Buglife - Notable invertebrates associated with lowland heathland

Notable invertebrates associated with lowland heathland Snails (Mollusca) Lymnaea glabra

RDB 2

Spiders and allies (Arachnida: Araneae and Pseudoscorpiones) Eresus sandaliatus BAP Priority RDB1 Uloborus walckenaerius BAP Priority RDB3 Alopecosa fabrilis RDB1 Altella lucida RDB1 Centromerus brevivulvatus RDB3 Cheiracanthium pennyi RDB2 Dipoena coracina RDB1 Dipoena erythropus RDB2 Dipoena melanogaster RDB2 Enoplognatha oelandica RDB3 Ero aphana RDB2 Gonatium paradoxum RDB2 Haplodrassus umbratilis RDB3 Lathys stigmatisata RDB3 Oxyopes heterophthalmus RDB2 Thanatus formicinus RDB2 Theridion pinastri RDBK Xysticus luctator RDB2 Zora armillata RDB3 Zora silvestris RDB2 Acartauchenius scurrilis Na Achaearanea riparia Nb Aelurillus v-insignitis Nb Agraecina striata Nb Agroeca cuprea Na Anelosimus aulicus Nb Araneus angulatus Nb Araniella displicata Na Bianor aurocinctus Na Coelotes terrestris Nb Crustulina sticta Nb Dipoena inornata Nb Dipoena prona Nb Dipoena tristis Na Drassyllus praeficus Nb Entelecara congenera Nb Episinus truncates Nb Ero tuberculata Nb Evarcha arcuata Nb Glyphesis cottonae Na Gnaphosa lugubris Na Haplodrassus dalmatensis Nb Hybocoptus decollatus Nb Hypsosinga sanguinea Nb Lathys nielseni Na Maro sublestus Nb Mecopisthes peusi Nb Micaria silesiaca Nb

Micaria subopaca Micrargus laudatus Neriene furtive Notioscopus sarcinatus Ozyptila scabricula Phaeocedus braccatus Philodromus collinus Philodromus emarginatus Scotina palliardi Sitticus saltator Steatoda albomaculata Talavera petrensis Tapinocyba mitis Theridiosoma gemmosum Thomisus onustus Trachyzelotes pedestris Trematocephalus cristatus Trichoncus saxicola Walckenaeria corniculans Walckenaeria incisa Wiehlea calcarifera Xerolycosa nemoralis Xysticus acerbus Xysticus robustus Zelotes longipes Zelotes petrensis Zilla diodia Zora nemoralis

Nb Nb Nb Nb Nb Nb Nb Nb Na Nb Nb Nb Nb Nb Nb Nb Na Nb Na Nb Na Nb Na Na Na Na Nb Nb

Dragonflies and damselflies (Odonata) Coenagrion mercuriale Leucorrhinia dubia Ischnura pumilio Orthetrum coerulescens Ceriagrion tenellum

BAP Priority N Local Local Local

True bugs (Hemiptera) Heteroptera Eysarcoris aeneus Rhopalus rufus Nysius helveticus Pachybrachius luridus Pygolampis bidentata Nabis brevis Amblytylus delicatus Myrmecoris gracilis Phytocoris insignis Legnotus picipes Odontoscelis lineola Spathocera dahlmanni Rhopalus maculatus Drymus pilipes Emblethis denticollis Graptopeltus lynceus Lasiosomus enervis

RDB3 RDB3 RDB3 RDB3 RDBK RDB3 RDB3 RDB3 RDB3 Nb Nb Na Nb Na pNb Nb Nb

Megalonotus dilatatus Megalonotus praetextatus Rhyparachromus pini Nabis pseudoferus Adelphocoris ticinensis Globiceps flavomaculatus Globiceps fulvicollis cruciatus Globiceps juniperi Monosynamma sabulicola Systellonotus triguttatus Tytthus geminus Pachycoleus waltli Micracanthia marginalis Hebrus pusillus Microvelia pygmaea Alydus calcaratus Coranus subapterus Eremocoris podagricus Ischnocoris angustulus Lamproplax picea Pachybrachius fracticollis Berytinus crassipes Berytinus montivagus Hallodapus rufescens Megacoelum beckeri Chartoscirta cocksi Gerris (Aquarius) najas Sigara semistriata

