.IntheSaltRange,thegenus Ussuridiscus isalsorestrictedtothelowerDienerian, butinPrimorye,itstypespecies Ussuridiscusvaraha isfoundinfourconsecutivebedsbelongingtothree differentzones(Shigeta&Zakharov2009),ranging fromtheupperGriesbachiantothemiddleDienerian,whereasitisrestrictedtoasinglezoneinthe SaltRange.ItspresenceintheupperGriesbachianis uncertain,asitsoccurrenceinthissubstageisbased

Ussuridiscus varaha

Bukkenites sakesariensis

Gyronites frequens Gyronites rigidus Gyronites plicosus

Gen. et sp. indet.

Ghazalaites roohii n. gen. et sp. Ussuridiscus ensanus Ambites atavus AmbitestenuisProptychit es amm onoides Mullericeras shigetai n. sp. Ambi tes dis cus Ambi tes subradi atus n. sp. Konin ckites khoo rensis Kingites davidsonian us Clypites typicus Konin ckites vetustus Suba ceri tes f riski n. gen. et sp. Pasht unites krafft i n. ge n. Flemingi tes barghavai

LMH-1...x : Local Maximal Horizon 1 to x RZ : Regional Zone

Flemingites bharghavai Fb KvKoninckites vetustus

KdKingites davidsonianus

AdAmbites discus

AaAmbites atavus

GfGyronites frequens

GpGyronites plicosus

GdGyronites dubius

LCLLower Ceratite Limestone

KM.LKathwaï Member - Limestone unit

KM.DKathwaï Member - Dolomitic unit

Limestone

Dolomite

Concretion

Sandy limestone

Shales

Observed occurence

Observed occurence Virtual occurence

Fig.8. CompositesectionofAmb:lithostratigraphy,ammonoidoccurrencesandbiostratigraphy.AMB53andAMB11werecollectedon adipslopeformedbythetwouppermostbedsoftheLowerCeratiteLimestone,withoutanyfurtherdistinction.

onbed1009ofShigetaandZakharov(2009,fig.15), whichyieldedapoorlypreservedspecimenassigned to Lytophiceras?sp.indet.However,thepresenceof Ussuridiscusvaraha inthemiddleDienerianisconfirmedbyitsassociationinbed1013ofShigeta& Zakharov(2009)with‘ Ambitoides’ fuliginatus (here re-assignedto Mullericeras), Proptychitesammonoides and‘ Gyronites’(herere-assignedto Ambites onthebasisofthebottleneckshapeoftheventer).

The Gyronitessubdharmus Zone,whichShigetaand Zakharov(2009)assignedtotheupperGriesbachian, containsthegenus Gyronites andthereforelikely correlateswiththelowerDienerianasdescribed here.However, Gyronitessubdharmus isabsentin theSaltRange,soacorrelationatthespecieslevel cannotbemade.Krystyn etal. (2004)placedtheir

Fig.9. SectionofMuntaNala,about2.5kmWestofthevillageofWargal(N32 27 07 ,E72 01 56.7 ;Kummel1966;Kummel&Teichert1966,1970,).ThicknessofKathwaiMemberandLowerCeratiteLimestoneamountstoca.6m.KM.d:KathwaiMember,dolomitic unit;KM.l:KathwaiMember,limestoneunit;LCL:LowerCeratiteLimestone;CM:CeratiteMarls.

‘ Pleurogyronites’bedsfromSpitiintotheupper Griesbachian.Thesebedsalsoyield Gyronites and thusbettercorrelatewithourlowerDienerian.AdditionalmaterialfromtheselayersinSpitiisdescribed inthecompanionpaperofthisvolume(Ware etal. 2018).Innorth-westernGuangxi(SouthChina), Bruhwiler etal. (2008)foundsomespecimensinitiallyascribedto‘ Koninckites’cf.‘ K.’ timorense but whichweresynonymizedwith Ussuridiscusvaraha byShigetaandZakharov(2009;anassignment whichwepartiallyconfirmhere),sotheir Proptychitescandidus bedsmaypartiallycorrelatewiththe lowerDienerianasdefinedhere(whereastheoriginal Proptychitescandidus ZoneofTozerishereconsideredasmiddleDienerian;seebelow).Thegenus Gyronites hasneverbeenreportedoutsideofthe Tethys,butconodontsfromthe Bukkenitesstrigatus ZoneoftheCanadianArcticindicatethatthetopmostpartofthiszonemayactuallyalreadybe Dienerian(Orchard2008)andthusmayinpartcorrespondtoourlowerDienerian.

Gyronitesdubius RegionalZone

Co-occurringspecies.– Bukkenitessakesarensis ( n 21) , Ghazalaitesroohii ( n 26), Gyronites dubius ( n 27) , Gyronitesrigidus ( n 1) , Kyoktites cf. K. hebeiseni ( n 2)and Ussuridiscusvaraha ( n 7) .

Occurrenceintheinvestigatedsections.– Thisregionalzoneisherebasedontheassociationfoundat thebaseoftheLowerCeratiteLimestoneinAmb.It alsooccursatthebaseoftheLowerCeratite

LimestoneinNammalNala,butitisthereincluded inacondensedlayer(LMH-3,Fig.4).

Correlation.–Gyronitesdubius and Gyronitesrigidus alsooccurinseverallocalitiesintheIndianHimalayas,wheretheywerefirstdescribedbyDiener (1897)andvonKrafftandDiener(1909),indicating furtherextensionofthisfaunatotheNorthern IndianMargin.Thisregionalzonecorrespondsto UA-zoneDI-1inWare etal. (2015)andpartially tothecondensedhorizonMH-G3inRomano etal. (2013).

Gyronitesplicosus RegionalZone

Co-occurringspecies.–Ghazalaitesroohii ( n 19), Gyronitesplicosus ( n 85) , Gyronitessitala ( n 2) , Proptychitesoldhamianus ( n 2), Proptychiteswargalensis ( n 3) , Ussuridiscusensanus ( n 8), Ussuridiscusornatus ( n 3) , Ussuridiscusventriosus ( n 1)

Occurrenceintheinvestigatedsections.– Thisregionalzonehasbeenrecognizedineveryinvestigated section,inseveralbedsinthemiddleoftheLower CeratiteLimestone,exceptinChiddruwhereit occursinthelowerthirdoftheLowerCeratite Limestone.

Correlation.–Gyronitessitala, Ussuridiscusensanus andsomespeciesheresynonymizedwith Gyronites plicosus and Proptychitesoldhamianus describedby Diener(1897)andvonKrafftandDiener(1909) occurinseverallocalitiesintheIndianHimalayas,

Ussurid iscus orna tus n. sp. Ghazala ites roohii n. gen. et sp Gyroni tes frequens Ussuridis cus ensanus Proptychites warg alensis Gyronites plicosus

Gyroni tes schwanderi n. sp. Ambites atavus Propty chites ammo noides Ambites discus Ambites su bradia tus n. sp. Pseudosagece ras sim pleloba tum n. sp. Clypites typicus Koninc kites vetus tus Shamarai tes ? sp. indet. Radioceras truncatum Koil oceras sa hibi n. sp.

