Notable invertebrates associated with maritime cliffs and slopes The listed species include only those which are particularly characteristic of cliffs. Many more species of grassland also occur on cliffs, particularly cliff tops. Snails and slugs (Mollusca) Soft cliffs: Truncatellina callicratis Paludinella littorina Abida secale Hard cliffs: Paludinella littorina Ponentina subvirescens
RDB 3 RDB 3 WCA (Schedule 5) Nb RDB 3 WCA (Schedule 5) Na
Spiders and allies (Arachnida: Araneae and Pseudoscorpiones) Callilepis nocturna RDB1 Clubiona genevensis RDB3 Episinus maculipes RDB3 Gnaphosa occidentalis RDB1 Hahnia candida RDB2 Haplodrassus umbratilis RDB3 Lathys stigmatisata RDB3 Neon robustus RDBK Acartauchenius scurrilis Na Aelurillus v-insignitis Nb Anelosimus aulicus Nb Argiope bruennichi Na Dipoena inornata Nb Dipoena prona Nb Episinus truncatus Nb Euophrys herbigrada Na Gnaphosa lugubris Na Liocranum rupicola Nb Micaria romana Nb Micrargus laudatus Nb Myrmarachne formicaria Nb Nigma puella Nb Ozyptila blackwalli Nb Porrhomma errans Nb Rugathodes bellicosus Nb Segestria bavarica Na Trachyzelotes pedestris Nb Trochosa robusta Nb Xysticus acerbus Na Xysticus robustus Na Zilla diodia Nb Millipedes (Diplopoda) Trachysphaera lobata Thalassisobates littoralis
RDBK Nb
Enantiulus armatus
Nb
Centipededes (Chilopoda) Lithobius tenebrosus Hydroschendyla submarina Schendyla peyerimhoffi Geophilus fucorum Geophilus pusillifrater Chalandea pinguis Arenophilus peregrinus
RDBK Nb Nb Nb Nb Nb unknown
Woodlice (Isopoda) Metatrichoniscoides celticus Miktoniscus patiencei Trichoniscoides albidus Trichoniscoides saeroeensis Halophiloscia couchi Armadillidium pulchellum Eluma purpurascens
RDBK Nb Nb Nb Nb Nb Nb
Grasshoppers, crickets, earwigs and cockroaches (Orthoptera/Dermaptera/Dictyoptera) Platycleis albopunctata Nb Tetrix ceperoi Na Gomphocerippus rufus Nb Ectobius lapponicus Nb Ectobius pallidus Nb Ectobius panzeri Nb *Tettigonia viridissima *Should be considered worthy of conservation effort, and this species, together with Platycleis albopunctata, is considered as part of the important invertebrate fauna to be found on sea cliff habitat in Devon
True Bugs (Hemiptera) Heteroptera Pyrrhocoris apterus Ischnodemus quadratus Macroplax preyssleri Pterotmetus staphyliniforms Trapezonotus ullrichi Piesma quadratus spergularia Lasicantha capucina Sciocoris cursitans Dicranocephalus agilis Heterogaster artemisiae Lasiosomus enervis Megalonotus praetextatus Rhyparachromus pini Tropistethus holosericeus Berytinus hirticornis Oncochila simplex
RDB1 RDB1 RDB3 RDB3 RDB3 RDB3 RDB3 Nb Nb Nb Nb Nb Nb Nb Nb Na
Myrmedobia coleoptrata Capsodes sulcatus Globiceps fulvicollis cruciatus Stronylocoris luridus Pachycoleus waltli Saldula arenicola Hebrus pusillus Enoplops scapha Corizus hyosyami Henestrias laticeps Beosus maritimus Kalama tricornis
Nb Nb Nb Nb Nb Na Cliff, exposed raw clay Nb Local Local Local Local Local
Leafhoppers, planthoppers, froghoppers, treehoppers & cicadas (Auchenorrhyncha) Ulopa trivia Nb Tettigometra impressopunctata Nb Flies (Diptera) Snail-killing flies, picture-wing flies, grass flies and allies (Acalyptrata) Platycephala umbraculata (Chloropidae) RDB2 Hoverflies (Syrphidae) Chrysotoxum elegans Eumerus sabulonum
RDB3
Volucella zonaria and V. inanis seem to utilise Paravespula colonies along the south coast cliffs. P. germanica is often very abundant at such ‘hot’ sites.
