An Annotated Catalogue of the Biodiversity of the Doward, Ian Draycott

An annotated catalogue of the biodiversity of the Doward Ian Draycott – December 2023

Purpose This publication is intended to be a reference point for all available information regarding the biodiversity of the Doward in south Herefordshire. It brings together thousands of individual species observations made over a period of more than a century from many visiting naturalists and local residents who have put their knowledge into the public domain. It is intended to be both a source of checklists for all the species groups found on the Doward and also to provide some general context about the plant and animal communities of this part of the Lower Wye Valley. It is a more detailed companion to the booklet ‘The Biodiversity of the Doward and Symonds Yat’ published by Herefordshire Wildlife Trust in August 2023 which is available from them and from some local outlets.

Biodiversity information is dynamic and changes over time. The information here is intended to reflect these changes be regularly updated as new observations are made and our knowledge of each species group expands. Please feel free to make your own contributions by emailing idraycott@yahoo.co.uk.

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Structure Following an introduction, the information is presented in three parts. Each part is then divided into sections representing the main taxa groups as summarised in the table below. The page numbers in the table lead to a more detailed directory of each section.

The species checklists usually ordered by taxonomic family but the format of each can vary and, in some cases, gives information about the national or local status of each species and source of the record.

Introduction

Page 3

Site definition and description Biodiversity Summary Information sources

Part 1. Plant and Fungal kingdoms

Page 6.

1.1 Vascular Plants 1.2 Bryophytes - Mosses and Liverworts 1.3 Fungi 1.4 Lichens 1.5 Other Kingdoms

Part 2. Invertebrates

Page 29.

2.1. Class Insecta – The insects 2.2. Class Arachnida – Spiders and allies 2.3. Classes Gastropoda and Bivalva – Slugs, Snails and Mussels 2.4. Other invertebrates

Part 3. Vertebrates

Page 87.

3.1 Mammals 3.2 Birds 3.3 Reptiles and Amphibians 3.4 Fish

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Introduction Site definition and description The Doward Hills is the area of around 4 square kilometres bounded to the south and east by the River Wye and by the A40 road to the North and West. It sits at the most southerly point of Herefordshire in the parish of Whitchurch and Ganarew and is all in the Lower Wye Area of Outstanding Natural Beauty.

The underlying geology is primarily a mixture of Carboniferous Limestone and Devonian Sandstone and there are many areas showing signs of former quarrying activity. These limestone rocks give rise to calcareous grassland and beech woodland and the sandstones can create more acidic conditions suiting several vegetation types from ranging from acid heath to oak and conifer woodland.

The Doward is that it is not an agricultural landscape. Over 60% of the Doward is now woodland with much of Lord’s Wood managed as commercial forestry. The rest is various sorts of grassland pasture and private landholdings of gardens and scrub. Nevertheless, some patches of species-rich calcareous grassland, ancient woodland, acid heathland and riparian wetland can still be found.

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Biodiversity Summary The following summary is a list of what is special about the biodiversity on the Doward. A more detailed account is given in the following parts of this publication and scientific names are given in the checklists.

Plantlife.

There are records of around 1,000 plant species of which 784 are vascular plants and the rest mainly bryophtes.

Dicot plants Unusual or regionally scarce plants include Yellow Bird’s-nest, Hutchinsinia, Narrow-leaved Bittercress, Small Scabious, Dropwort, Wild Clary, Rest Harrow, Wild Madder, Deadly Nightshade, Slender St John’s-wort and Great Lettuce. There is also one micro-species of Hawkweed Hieracium subplanifolium found here that is endemic to the Doward.

Monocot plants The highlights within this group include at least 10 species of orchid including good populations of Greater Butterfly, Bird’snest Orchids, White Helleborine and also a few Pyramidal and Fly Orchids. Some nationally scarce limestone sedges exist including Fingered, Dwarf and Soft-leaved Sedges. Unusual grasses include Mountain Mellick, Blue Moor Grass, Wood Barley and Flattened Meadow-grass.

Trees and shrubs Within the hedgerows and shrub layer Purging Buckthorn, Wayfaring Tree, Spindle and Spurge Laurel can be found. The woodlands hold many notable veteran trees especially beech and have a significant population of both species of native Lime trees as well as Wild Service Trees.