LMH-1...x : Local Maximal Horizon 1 to x RZ : Regional Zone

KvKoninckites vetustus

AdAmbites discus

AaAmbites atavus

GfGyronites frequens

GpGyronites plicosus

LCLLower Ceratite Limestone

KM.LKathwaï Member - Limestone unit

KM.DKathwaï Member - Dolomitic unit

Limestone

Sandy limestone Shales Dolomite

Concretion

Observed occurence

Observed occurence Virtual occurence

Fig.10. CompositesectionofWargal:lithostratigraphy,ammonoidoccurrencesandbiostratigraphy.WAR11andWAR100werecollectedonadipslopeformedbyseveralbedsinthemiddlepartoftheLowerCeratiteLimestone,withoutanyfurtherprecision.

indicatingthebroaderextensionofthisfauna throughouttheNorthernIndianMargin.ThisregionalzonecorrespondstohorizonMH-D1(Romano etal. 2013)andtotheUA-zoneDI-2(Ware etal. 2015).

Remarks.–Gyronitessitala, Proptychitesoldhamianus, Ussuridiscusornatus and Ussuridiscusventriosus wereonlyfoundinNammal(LMH-4,Fig.4), whereas Proptychiteswargalensis hasonlybeenfound inWargal(LMH-1,Fig.10),thusformingtwodistinctlocalmaximalhorizons.Thesetwolocalmaximalhorizonsareherelumped,asthisdistinction maybetheresultofthescarcityofsomeoftheir respectivecharacteristicspecies.Moreover,thesetwo

localmaximalhorizonshavenotbeenfoundin sequenceanywhere,thusprovidinganadditional argumenttomergethem.

Gyronitesfrequens RegionalZone

Co-occurringspecies.–Ghazalaitesroohii ( n 1), Gyronitesfrequens ( n 90) , Gyronitesschwanderi ( n 2) , Proptychitesoldhamianus ( n 1), Ussuridiscusensanus ( n 4).

Occurrenceintheinvestigatedsections.– Thisregionalzoneoccursineveryinvestigatedsection:inthe twotopmostbedsoftheLowerCeratiteLimestone inNammalNala,inthepenultimatebedofthe

Salt Range

This work, Brühwiler et al. 2012

Shamaraites rursiradiatus RZ

Flemingites bhargavai RZ

Awanites awani RZ

Koninckites vetustus RZ

Kingites davidsonianus RZ

Vavilovites cf. V. sverdrupi RZ

Ambites lilangensis RZ

Ambites superior RZ

Ambites discus RZ

Ambites radiatus RZ

Ambites atavus RZ

Gyronites frequens RZ

Gyronites plicosus RZ

Gyronites dubius RZ

? Ophiceras sakuntala RZ?

Ophiceras connectens RZ

Hypophiceras cf. H. gracile RZ

Spiti (Himalaya)

Brühwiler et al. 2010

Krystyn et al. 2004, 2007

Kashmiritidae gen. nov. A bed

Flemingites bhargavai beds

Prionolobus rotundatus beds

Kingites lens beds

Fuchsites markhami beds

Ambites lilangense Zone

“Pleurogyronites” planidorsatus Zone

Discophiceras Zone

Ophiceras tibeticum Zone

Otoceras woodwardi Zone

NW Guangxi (South China)

Brayard & Bucher 2008

Brühwiler et al. 2008

Kashmirites kapila beds

Clypites sp. indet. beds

South Primorye

Markevich & Zakharov 2004

Shigeta et al. 2009

Arctic Canada & British Columbia Tozer 1994

Paranorites varians Zone Hedenstroemia hedenstroemi Zone

Clypeoceras spitiense Zone

Vavilovites sverdrupi Zone

Proptychites candidus beds

Ophiceras beds

Ambitoides fuliginatus Zone

Proptychites candidus Zone

Gyronites subdharmus Zone

Lytophiceras Zone

Bukkenites strigatus Zone

Ophiceras commune Zone

Otoceras boreale Zone

Fig.11. BiostratigraphicalsubdivisionsoftheGriesbachian,DienerianandearliestSmithianoftheSaltRangeandcorrelationwith zonationsofotherregions.Thickverticalblackbarsindicateuncertaintyintervalsforcorrelations.The O.sakuntala beds,indicatedwith questionmarksandadashedframe,correspondtoahorizonidentifiedonlyinthecondensedhorizonofNammalNala(LMH-3,Fig.4). Seetextfordetails(RZ:RegionalZone).

LowerCeratiteLimestoneinAmbandWargal,and inonebedinthemiddleoftheLowerCeratiteLimestoneinChiddru.

Correlation.– SomespeciesdescribedbyvonKrafft &Diener(1909)areheresynonymizedwith Gyronitesfrequens,thusindicatingtheextensionofthis regionalzoneintheIndianHimalayas.SomespecimensdescribedbyWang&He(1976)andWaterhouse(1996)mayalsobelongto Gyronitesfrequens, indicatingthatthisregionalzonemayalsoexpandto NepalandTibet.Thisregionalzonecorrespondsto thehorizonMH-D2inRomano etal. (2013)andto UA-zoneDI-3ofWare etal. (2015).

Remarks.– InNammalNala,thisregionalzoneis representedbytwodistinctlocalmaximalhorizons (LMH-5andLMH-6,Fig.4).Theyareonlydifferentiatedbytworarespecies( Gyronitesschwanderi and Ghazalaitesroohii)andarethuslumped here.

MiddleDienerianammonoidfaunas

Dependingontheareas,middleDienerianammonoidsoccureitherintheLowerCeratiteLimestone oratthebaseoftheCeratiteMarls.Themiddle Dienerianisheredefinedbytheoccurrenceof

Ambites,agenuswhichisgenerallyveryabundant andwithaworldwidedistribution.Thegenus Mullericeras ismostlyrestrictedtothemiddleDienerian,exceptforonespecies( Mullericerasspitiense) whichextendsupintothelateDienerian.InSpiti, Ambites isveryabundantinthelowerhalfofthe ‘ Ambites beds’,inthe Ambiteslilangense zoneand atthebaseofthe Fuchsitesmarkhami beds(Bruhwiler etal. 2010a).Severalspeciesof Ambites were foundindifferentsectionsinnorth-western Guangxi(Bruhwiler etal. 2008),whichwereall consideredasbelongingtothe Proptychitescandidus beds.Inthiswork,theauthorsdidnotprovideany moreprecisesubdivisionsbecauseofthescattered occurrenceofthefaunas,butacomparisonwith thefaunasfromtheSaltRangeshouldallowthe refinementofthebiozonationforthisarea.In SouthPrimorye, Ambites ispresentinbed1013of Shigeta&Zakharov(2009),correspondingtotheir ‘ Ambitoides’ fuliginatus Zone.