Nb Nb Nb Nb Nb Nb Nb Nb Nb Nb Nb Nb Na Nb Nb Local Local Local Local Local Local Local Local Local Local Local Local Local

Leafhoppers, planthoppers, froghoppers, treehoppers & cicadas (Auchenorrhyncha) Aphrophora alpina Nb Scleroracus decumanus Nb Zygina rubrovittata Nb Grasshoppers, crickets, earwigs and cockroaches (Orthoptera/Dermaptera/Dictyoptera) Decticus verrucivorus Stethophyma grossum Chorthippus vagans *Platycleis albopunctata Metrioptera brachyptera Nemobius sylvestris Tetrix ceperoi Omocestus rufipes Ectobius lapponicus Ectobius pallidus Ectobius panzeri

BAP Priority RDB2 BAP Priority RDB2 RDB3 Nb *Coastal heath only Nb Na Na Nb Nb Nb Nb

Flies (Diptera) Snail-killing flies, picure-wing flies, grass flies and allies (Acalyptrata) Odinia ornata (Odiniidae) RDB2 Pherbellia knutsoni (Sciomyzidae) RDB3 Themira biloba (Sepsidae) RDBI Urophora spoliata (Tephritidae) RDB3 Chyliza vittata (Psilidae) N

Geomyza subnigra Opomyza lineatopunctata Opomyza punctata Micropeza lateralis Psilopa marginella Rhopalopterum femorale Trachysiphonella ruficeps Trixoscelis marginella

(Opomyzidae) (Opomyzidae) (Opomyzidae) (Micropezidae) (Ephydridae) (Chloropidae) (Chloropidae) (Heleomyzidae)

N N N N N N N N

Soldier flies, bee flies, robber flies & allies (Larger Brachycera) Asilus crabroniformis BAP Priority Bombylius minor BAP Priority Thyridanthrax fenestratus BAP Priority Machimus arthriticus RDB1 Villa venusta RDB2 Eutolmus rufibarbis RDB3 Lasiopogon cinctus Priority Ogcodes gibbosus Priority Ogcodes pallipes Priority Hoverflies (Syrphidae) Chrysotoxum octomaculatum Pelecocera tricincta Chrysotoxum vernale Cheilosia mutabilis Eupeodes bucculatus Microdon analis Microdon myrmicae Orthonevra geniculata Paragus tibialis Sphaerophoria virgata

RDB2 RDB3 RDB1 N N N N N N N

Didea intermedia may occur where Pinus is invading. Callicera aurata and Psilota anthracina may occur where old trees are present within heathland. Chrysotoxum elegans can be found on breck ‘chalk-heath’. Triglyphus primus can occur where Artemisia vulgaris is present on heath edges. Volucella inanis occurs on some heaths in the home counties where good Paravespula populations are present; (V. zonaria may also occur too). Xylota tarda may occur where there are stands of Populus tremula. Platycheirus discimanus occurs especially at heathland-woodland interfaces.

Eupeodes nitens may also occur on heathland.