Soldier flies, bee flies, robber flies & allies (Larger Brachycera) Neomochtherus pallipes [status not known] Thereva strigata RDB3 Bombylius discolor N Oxycera morrisii N Oxycera pardalina N Oxycera pygmaea N Spania nigra N Stratiomys potamida N Vanoyia tenuicornis N Craneflies (Tipulidae) Symplecta scotica Arctoconopa melampodia Helius hispanicus Dicranomyia lackschewitzianus Dicranomyia goritiensis
RDB1 RDB2 RDBK RDBK RDB3
Blowflies, dung flies and allies (Calyptrata) Bithia modesta (Tachinidae) Eloceria delecta (Tachinidae) Macronychia griseola (Sarcophagidae) Sarcophila latifrons (Sarcophagidae)
RDB2 N RDB3 N
Orchisia costata (Muscidae) Coenosia atra (Muscidae) Lispocephala falculata (Muscidae) Beetles (Coleoptera) Ground beetles (Carabidae) Cicindela germanica Lionychus quadrillum Bembidion nigropiceum Species Statement Tachys micros Species Statement Chlaenius nitidulus Drypta dentata Harpalus honestus Trechus subnotatus Dyschirius obscurus Cymindis axillaris Nebria livida Amara praetermissa Asaphidion pallipes Bembidion saxatile Harpalus serripes Tachys bistriatus
RDB2 N N
BAP Priority RDB3 BAP Priority RDB3 Na Na RDB1 RDB1 RDB1 RDB1 RDB2 Na Na Nb Nb Nb Nb Nb
Leaf beetles (Chrysomelidae) Psylliodes luridipennis Longitarsus aeruginosus Calomicrus circumfusus Cassida hemisphaerica Chrysolina haemoptera Longitarsus absynthii Longitarsus anchusae Longitarsus brunneus Longitarsus dorsalis Longitarsus fowleri Longitarsus nasturtii Longitarsus nigrofasciatus Longitarsus plantagomaritimus
BAP Priority RDB 2 Endemic RDB1 Notable A Notable A Notable B Notable A Notable B Notable B Notable B Notable A Notable B Notable A Notable
Weevils (Curculionoidea) Pachytychius haematocephalus Cathormiocerus britannicus Cleopomiarus micros Smicronyx caecus Smicronyx reichi Tychius crassirostris Baris analis Ceutorhynchus insularis Otiorhynchus ligustici Barypeithes pyrenaeus Sitona gemellatus Cathormiocerus attaphilus
BAP Priority BAP Priority RDB3 RDB3 RDB3 RDB IK RDB2 RDB1 RDB2 RDB Ind RDB1 RDB1
Cathormiocerus maritimus Cathormiocerus myrmecophilus Cathormiocerus socius Hypera ononidis Hypera pastinacae Anchonidium unguiculare Aizobius sedi Helianthemapion aciculare * Stenopterapion intermedium Diplapion stolidum Cyanapion gyllenhali Catapion curtisi Protapion laevicolle Protapion schoenherri Grypus equiseti Smicronyx jungermanniae Orthochaetes insignis Pseudostyphlus pillumus Sibinia sodalis Tychius breviusculus * Tychius pusillus Tychius schneideri Tychius squamulatus Baris laticollis Calosirus terminatus Ceutorhynchus resedae Trichosirocalus dawsoni Trichosirocalus horridus Zacladus exiguus Mononychus punctumalbum Barypeithes sulcifrons Sitona ononidis Sitona waterhousei Tanymecus palliatus Trachyphloeus alternans Trachyphloeus aristatus Trachyphloeus asperatus Trachyphloeus bifoveolatus * Trachyphloeus digitalis Trachyphloeus rectus Trachyphloeus spinimanus Larinus planus Rhinocyllus conicus
RDB3 RDB3 RDB2 RDB IK RDB1 RDB2 Nb Na Nb Nb Na Na Na Nb Nb Nb Na ** Na Nb Nb Nb Na Nb Nb Nb Na Nb Na Nb Nb Nb Nb Nb Nb Nb Na Na Nb Nb Na
* Species recently recognised or added to the British List, but which merit at least Notable status. ** This species merits RDB status in the opinion of many specialists.