CorrelationwiththeDienerianbiozonation establishedbyTozer(1965,1994)isnotclear. Thefaunasfromnorth-easternBritishColumbia describedbyTozer(1963,1994)aredifferent fromthosedescribedbyTozer(1961,1994)from ArcticCanada.Hencecorrelationbetweenthese twobasinsisuncertain.WhatTozerreferredtoas Proptychitescandidus inBritishColumbia,on

Fig.12. BiostratigraphicalcorrelationsofChiddru,Wargal,AmbandNammalNalasections.NotethediachronismoftheLowerCeratite Limestonebetweenthedifferentsections.

whichhiscorrelationwithArcticCanadaisbased, isinouropinionnotconspecificwiththespecies originallydescribedbyTozer(1961)inArctic Canada.

Moreover,severalspeciesclearlybelongingto thegenus Ambites occurinBritishColumbia, indicatingthattheCandidusZoneofthisregion ismiddleDienerian.However,Tozerassigned

onlyonespecies( Ambitesferruginus)fromArcticCanadato Ambites.Thelatterdiffersfrom allother Ambites speciesbyitsinvolutecoiling andunusuallythicktrapezoidalwhorlsection. Hence,thecorrelationoftheCandidusZoneof ArcticCanadawithourmiddleDieneriancannotbeconfirmed.ThezonationofTozer(1994) isbasedonscatteredoccurrencesofthese

DIENERIAN GRIESB. late late middle early e.

Ophiceras connectens RZ ? Ophiceras sakuntala RZ? Gyronites plicosus RZ Gyronites frequens RZ Ambites atavus RZ Ambites radiatus RZ Ambites discus RZ

Hypophiceras cf . H. gracile RZ Gyronites dubius RZ Ambites superior RZ Ambites lilangensis RZ

Vavilovites cf . V. sverdrupi RZ Kingites davidsonianus RZ Koninckites vetustus RZ Awanites awani RZ

Hypophiceras aff. H gracile

Ophiceras connectens

Ophiceras sakuntala

Kyoktites hebeiseni n. gen. et sp.

Ussuridiscus? sp. indet.

Ophiceratidae? n. gen. A n. sp.

Gyronites dubius

Gyronites rigidus

Kyoktites cf. K. hebeiseni n. gen. et sp.

Ussuridiscus varaha

Ophiceratidae? gen. et sp. indet. Gen. et sp. indet.

Ghazalaites roohii n. gen. et sp.

Gyronites sitala

Gyronites plicosus

Proptychites wargalensis

Ussuridiscus ventriosus n. sp.

Ussuridiscus ornatus n. sp.

Proptychites oldhamianus

Ussuridiscus ensanus

Gyronites frequens

Gyronites schwanderi n. sp.

Ambites atavus

Ambites tenuis n. sp.

Mullericeras shigetai n. sp.

Proptychites ammonoides

Ambites radiatus

Ambites bojeseni n. sp.

Ambites discus Bukkenites sakesariensis

Ambites? sp. indet.

Ambites subradiatus n. sp.

Ambites superior

Mullericeras indusensis n. sp.

Ambites bjerageri n. sp.

Ambites cf. A impressus

Ambites lilangensis

Proptychites lawrencianus

Proptychites cf. P pagei

Mullericeras niazii n. sp.

Mullericeras spitiensis

Vavilovites cf. V sverdrupi

Koninckites khoorensis

Pseudosageceras simplelobatum n. sp.

Clypites typicus

Kingites davidsonianus

Subacerites friski n. gen. et sp.

Koninckites vetustus

Pashtunites kraffti n. gen.

Kingites korni

Koiloceras sahibi n. sp.

Xenodiscoides? sp. indet.

Shamaraites ? sp. indet.

Radioceras truncatum

Awanites awani n. gen. et sp.

Fig.13. RangechartshowingthebiostratigraphicaldistributionofGriesbachianandDienerianammonoidspeciesintheSalt Range.DashedthicklinescorrespondtospeciesofuncertainagefoundinthecondensedhorizonofNammalNala(RZ:Regional Zone).

faunas,oftenwithoutsuperpositionalinformation.Additionalinvestigationsintheseareas wouldprobablyallowtheconstructionofa moredetailedbiozonationandmoreaccurate correlations.

Ambitesatavus RegionalZone

Co-occurringspecies.– Ambitesatavus ( n 74) , Ambitestenuis ( n 5) , Mullericerasshigetai ( n 1), Proptychitesammonoides ( n 1).

Kyoktites n. gen.

Ophiceratidae n. gen. A

Ghazalaites n. gen. ?

Hypophiceras Ophiceras Gyronites

Ambites

Vavilovites

Koninckites

Pashtunites n. gen.

GRIESB. late late middle early e.

Koninckites vetustus RZ

Kingites davidsonianus RZ

Vavilovites cf. V. sverdrupi RZ

Ambites lilangensis RZ

Ambites superior RZ

Ambites discus RZ

Awanites awani RZ DIENERIAN

Ambites radiatus RZ

Ambites atavus RZ

Gyronites frequens RZ

Gyronites plicosus RZ

Gyronites dubius RZ

? Ophiceras sakuntala RZ?

Ophiceras connectens RZ

Hypophiceras cf. H. gracile RZ

Occurrenceintheinvestigatedsections.– Thisregionalzonehasbeendocumentedineverystudied locality:atthebaseoftheCeratiteMarlsin NammalNala,inthetopmostbedoftheLower CeratiteLimestoneinAmbandWargal,andina bedinthemiddleoftheLowerCeratiteLimestoneinChiddru.

Correlation.–Ambitesatavus hassofaronlybeen recordedfromtheSaltRange.ThisregionalzonecorrespondstothehorizonMH-D3inRomano etal. (2013)andtoUA-zoneDI-4inWare etal. (2015).

Ambitesradiatus RegionalZone

Co-occurringspecies.– Ambitesbojeseni ( n 18) , Ambitesradiatus ( n 104) , Mullericerasshigetai ( n 3), Proptychitesammonoides ( n 2).

Occurrenceintheinvestigatedsections.– ThisregionalzonehasonlybeenrecognizedinNammalNala, inagroupofnodularbedsabout1mabovethebase oftheCeratiteMarls.

Radioceras

Awanites n. gen.

Bukkenites Koiloceras

Kingites

Proptychites

Shamaraites ? Xenodiscoides ?

Ussuridiscus

Mullericeras

Pseudosageceras

Clypites

Subacerites n. gen.

Fig.14. RangechartshowingthebiostratigraphicaldistributionofGriesbachianandDienerianammonoidgenera(groupedbyfamilies) intheSaltRange.DashedthicklinescorrespondtogeneraofuncertainageoccurringinthecondensedhorizonofNammalNala.Grey thicklinescorrespondtovirtualoccurrences(RZ:RegionalZone).