Craneflies (Tipulidae) Nephrotoma sullingtonensis Nephrotoma crocata Prionocera pubescens Tanyptera nigricornis Tipula marginella Tanyptera atrata Tipula livida Tipula holoptera Tipula yerburyi Erioptera nielseni Phylidorea abdominalis Orimarga juvenilis Phalacrocera replicata

RDB1 RDB3 RDB2 RDB3 RDB3 N N N N N N N N

Nephrotoma scurra Tipula cava Tipula helvola Limonia dilutior Dicranomyia affinis Ptychoptera minuta

N (demoted from Notable)

Blowflies, dung flies, flesh flies and allies (Calyptrata) Blaesoxipha rossica (Sarcophagidae) Coenosia vibrissata (Muscidae) Cordilura picipes (Scathophagidae) Cylindromyia brassicaria (Tachinidae) Erynnia ocyptera (Tachinidae) Exorista glossatorum (Tachinidae) Freraea gagatea (Tachinidae) Gasterophilus nasalis (Gasterophilidae) Gasterophilus pecorum (Gasterophilidae) Gonia divisa (Tachinidae) Gymnosoma rotundatum (Tachinidae) Helina crinita (Muscidae) Helina tetrastigma (Muscidae) Hydrotaea glabricula (Muscidae) Hypoderma bovis (Oestridae) Hypoderma lineatum (Oestridae) Hippobosca equina (Hippoboacidae) Macronychia griseola (Sarcophagidae) Miltogramma germari (Sarcophagidae) Oestrus ovis (Oestridae) Orchisia costata (Muscidae) Phaonia jaroschewskii (Muscidae) Subclytia rotundiventris (Tachinidae) Blaesoxipha gladiatrix (Sarcophagidae) Coenosia atra (Muscidae) Eurithia intermedia (Tachinidae) Gasterophilus intestinalis (Gasterophilidae) Pterella grisea (Sarcophagidae) Sarcophila latifrons (Sarcophagidae) Scathophaga scybalaria (Scathophagidae)

RDB3 RDB3 RDB3 RDB1 RDB2 RDB2 RDB3 RDB1 RDB1 RDB3 RDB3 RDB2 RDB3 RDB3 RDB2 RDB2 RDB K RDB3 RDB3 RDB3 RDB2 RDB2 RDB3 N N N N N N N

Beetles (Coleoptera Ground beetles (Carabidae) Poecilus kugelanni Tachys edmondsi Amara famelica Cicindela sylvatica Anisodactylus nemorivagus Species Statement Cymindis macularis Elaphropus walkerianus Sericoda quadripunctata Agonum sexpunctatum Amara infima Cymindis axillaris Bembidion nigricorne Carabus nitens Notiophilus quadripunctatus

BAP Priority RDB1 BAP Priority RDB1 BAP Priority RDB3 BAP Priority Na Na RDB1 RDB1 RDB1 Na Na Na Nb Nb Nb

Poecilus lepidus Pterostichus quadrifoveolatus

Nb Nb

Leaf beetles (Chrysomelidae) Cryptocephalus coryli Cryptocephalus decemmaculatus Cryptocephalus biguttatus Altica ericeti Calomicrus circumfusus Cassida nebulosa Chaetocnema aerosa Chaetocnema subcoerulea Longitarsus agilis Luperus flavipes Mantura chrysanthemi Orsodacne lineola

BAP Priority RDB 1 BAP Priority RDB 2 RDB 2 Notable B Notable A RDB K RDB K Notable B Notable A Notable B Notable A Notable B

Weevils (Curculionoidea) Procas granulicollis Apion rubiginosum Dieckmanniellus gracilis Orchestes calceatus Bagous brevis Bagous collignensis Bagous czwalinae Bagous frit Pelonomus olssoni Strophosoma fulvicorne Coniocleonus nebulosus Perapion affine Exapion genistae Sibinia pyrrhodactyla Tychius parallelus Bagous limosus Bagous lutulosus Pelonomus canaliculatus Caenopsis fissirostris

BAP Priority (RDB Ind) RDB3 dry/open RDB3 wet heath ?* scrub RDB1 wet heath RDB3 ponds RDB1 ponds RDB3 wet heath/ponds RDB3 wet heath RDB2/3dry heath RDB3 dry heath Na dry heath Na dry/humid Nb open Na scrub Nb ponds Na wet/open Nb ponds Nb dry/humid