Rove beetles and allies (Staphylinidae/Scydmaenidae/Silphidae) Stenichnus poweri RDBK Silpha obscura RDB2
Bledius crassicollis Bledius dissimilis Bledius filipes Bledius occidentalis Thinobius brevipennis Medon pocoferus Scopaeus laevigatus Scopaeus minutus Astenus procerus Astenus subditus Stenichnus pusillus Lesteva hanseni Carpelimus halophilus Medon ripicola Astenus immaculatus Erichsonius signaticornis Staphylinus pedator Mycetoporus piceolus Tachyporus formosus Lamprinodes saginatus Claviger testaceus Bledius fergussoni Bledius limicola Bledius longulus Bledius praetermissus Stenus canaliculatus Stenus guttula Stenus guynemeri Stenus nitidiusculus Stenus picipennis Dianous coerulescens Lathrobium multipunctum
RDBI RDBI RDB1 RDBK RDBK RDBI RDBI RDB1 RDBK RDBI N N N N N Nb Na N Na Na N Local Local Local Local Local Local Local Local Local Local Local
Click beetles (Elateridae) Anostirus castaneus Cardiophorus vestigialis
RDB1 RDB2
Dung beetles and chafers (Scarabaeidae) Aphodius subterraneus Psammodius porcicollis Aphodius sordidus Amphimallon ochraceus Cetonia aurata
RDB1 Extinct Na Na Local
Skin beetles (Trogidae) Trox perlatus
RDB1
Jewel beetles (Buprestidae) Aphanisticus pusillus
Nb
Earth-boring dung beetles (Geotrupidae)
Geotrupes vernalis Long-horned beetles (Cerambycidae) Anoplodera livida Caddis flies (Trichoptera) Adicella filicornis Ernodes articularis Tinodes unicolor Hydroptila valesiaca Plectrocnemia brevis Ants, bees and wasps (Hymenoptera: Aculeata) Anergates atratulus Lasioglossum angusticeps Osmia xanthomelana Nomada errans Bombus humilis Stromgylognathus testaceus Euodynerus quadrifasciatus Mimumesa atratina Mimumesa littoralis Nysson interruptus Colletes hederae Hylaeus spilotus Andrena alfkenella Andrena niveata Andrena proxima Andrena rosae Andrena simillima Lasioglossum laticeps Sphecodes niger Sphecodes spinulosus Nomada conjungens Nomada fulvicornis Nomada guttulata Nomada hirtipes Nomada sexfasciata Anthophora retusa Chrysis viridula Pseudospinola neglecta Tiphia femorata Methocha articulata Monosapyga clavicornis Temnothorax albipennis Ponera coarctata Cryptocheilus notatus Priocnemis agilis Priocnemis gracilis Arachnospila minutula Aporus unicolor
Nb
Local
RDB3
BAP Priority BAP Priority BAP Priority BAP Priority BAP Priority RDB3 RDB3, RDB2 RDB2 RDB3 RDB3, RDB2 Very rare RDB3 RDB3 RDB2 RDB3, RDB3 RDB3, RDB2 RDB3, RDB2 RDB2 RDB3 RDB3, RDB2 RDB3, RDB2 RDB3 RDB1 RDB3 RDB1 RDB3, RDB1 Scarce Scarce Scarce Nb, Scarce Nb, Scarce RDB3, Na, Scarce Nb, Rare RDB3, RDB2, Rare Nb, Scarce RDB3, Nb, Scarce Nb, Scarce Na, Rare
Odynerus melanocephalus Crossocerus palmipes Mimumesa unicolor Podalonia hirsuta Nysson dimidiatus Didineis lunicornis Gorytes laticinctus Argogorytes fargeii Cerceris ruficornis Andrena bimaculata Andrena bucephala Andrena fulvago Andrena hattorfiana Andrena humilis Andrena labiata Andrena nigriceps Andrena nitidiuscula Andrena pilipes Andrena trimmerana Lasioglossum puncticolle Lasioglossum xanthropus Sphecodes crassus Sphecodes reticulatus Sphecodes rubicundus Stelis ornatula Coelioxys concoidea Nomada flavopicta Nomada integra Eucera longicornis Anthophora quadrimaculata
Na, Rare Nb, Scarce RDB3, Na, Rare Nb, Scarce Nb, Scarce RDB3, Na, Scarce RDB3, Rare RDB3, Na, Rare Scarce Nb, Scarce RDB3, Na, Rare RDB3, Na, Rare RDB2, RDB3, Scarce Nb, Scarce RDB3, Na, Scarce Nb, Scarce RDB3, Scarce Nb, Scarce Nb, Scarce Nb, Scarce Nb, Scarce Nb, Scarce RDB3, Na, Scarce Na, Rare RDB3, Rare Scarce Nb, Scarce Na, Rare Na, Rare Nb, Scarce