Correlation.–Ambitesradiatus alsooccursinJinya innorth-westernGuangxi,whereitwasoriginally describedbyBruhwiler etal. (2008),extendingthis faunatotheSouthChinablock.Thisregionalzone correspondstothehorizonMH-D4byRomano etal. (2013)andtoUA-zoneDI-5ofWare etal. (2015).

Ambitesdiscus RegionalZone

Co-occurringspecies.– Ambitesdiscus ( n 163) , Ambitessubradiatus ( n 8) , Mullericerasshigetai ( n 5), Proptychitesammonoides ( n 34).

Occurrenceintheinvestigatedsections.– Thisregionalzonehasbeenrecordedineverystudiedlocality:in agroupofbedsabout2mabovethebaseoftheCeratiteMarlsinNammalNala,atthebaseoftheCeratite MarlsinAmbandWargal,andinabedinthemiddle oftheLowerCeratiteLimestoneinChiddru.

Correlation.– SomespeciesdescribedbyDiener (1897)andvonKrafftandDiener(1909)fromseveral localitiesintheIndianHimalayasarehere

synonymizedwith Ambitesdiscus,thusenlargingthe distributionofthisregionalzonetothisregion.Some specimensdescribedbyWang&He(1976)andMu etal. (2007)couldpossiblybelongto Ambitesdiscus, thussuggestingthepresenceofthe Ambitesdiscus ZoneinTibetandnorth-westernGuangxi,respectively.OnespeciesfromtheCandelariaHills (Nevada: Ambites aff. A.radiatus;Ware etal. 2011)is heresynonymizedwith Ambitessubradiatus,thus indicatingthecorrelationofthe Ambitesdiscus Zone withtheCandelariaFormationinNevada.The Ambitesdiscus regionalZonecorrespondstothehorizonMH-D5inRomano etal. (2013)andtoUA-zone DI-6inWare etal. (2015).

Ambitessuperior RegionalZone

Co-occurringspecies.– Ambitessuperior ( n 48) , Mullericerasindusense ( n 6) , Proptychites ammonoides ( n 4).

Occurrenceintheinvestigatedsections.– ThisregionalzonehasonlybeenrecognizedinNammalNala area,inabedabout3mabovethebaseoftheCeratiteMarls.

Correlation.– Onespecimendescribedbyvon KrafftandDiener(1909)maybelongto Ambites superior,indicatingthatthisregionalzonemay alsoexpandtotheIndianHimalayas.ThisregionalzonecorrespondstothehorizonMH-D6in Romano etal. (2013)andtoUA-zoneDI-7in Ware etal. (2015).

Ambiteslilangensis RegionalZone

Co-occurringspecies.– Ambitesbjerageri ( n 32) , Ambiteslilangensis ( n 64) , Mullericerasniazii ( n 11) , Mullericerasspitiense ( n 2), Proptychiteslawrencianus ( n 35) , Proptychites cf. P pagei ( n 5)

Occurrenceintheinvestigatedsections.– ThisregionalzonehasonlybeenidentifiedinNammalNala, inabedabout4mabovethebaseoftheCeratite Marls.

Correlation.– Onespecies,originallyascribedto Gyronitesfrequens byBruhwiler etal. (2008)and heresynonymizedwith Ambitesbjerageri n.sp., occursinShangganinnorth-westGuangxi,indicatingthecorrelationofthisregionalzonetoSouth China. Ambiteslilangensis anditssynonymshave beenfoundinvariouslocalitiesworldwide:inSpiti ValleyinIndia(vonKrafft&Diener1909),inBritish

Columbia(Tozer1994),inNepal(Waterhouse 1996)andinNevada(Ware etal. 2011).SomespecimensdescribedbyWang&He(1976)mayalso belongto Ambiteslilangensis,indicatingthatthis regionalzonemayalsobepresentinTibet.This regionalzonecorrespondstothehorizonMH-D7in Romano etal. (2013)andtoUA-zoneDI-8inWare etal. (2015).

LateDienerianammonoidfaunas

UpperDienerianammonoidsoccurintheupper partofthelowerthirdoftheCeratiteMarlsin Nammal,AmbandWargal,andinthesecondhalf oftheLowerCeratiteLimestoneinChiddru.The upperDieneriansubstageisheredefinedbythefirst occurrenceofParanoritidae,asexemplifiedbythe typicallyDieneriangenera Koninckites, Vavilovites and Awanites.Sagecerataceaealsoappearinthis interval,andtheraregenus Clypites hassofaronly beendocumentedinthistimeinterval.Theupper DieneriansubstagecaneasilybeidentifiedinSpiti, whereParanoritidaeoccurinabundanceinthe upperpartofthe‘ Ambites beds’ofBruhwiler etal. (2010a),atthetopofthe‘ Fuchsitesmarkhami beds’,inthe‘ Kingiteslens beds’andinthe‘ Prionolobusrotundatus beds’.Aspeciesdescribedas Proptychites sp.indet.byBruhwiler etal. (2008) fromYupingsectionisheresynonymizedwith Koninckiteskhoorensis,thusindicatingthepresenceof thebaseoftheupperDienerianinthe‘ Proptychites candidus beds’ofnorth-westernGuangxi.Inthe samearea,the‘ Clypites sp.indet.beds’ sensu Brayard&Bucher(2008)fromWailisectionarealso mostlikelylateDienerianinageasthisgenusisso faronlyknowninthisinterval.InPrimorye(Shigeta&Zakharov2009),onespeciesoriginally ascribedto Clypeocerasspitiense isheresynonymizedwith Clypitestypicus,aSagecerataceae whichissofaronlyknownfromthelateDienerian. Ambitoidesorientalis,withwhichitco-occurs,hasa suturelinetypicalofParanoritidae,sothe Clypeocerastimorense zoneofPrimoryeismostlikely upperDienerian.Thegenus Vavilovites ishereconsideredasarepresentativeofParanoritidae,witha typicalrangeintheupperDienerian.Therefore,the Vavilovitessverdrupi ZoneofTozer(1994)correlatesatleastinpartwiththeupperDieneriansubstage.However,correlationofthethreesubzones distinguishedinthe Vavilovitessverdrupi Zoneis difficult.Thefirstsubzoneisbasedonasinglecharacteristicspecies( Koninckitesdimidiatus Tozer 1994)forwhichnoequivalentisknownanywhere else.Thesecondsubzonecontainsthetypespecies of Vavilovites,andthusclearlybelongstotheupper