Rove beetles and allies (Staphylinidae/Scydmaenidae/Silphidae) Scaphisoma assimile RDBI Oxytelus piceus RDBK Stenus subdepressus RDBI Paederus caligatus RDB3 Staphylinus caesareus RDBI *Erichsonius ytenensis RDBI *No verified post-1940 records are available for this species, and it may deserve a higher conservation status

Aclypea opaca Scaphisoma boleti Bledius femoralis Staphylinus fortunatarum Staphylinus ophthalmicus Mycetoporus punctus Bledius gallicus Bledius opacus

Na Nb Na Nb Na N Local Local

Stenus geniculatus Staphylinus erythropterus Staphylinus aeneocephalus Staphylinus brunnipes Mycetoporus clavicornis Bolitobius cingulatus Tachyporus atriceps Ocalea badia

Local Local Local Local Local Local Local Local

Jewel beetles (Buprestidae) Aphanisticus emarginatus Melanophila acuminata

RDB1 Local

Dung beetles and chafers (Scarabaeidae) Aphodius consputus Onthophagus fracticornis Aphodius paykulli Aphodius distinctus Cetonia cuprea

RDB3 RDBK Nb Nb Nb

Click beetles (Elateridae) Ampedus sanguinolentus Sericus brunneus

Na Local

Earth-boring dung beetles (Geotrupidae) Geotrupes vernalis Odontaeus armiger Geotrupes pyrenaeus

Nb Na Local

Skin beetles (Trogidae) Trox sabulosus

Na

Caddis flies (Trichoptera) Hagenella clathrata ?Limnephilus pati ?Limnephilus tauricus Limnephilus elegans Oligotricha striata

RDB1 RDB1 RDB1 Local & Regionally notable Regionally notable

Caddis associated with wet woodland, ditches and streams, and large, often artificial, ponds within the lowland heath are not listed. If there are any uncommon species associated with those habitats they will have been picked up elsewhere with the relevant habitat.

Ants, bees and wasps (Hymenoptera: Aculeata) Chrysis fulgida Anergates atratulus Formica candida Formica exsecta Formica rufibarbis Homonotus sanguinolentus Psedepipona herrichii Cerceris quinquefasciata Andrena ferox

BAP Priority BAP Priority BAP Priority BAP Priority BAP Priority BAP Priority BAP Priority BAP Priority BAP Priority

Andrena gravida Nomada ferruginata Bombus sylvarum Chrysis gracillima Myrmica hirsuta Myrmica karavajevi Temnothorax interruptus Stromgylognathus testaceus Ceropales variegata Symmorphus connexus Symmorphus crassicornis Tachysphex unicolor Crossocerus vagabundus Ectemnius borealis Mimumesa spooneri Mimesa bicolor Passaloecus turionum Nysson interruptus Hylaeus gibbus Andrena alfkenella Andrena niveata Andrena proxima Andrena rosae Andrena ruficrus Halictus confusus Lasioglossum pauperatum Lasioglossum semilucens Lasioglossum sexnotatum Sphecodes scabricollis Stelis breviscula Heriades truncorum Coelioxys quadridentata Nomada fulvicornis Nomada guttulata Nomada hirtipes Nomada roberjeotiana Nomada signata Anthophora retusa Cleptes nitidulus Cleptes semiauratus Hedychridium coriaceum Hedychridium cupreum Hedychridium roseum Hedychrum niemelai Chrysis illigeri Chrysis viridula Chrysura radians Pseudospinola neglecta Tiphia femorata Methocha articulata Mutilla europaea Smicromyrme rufipes Monosapyga clavicornis Formica sanguinea Tapinoma erraticum/ambiguum