Dienerian.Thethirdoneischaracterizedbytwo speciesbelongingtogenerawhichdonotoccurin theSaltRange: Heibergitesheibergensis and‘ Kingites’discoidalis.Theassignmentofthelatterto Kingites ishererejected(seetaxonomicdescriptions)andnocorrelationscanbemadeonthebasis ofthesetwotaxa.Tozer(1994,p.24)alsodescribes somespeciesasbeingderived‘frombedsinthe ToadFormationthatcloselyfollowSubzone2 , amongwhich Flemingitesreticulatus and Xenodiscoidesscapulatus areundoubtedlyofSmithianaffinity.Thisfaunamaythuscorrelatewiththe lowermostSmithianasdefinedbyBruhwiler etal. (2010b).InEasternVerkhoyansk(Siberia;Dagys& Ermakova1996),theDienerianStageisdividedinto threezones.Thefirstoneisaclearcorrelativeof the Vavilovitessverdrupi Zone,andisthusupper Dienerian.Thesecondzoneissubdividedintotwo subzonescharacterizedby Vavilovitessubtriangularis and Vavilovitesumbonatus,respectively.Theassignmentofthesetwospeciestothegenus Vavilovites is uncertainbecauseoftheirdifferentwhorlsections. Otherco-occurringspeciesarealsoreferredtoas Vavilovites andareagainrestrictedtothisregion. Therefore,norobustcorrelationscanbeestablished betweentheSubtriangularisZoneofEastern VerkhoyanskandtheSaltRange.Thethirdzoneis characterizedby Kingites? korostolevi,aspecies whoseassignmentto Kingites ishererejected(see taxonomicdescriptions).Italsocontains Sakhaitoidesallarense, Sakhaitoidesverkhoyanicum and Episagecerasantiquum,threespeciesforwhichno equivalentareknownanywhereelse,thusmaking itsassignmenttotheupperDienerianuncertain.

Vavilovites cf. V.sverdrupi RegionalZone

Co-occurringspecies.– Koninckiteskhoorensis ( n 54), Mullericerasspitiense ( n 11) , Pseudosagecerassimplelobatum ( n 1), Vavilovites cf. V sverdrupi ( n 5)

Occurrenceintheinvestigatedsections.– ThisregionalzonehasonlybeenrecognizedinNammal,in twoconsecutivebedsca.4mabovethebaseofthe CeratiteMarls.

Correlation.– Thetwospecimensassignedto Vavilovites sp.indet.fromfloatedblocksintheCandelariaHills(Nevada;Ware etal. 2011)arehere synonymizedwith Vavilovites cf. V. sverdrupi,thus documentingthisregionalzoneinNevada.This regionalzonecorrespondstothehorizonMH-D8in Romano etal. (2013)andtoUA-zoneDI-9ofWare etal. (2015).

Remarks.– ThetwobedsofNammalassignedtothis regionalzonecorrespondtotwodifferentlocalmaximalhorizons(LMH-12andLMH-13;Fig.4).The firstonecontainsonlythirteenspecimensbelonging to Vavilovites cf. V. sverdrupi and Mullericerasspitiense.Thesecondlocalmaximalhorizoncontains58 specimens,including54 Koninckiteskhoorensis,three Vavilovites cf. V. sverdrupi andone Pseudosageceras simplelobatum.Becauseofthesmallsamplesizeofthe firsthorizonandthescarcityof Mullericerasspitiense (thespecieswhichdistinguishesthefirstlocalmaximalhorizonfromthesecondone),thesetwolocal maximalhorizonsarelumpedtogether.

Kingitesdavidsonianus RegionalZone

Co-occurringspecies.– Clypitestypicus ( n 11), Koninckiteskhoorensis ( n 776) , Kingitesdavidsonianus ( n 75) , Pseudosagecerassimplelobatum ( n 50), Subaceritesfriski ( n 1) .

Occurrenceintheinvestigatedsections.– Thisregionalzoneoccursinthreesuccessivebedsbetween5 and6mabovethebaseoftheCeratiteMarlsinNammal.IthasalsobeenidentifiedinAmb,inabedabout 4mabovethetopofLowerCeratiteLimestone.

Correlation.– InSpitivalley,the Kingiteslens beds (Bruhwiler etal. 2010a)areanexactcorrelativeof thisregionalzone.Thisregionalzonecorrespondsto thehorizonMH-D9inRomano etal. (2013)andto UA-zoneDI-10ofWare etal. (2015).

Remarks.– InChiddru,asingleisolatedspecimenof Koninckiteskhoorensis hasbeenfoundca.1mbelow thetopoftheLowerCeratiteLimestone(LMH-6, Fig.6),withoutfurtherassociatedspecies.Itmay belongtothisregionalzoneortothepreviousone.

Koninckitesvetustus RegionalZone

Co-occurringspecies.–Clypitestypicus ( n 11), Kingiteskorni ( n 14), Koilocerassahibi ( n 7) , Koninckitesvetustus ( n 189) , Pashtuniteskraffti ( n 11) , Pseudosagecerassimplelobatum ( n 16), Radiocerastruncatum ( n 9).

Occurrenceintheinvestigatedsections.– The Koninckitesvetustus Zonehasbeenidentifiedinevery studiedlocality:inNammalNalaca.7mabovethe baseoftheCeratiteMarls,inAmbca.5mabovethe baseoftheCeratiteMarls,inWargalca.4mabove thebaseoftheCeratiteMarls,andinChiddruatthe topoftheLowerCeratiteLimestone.

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the Society for six months and who were in the employment of the Society at the date of commencing the fund, if they were sixteen years of age or upwards and not over fifty years of age in the case of males or forty-five years of age in the case of females, should be members of the fund. The proposed scale of contributions to the scheme was 5 per cent. of the wages or salary received, and contributors were to be eligible at sixty years of age to retire on pension if they so desired. At the age of sixty-five for males and fifty for females they would be eligible to receive annuities ranging from 25 per cent. of their salaries, after ten years’ payment of contributions, to 85 per cent. of their salaries after having paid contributions for fifty-one years. To assist in launching the scheme it was proposed that the Society should make an initial contribution of £10,000. It was also proposed that the superannuation fund should be managed by a committee of seven, which committee should consist of the chairman and three directors for the time being of the U.C.B.S. and three representatives of the employees, who must have at least three years’ service with the Society.

When the scheme was brought forward again at the September meeting of the Society one of the amendments sent in was from St George Society, and called for the rejection of the scheme in its entirety. The motion to reject the scheme was seconded by a representative of the employees, who referred to the “autocratic” methods of the directors in adopting this scheme and bringing it forward without consulting the workers. The result was that the scheme was disapproved. The scheme which was brought forward by the S.C.W.S. for the superannuation of their employees suffered the same fate.

THE INTERNATIONAL CO-OPERATIVE CONGRESS.

It was as the result of an invitation which came from the Scottish National Co-operative Conference, held in Kilmarnock in 1910, that the International Co-operative Congress held in Hamburg in the autumn of that year decided to come to Glasgow for the 1913 Congress rather than go to Vienna, the claims of which city were strongly urged by Dr Beno Karpeles, in opposition to the claims of Glasgow, as set forth by Mr James Deans.

The year 1913 was a big year for Scottish Co-operators, for not only had they to entertain the International Congress in the autumn, but Scotland was also the location of the British Congress in the summer, and pessimists were not wanting who thought that the Co-operators of Scotland had undertaken a task which was too heavy for them when they decided to entertain two so great Congresses as the British and the International in one year. Events proved that the pessimists were wrong, however. One of the first duties which fell to the lot of the committee which was appointed to make preparations for the International Congress was that of finding a suitable house for it, but they did not require to look far afield. The U.C.B.S. were the owners of the best hall in the city for the purpose for which it was required. In St Mungo Hall there was ample accommodation for the Congress itself, while in the adjoining halls ample space for dining the delegates could be provided. The Bakery was just across the street, and cooking and waiting facilities were all that could be desired. So it was decided that in St Mungo Hall the Congress should be held.

The U.C.B.S. undertook all the work of catering for the delegates at luncheon each day. They also undertook the provision of the luncheon which was given by the reception committee on the Saturday, and they themselves provided the entertainment for one of the afternoons of Congress. It was universally acknowledged that the International Congress of 1913 was the best International Cooperative Congress held, and to this happy result the U.C.B.S. contributed no small share. The event was one which will not readily

be forgotten by those privileged to take part. Alas, that the expressions of fraternity so freely uttered then should have been so soon made of no avail by the outbreak of war.

THE HOLIDAY CAMP.

It is to Mr John Dewar, for many years president of the Renfrewshire Co-operative Conference Association, that the idea of a Co-operative holiday camp owes its origin. For many years Mr Dewar was an enthusiastic Volunteer, and his experiences under canvas during the annual training periods of his regiment impressed him with the value of this form of holiday. Associated with him in his propaganda for a camping association on Co-operative lines was Mr James Lucas, at that time president of the Glasgow and Suburbs Conference Association, and latterly, also, Mr John Paton, of the Renfrewshire Conference council, who had been converted to the idea as the result of a visit to Douglas, I.O.M., where he had seen the huge city under canvas which for a number of years housed thousands of holiday-making Lancashire lads every summer.

By 1910 these gentlemen had been able to get their organisation so far advanced that they had selected a site on the Ayrshire Coast for their first camp, and had made arrangements with the farmer who rented the land. At the last moment, however, the landowner stepped in and vetoed the whole proceedings. This put an end to doing anything further with regard to a camp during that year, but the search for a suitable site continued and, at length, the little farm of Roseland, situated on Canada Hill, Rothesay, overlooking the Bay, was secured. The farm was for sale, but the committee in charge of the arrangements considered that purchase was too bold an initial step to take, so they leased the farm for six months; securing an option to purchase at the end of that period if they wished.

Here, in the summer of 1911, the first Scottish Co-operative holiday camp was established. It was rather a primitive affair, that first camp. The cooking was done in the little farmhouse, while the campers had their meals in a large marquee. The U.C.B.S. directors took a keen interest in the camp from the very beginning. The catering was done by them, and the catering staff were housed in the little farmhouse.

Primitive though the arrangements were, they appealed to the campers, who were unanimous in their praise of the beautiful situation, the pure air, the perfect catering, and the small outlay for which they secured a perfect holiday. Thus encouraged, the committee which had promoted the camp proceeded to organise a Co-operative society to work it, and in this Co-operative society the U.C.B.S. took out twenty-five shares. The farm was purchased for £600, and in September the Baking Society increased the number of their shares to 100.

In 1912 the camp was much better organised than in 1911, but it was still far from being what its promoters desired to see it. They were hampered for lack of funds, however, as the Co-operative societies were showing caution and a lack of faith in the enterprise, and were not providing the capital necessary to work it properly as readily as had been expected. The only fault which the committee found with the site lay in the fact that in dry summers the water supply was inadequate. The summer of 1912 also showed them that it was desirable that something more impervious to rain than a marquee was desirable for the gatherings of campers and, in order that these two defects might be put right, they applied to the U.C.B.S. for a loan of £1,000 on the security of the property. This loan was granted them, and so good use did they make of the power which it gave them that, before the time came for opening the camp in 1913, they had put down a huge storage tank for water, capable of storing 20,000 gallons; and had erected a dining hall large enough to dine several hundred persons.

The camp was a very great success in the third year. Its popularity was so great that the committee found it quite impossible to provide accommodation for all who wished to avail themselves of its facilities for holiday making, and this has been the case in each succeeding year, notwithstanding the influence of the war. At the end of the third season, however, the committee of the association came to the conclusion that, if the camp was to be made the success they believed it was capable of becoming, some rearrangement of its management would require to be made, so they invited the Baking Society to take it over as a going concern and work it themselves.

They explained to the directors of the Baking Society that they were not taking this step because they disbelieved in its success, but

solely on the ground that they considered that dual control was not good for discipline and did not make for good management.

The directors of the Baking Society promised to consider the matter, and the result was they brought forward to the quarterly meeting, held in March 1914, a recommendation that the camp should be taken over, and this recommendation was accepted by the delegates. Since then the camp has been managed by the U.C.B.S.

In 1914 accommodation was provided for 250 persons, and it is extremely probable that greatly increased accommodation would have been provided before now had it not been for the intervention of the war which, by providing another and much more strenuous form of camping for the past and prospective frequenters of Canada Hill, prevented for the time being such further developments. Doubtless, however, with the return of Europe to sanity, such developments will take place; until, before many years are past, almost the whole of the Society’s seven-acre estate will be covered in the summer and autumn months with the picturesque pyramids of white canvas.

THE SOCIETY’S PROGRESS.

In all its branches, with the exception of the tearooms, the progress of the Society during these four years had been remarkable. In 1913, however, the London Street tearoom was closed, and as soon as the lease of the Union Halls expired they also were given up. At the beginning of the period the output had averaged 3,820 sacks per week, while at its end the average output was 4,648, an increase of 848 sacks per week in four years. The aggregate sales for the year which ended in July 1914 were £692,600. Truly, the Society had travelled far from the days when a small two-oven bakehouse sufficed for all its output. The membership now consisted of 201 societies; which was also a contrast to the eight small struggling societies which had banded themselves together in the last days of December 1868 to form the Federation.

The time had now arrived when the Federation was to be put to a more severe test than ever before since it had attained to years of maturity. Like a thunderclap the war storm which had been gathering over Europe during the month of July burst on an astonished world which had almost come to believe war on such a gigantic scale impossible, and many were the doubts expressed, even by sincere wellwishers of the Co-operative movement, as to how it would weather the storm. The next chapter will tell how one federation kept the Co-operative flag flying and added to the laurels it had gained by its devoted and loyal service to the interests of the common people.

CHAPTER

XVIII.

BAKING UNDER WAR CONDITIONS.