BAP Priority BAP Priority BAP Priority RDB2, RDB2 RDB3, RDBK, Very rare RDB3, RDBK, Very rare RDB3, RDB3 RDB3, RDB3 RDB1, RDB1 RDB3, RDB3 RDB3, RDB3 Very rare RDB1, RDB+, Very rare RDB3, RDB3 RDB3, RDB3 RDB2, RDB2 Very rare RDB3, RDB2 RDB3, RDB3 RDB3, RDB3 RDB2, RDB2 RDB3, RDB3 RDB3, RDB2 RDB3, RDB3 RDB3, RDB3 RDB3, RDB3 Not listed, RDB3 Appendix, RDB1 RDB3, RDB3 RDB1, RDBK, Very rare RDB3, RDBK, Very rare RDB3, RDB3 RDB3, RDB3 RDB1, RDB1 RDB3, RDB3 RDB3, RDB3 RDB3, RDB2 RDB3, RDB1 RDB3, Na, Rare Nb, Scarce RDB3, RDB3, Rare Nb, Scarce Scarce Not listed, RDB3, Scarce Nb, Scarce Scarce Na, Rare Scarce Scarce Nb, Scarce Nb, Scarce Nb, Scarce Nb, Scarce Nb, Scarce Nb, Scarce

Cryptocheilus notatus Priocnemis coriacea Priocnemis hyalinata Caliadurgus fasciatellus Anoplius viaticus Arachnospila minutula Arachnospila wesmaeli Evagetes dubius Aporus unicolor Ceropales maculata Eumenes coartctata Odynerus melanocephalus Tachysphex nitidus Crabro scutellatus Crossocerus leucostomus Crossocerus palmipes Oxybelus argentatus Oxybelus mandibularis Mimesa bruxellensis Psenulus schencki Pemphredon morio Diodontus insidiosus Diodontus tristis Ammophila pubescens Podalonia affinis Podalonia hirsuta Nysson dimidiatus Gorytes laticinctus Argogorytes fargeii Cerceris ruficornis Colletes marginatus Hylaeus signatus Andrena apicata Andrena argentata Andrena bimaculata Andrena bucephala Andrena falsifica Andrena florea Andrena humilis Andrena labiata Andrena marginata Andrena minutuloides Andrena nigrospina Andrena tibialis Andrena trimmerana Andrena varians Lasioglossum brevicorne Lasioglossum prasinum Lasioglossum quadrinotatum Sphecodes crassus Sphecodes longulus Sphecodes miniatus Sphecodes reticulatus Sphecodes rubicundus Dasypoda hirtipes

RDB3, RDB2, Rare Na, Rare Nb, Scarce Scarce Scarce Nb, Scarce RDB3, Na, Rare Nb, Rare Na, Rare Rare Na, Scarce Na, Rare Scarce Na, Scarce RDB3, Na, Rare Nb, Scarce Na, Scarce Na, Scarce Na, Rare RDB3, Na, Rare RDB3, Nb, Scarce RDB3, RDB3, Scarce Scarce Scarce RDB3, RDB3, Rare Nb, Scarce Nb, Scarce RDB3, RDB3, Rare RDB3, Na, Rare Scarce RDB3, Na, Scarce Nb, Scarce Nb, Scarce Na, Rare Nb, Scarce RDB3, Na, Rare Na, Rare RDB3, RDB3, Scarce Nb, Scarce RDB3, Na, Scarce Na, Scare Na, Rare Rare RDB3, Na, Scarce Nb, Scarce Nb, Scarce RDB3, RDB3, Scarce Scarce Na, Rare Nb, Scarce Na, Rare Nb, Scarce RDB3, Na, Scarce Na, Rare Nb, Scarce

Macropis europaea Stelis ornatula Megachile dorsalis Coelioxys concoidea Nomada argentata Nomada baccata Nomada flavopicta Nomada integra Eucera longicornis Anthophora quadrimaculata Ceratina cyanea

RDB3, Na, Scarce RDB3, RDB3, Rare Nb, Scarce Scarce Not listed, RDB3, Rare Na, Rare Nb, Scarce Na, Rare Na, Rare Nb, Scarce RDB3, RDB3, Rare