THE EARLY DAYS OF WAR—KEEPING DOWN PRICES: THE DIRECTORS’ VIEW—HALF-PAY TO SOLDIER EMPLOYEES —ARMY CONTRACTS—HORSES AND MOTORS COMMANDEERED—PRICE OF BREAD ADVANCED— DEATH OF BISCUIT FACTORY MANAGER—MORE ARMY CONTRACTS—EXTENSIONS AND ALTERATIONS—M‘NEIL STREET BLOCK COMPLETED—BUSINESS ORGANISATION —TEABREAD AND PASTRY SHOPS—THE PRICE OF BREAD —INCREASING OUTPUT—BISCUIT AND TEABREAD TRADE—THE SOCIETY AND ITS WORKERS—DELIVERY DIFFICULTIES—DEATH OF MR DUNCAN M‘CULLOCH.

On the fourth day of August 1914, Great Britain entered on a new era. There are not awanting cynics who declare that Great Britain is hardly ever out of war, but however that may be, there never has been in the world’s history a war which seemed in its early days to be fraught with so dire consequences for civilisation as did that into which the nations of Europe plunged in those early days of August 1914. For the first few days it seemed as if chaos had suddenly developed. The people seemed to go mad with panic, and rushed to purchase goods wherever these could be procured, careless of the prices which they were called on to pay. To the traders, large and small, it must have seemed like the beginning of a trading millennium, and they took immediate advantage of the position in which they found themselves. Prices doubled, trebled, and quadrupled; yet the eager rush to buy continued. The poor bought what they could, but they soon reached the end of their meagre

resources. Not so the rich, however. Amongst them, the panic seemed to be even greater than amongst the poorer sections of the community, and they provisioned their houses as if for a siege, careless as to who must want if only they had plenty in store.

EDUCATIONAL DEPUTATION TO ENGLAND

EDUCATIONAL DEPUTATION TO ENGLAND

The bakers of Glasgow were also the victims of the profiteering craze which had suddenly enmeshed the trading public, and at a meeting which took place on the afternoon of 3rd August, they proposed that the price of bread should be raised. Mr James Young, manager of the U.C.B.S., had been present at this meeting and had opposed the proposal, with the result that it was not carried out. This action of his was homologated by his board at a special meeting which was held the same evening, while at another meeting, held three days later, the committee decided that they would not raise the price of bread in the meantime. The Baking Society found itself in a very favourable position at this time. In M‘Neil Street there were supplies of flour which would keep the bakery going for nine weeks, while Clydebank had supplies for six, and Belfast for seven weeks. They were thus in a position to view the situation with more or less equanimity. The S.C.W.S. were also in a good position so far as supplies of flour and wheat were concerned, having some nine or ten weeks’ supplies either on hand or on board ship at the outbreak of war, and the rapidity with which the German fleet was bottled up within its own harbours gave security of passage for the time being to food-carrying vessels. Nevertheless, the general panic of that first week of war affected the wheat and flour trade as much as it did

many others, and it was not long before flour was selling at a price which had never been asked for it in the whole course of the Federation’s history.

These were conditions under which the two great Co-operative federations were able to show to advantage, and they were conditions also which proved the wisdom of the Wholesale Society in establishing its own purchasing agency in Canada; for while outside millers were asking as much as 65/ a sack for flour in the first week of the war, the highest price quoted by the Wholesale Society was 37/, and ordinary flour was sold by them at 1/6 per sack advance on normal price. Thus the two federations were in a position to see to it between them that on this side of the Atlantic, at least, no undue advantage was taken of the circumstances created by the war. At the meeting of the Baking Society’s board, which took place on 1st September, the committee placed on record their appreciation of the manner in which the crisis brought about by the war had been met by the manager, heads of departments, and employees generally, and desired the manager to convey this expression of the board’s appreciation to the heads of departments and employees. At the quarterly meeting of the Society, the directors had a similar compliment paid to themselves; Mr Shaw, Cambuslang Society, moving, and Mr Glasse, S.C.W.S., seconding a motion which received the unanimous endorsement of the delegates, thanking the directors for the splendid lead which they had given to the purveyors of bread in the city and far beyond. Mr Shaw drew attention to the importance and magnificence of what had been done, and thought that the nation was entitled to recognise what the Co-operative movement had done, not only now but at all times, in keeping down prices and in keeping goods pure.

Perhaps the position in which the Federation was placed by the war and the view of it taken by the directors can be explained most clearly in the words of Mr Gerrard himself. Addressing the September quarterly meeting, he stated that the profits for the halfyear which had ended in July had been a little better than usual, but [3]“the future was so indefinite that no one could foresee with any accuracy what would happen. They had been blamed, and were still being blamed in certain quarters, for not increasing the price of bread, but they had resolutely refused, thinking it their duty as a

Federation to keep the price of bread and other commodities as low as possible. Notwithstanding, he understood the price of biscuits had been increased in some quarters, and this had brought them some trade that they should have had before. The way in which prices were rushed up at the commencement of the war had seemed to the board most unseemly and unpatriotic, and seemed to show that everyone was trying to make as much as possible out of the war. It was unfortunate that the price of bread would require to be increased before the war was over, but until that step was absolutely necessary they would not increase it, and then only as far as was necessary. They had received several contracts for bread, and the manner in which the manager, heads of departments, and the general body of the employees acted under the stress of these circumstances was worthy of all praise.”

3. (The Scottish Co-operator, 25th September 1914.)

HALF-PAY TO SOLDIER EMPLOYEES.

At the outbreak of war, a number of the employees had been members of the Territorial Forces and had been called up at once, and the board decided to grant them half-pay. Later, when the call for more men came, many of the employees joined up, and the directors decided that those who enlisted should be placed on the same footing as regarded half-pay as were those who had been called up at the outbreak of war. At this quarterly meeting the directors were accused by a representative of the Bonus Investment Society of not being so generous to their employees who were serving with the Colours as were some other Co-operative societies, but the chairman was able to satisfy the delegates that, in those cases where generosity was most necessary—those of wives with large families—the Baking Society’s scheme was the one which gave the wife the larger total income.

This meeting was the one at which the method of giving the minutes in the form of synopses was first adopted, and one of the delegates took occasion to compliment the secretary on having produced synopses of the various minutes which gave the delegates enough information about the business done to enable them to find out what was taking place. At this meeting also a grant of £100 was made to the Belgian Relief Fund, the chairman stating that further grants could be made if occasion demanded. Other donations made were:—£500 to the War Relief Fund, £100 to the Lord Mayor of Belfast’s Fund, and £100 to the fund being raised by the Provost of Clydebank. It was urged very strongly by several of the delegates that the administration of the relief fund should be in the hands of the Government. In the minute of the board meeting of 2nd October, the fact is noted that during the past quarter the Society’s motors had covered a distance of 125,015 miles. It had certainly been a busy quarter.

ARMY CONTRACTS.

Immediately the war broke out, the United Baking Society were inundated with Army contracts. In the first few days they had to make arrangements for supplying camps all over the country, in addition to those for which they had held the contracts. These new camps included those at Perth, Inverness, Falkirk, and Dunblane, and all were supplied at current prices. Yet, notwithstanding the fact that the Baking Society were doing much work for the Army in that first week of the war, they were not allowed to escape from some of the other discomforts which come to the owners of horses and vehicles in a nation at war. On the 6th of August 18 horses, two motors, and one lorry were commandeered for military purposes. Other large bakers suffered in the same way, and on behalf of all of them a telegram was sent to the Officer in Command, pointing out the importance of the work which was being done by the baking firms in the distribution of food, and complaining that serious hardship to the people would ensue if bakers’ horses were removed. With this telegram the committee of the Baking Society associated themselves. To enable them to overtake their work, the Society decided to purchase several motors which had been offered to them, while a circular-letter, explaining the position, was sent out to the societies. That the War Office officials were not ungrateful for the assistance which had been rendered them in the emergency of mobilisation was shown by the fact that at the board meeting which was held on 7th August, letters of congratulation on the manner in which the service of bread to two camps had been carried out were read to the committee.

Among the minor results due to the war may be mentioned the cancelling of a number of social meetings, excursions, marriages, etc., and the closing down of the holiday camp at the end of August. At the beginning of October, another attempt was made by the Glasgow Master Bakers’ Association to have the price of bread raised, but again the directors of the Baking Society blocked the way. At the same time, however, the committee came to the decision that, in view of all the circumstances, they would not raise any objection

should another overture be made. At the beginning of October, the situation was again reviewed and, in view of the fact that by the beginning of the year the price of flour would be greatly in advance of that then being used, it was agreed to resist no longer a slight advance in the price of bread. For three months the Society had been the means of keeping the price of bread at the rate at which it was being sold at the outbreak of war. If we assume that their action influenced only two hundred thousand households, and that the average consumption of bread in each household was only one 2–lb. loaf per day, this action of the Baking Society was responsible for saving to these householders in three months no less a sum than £21,250, and probably double that sum, for that only supposes an advance of one farthing per loaf. The first advance in the price of bread took place on 16th November. In Belfast the price of bread had been advanced ¼d. per 2–lb. loaf on 19th October.

DEATH OF BISCUIT FACTORY MANAGER.

Mr John Gilmour took charge of the Society’s biscuit factory shortly after it was commenced, and he continued to manage it until his death, which took place on 9th October 1914. He was highly respected by both directors and workmen for his devotion to business and his tact and kindness in dealing with those under him. He was succeeded by Mr William Ninian, who had the distinction of being a Bakery-trained man and who was acting as Mr Gilmour’s assistant at the time of that gentleman’s death.

At the quarterly meeting, the chairman made fitting and sympathetic reference to the loss which the Society had sustained through the death of Mr Gilmour. He also referred to the retiral from active work of one of the Society’s bread bakers, Mr W. Lees, who had been in the employment of the Society for forty and a half years. During the quarter an electrically-driven motor had been at work in Glasgow for the Society. It was proving very satisfactory and the board, he said, considered that before long these electrically-driven cars would replace horses on the streets, although petrol-driven cars would still be used for the long journeys.

At this quarterly meeting, some of the delegates wished to know why the board had decided that they would no longer resist an increase in the price of bread seeing that in their minutes they stated that their contract for flour was still unbroken. The chairman, in replying, pointed out that flour at the moment was about 10/ higher in price than before the war, and the board thought it better that the price should be raised by a halfpenny now than that it should be put a penny or three halfpence later, when the stock of cheap flour was completely exhausted. He also stated that eighty-six of the Society’s employees were then serving in the Army. At this quarterly meeting, also, the question of granting relief to what were termed “innocent enemy aliens,” in other words, British women who had married subjects of nations at war with Britain, and who through the operation of war were left destitute in this country, was raised through an appeal on their behalf issued by the International Co-

operative Alliance executive. A proposal was made to grant £25 to the fund which was being raised, but this was defeated by a two to one majority. The cake show was held this year as usual, but, as was to be expected, the sales were not so great as in some former years. The price had been advanced on the average one penny per pound. In their report to the delegates to the December quarterly meeting, the directors stated that they would regard it as an instruction to make no increase in the prices of their goods so long as they were able to work without actual loss.

MORE ARMY CONTRACTS.

In September, an arrangement had been entered into with the Army Authorities by which the Government supplied the flour and the Baking Society baked it into bread for the troops. In addition, several contracts for biscuits for the Army were received. The first three of these totalled 200 tons. The contracts for the baking of bread for the Army continued until early in 1916, when an intimation was received from the Quartermaster of the Scottish Command that the contract would cease, as the Government were now erecting field bakeries for themselves. Two months later, however, another contract was entered into with the Scottish Command on the same terms as formerly. The contracts carried out by Clydebank and Belfast Branches have already been referred to.

EXTENSIONS AND ALTERATIONS.

For the whole of the first year of war and almost the whole of the second year, the building department of the Society was kept busy with the extension to the M‘Neil Street premises. The principal reason for this extension was the necessity for securing more space for the biscuit factory, and as soon as practicable this extension was carried through. A new travelling oven was installed early in 1915 at a cost of £175, and at the same time a new biscuit cutting machine was got at a cost of £300. A gas-fired travelling oven was also installed before the end of the year and another one, built in accordance with alterations suggested by the bakery staff, was installed in October 1916. In the spring of 1915, it was decided to cover over the space between bakehouses Nos. 11 and 12 for the purpose of providing dressing rooms for the workers, and at the same time it was decided to put fans in each flat for ventilation purposes.

M‘NEIL STREET BLOCK COMPLETED.

Parts of the new block at the Adelphi Street and South York Street corner were occupied as they were completed. Particularly, the ground floor was occupied as an extension to the biscuit factory, but it was not until June 1916 that the entire building was ready for the official opening ceremony. This was of a very modest character, for, as the chairman stated, the directors did not think the circumstances of the time lent themselves to the celebration of the opening of their splendid addition by means of a grand function, and in this he believed the board had interpreted the wishes of the delegates.

The dressing room for the female workers, which was situated on the fourth flat, was fitted up in an excellent manner. Each worker had a locker of her own, made of cast steel, of which she alone held the key. Fourteen wash-hand basins were fitted up and eight spray baths. There was also an excellently appointed sickroom, where a girl who became unwell suddenly could rest, and also a room fitted up with the appropriate scientific appliances, where accidents could be treated at once.

At the quarterly meeting, which took place on the day on which the new wing of the building was opened, the chairman stated that the total cost of the land and the buildings on it had been £11,800, while the total cost of the bakery buildings, including the addition, had been £131,000; which had been depreciated to the extent of £52,000.

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