Most notable of all are fourteen species of Sorbus trees (i.e., Whitebeams and allies) two of which, Sorbus herefordensis and S. greenii, are endemic to the Doward with several others are globally restricted to the Lower Wye Valley.

Non-flowering plants There are 17 ferns present in the area including Adder’s Tongue Fern and the rare pachyrachis form of Maidenhair Spleenwort. 258 bryophyte species are recorded including at least 8 that are classed as nationally scarce.

Fungi.

The full list of fungi for the 10km square exceeds 1,500 species (NBN) and includes national scarcities such Devils Bolete, Hollow Bolete, Terracotta Hedgehog, Giant Clubs, Fluted Bird’s-nest, Lacquered Bracket, Snakeskin Grissette, Earthstars, Earth Tongues and 9+ Waxcaps.

There are also records of several nationally notable lichen species.

Vertebrates.

Birds: 93+ species recorded. Regular breeding Wood Warbler*, Tree Pipit*, Lesser Spotted Woodpecker*, Marsh Tit*, Common Redstart, Goshawk, Peregrine with occasional breeding Pied flycatcher* and possibly Hawfinch* and Woodcock*. (*denotes Red Listed as Birds of Conservation Concern) Mammals: 34 species recorded. Notable for bats with Greater and Lesser Horseshoe Bats plus records of Western Barbastelle, Whiskered and Natterer’s Bats. A good population of Hazel Dormouse and occasional visiting Wild Boar and Otter. The Polecat is known to occur and there are reports of Pine Martens with radio-tracking collars passing through.

Reptiles and amphibians: 7 species recorded including old records of Great Crested Newt.

Fish: The River Wye and adjoining watercourses hold at least 25 species including Atlantic Salmon, Allis Shad and 3 species of Lamprey and is an important breeding ground for the nationally scarce Twaite Shad.

Invertebrates Over 2,100 invertebrate species recorded.

Lepidoptera – Grizzled Skipper, Wood White, White Admiral and possibly still Pearl-bordered Fritillary all occur. Over 968 moth species have been recorded of which 15 are listed as Nationally Scarce B. These include Great Oak Beauty, Ruddy Carpet, Bilberry Pug and Drab Looper (a Biodiversity Action Plan species). A long list of other species found here are currently regarded as nationally uncommon or local.

Other nationally scarce or rare insect species include several saproxylic beetles such as Cosnard’s Net-winged Beetle (Endangered RDB1) and the Welsh Longhorn Beetle, various woodland wasps such as Sabre Wasp and Giant Woodwasp, the Bee-wolf, Eared Leafhopper, Spurgebug, Club-tailed Dragonfly, Yellow Mayfly, several tufa specialist craneflies, Giant Fungus Gnat, several Bee-flies, Tachinid flies and the Southern Bristletail.

Other scarce invertebrates include Green Huntsman Spider, the Cave Spider, Lapidary and Wrinkled Snails and the Greater Pellucid Snail. There are two nationally rare woodlouse species.

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Information sources An important internet portal to access distribution data on all types of wildlife is the National Biodiversity Network in what is now known as the NBN Atlas (formerly NBN Gateway). This was originally set up by the Biological Records Centre (BRC) who are part of the Centre for Ecology and Hydrology (CEH) and are associated with the Natural Environment Research Council and the Joint Nature Conservation Committee (JNCC). They access species records mainly from a set of ‘National Recording Schemes’ which are run by experts and enthusiasts to collate the occurrence and distribution information of species belonging to a particular taxonomic group. By putting this ‘data’ online the public as well as professional ecologists can access detailed information about the wildlife in their local area. Similar data sources can also be found in iRecord and the Welsh system ADERYN but these can include unverified records which have to be filtered out. Local Records Centres such as the one in Hereford, also have their own systems but public access to data is more complicated.

In this publication I have used the Ordnance Survey geographical reference system which divides the UK into labelled kilometre squares. The Doward is the main part of the 10 km square labelled SO51 or the 2km square SO51M. More precisely it is comprised of parts of eight 1km squares of which SO5415, SO5414, SO5416, SO5515 are the most important. Being sited at the most southerly point of Herefordshire the information for the square SO51M also goes across the River Wye into Monmouthshire and covers part of Lady Park Wood National Nature Reserve. In this publication I have tried to exclude this as much as possible, but many older records do not have the spatial resolution to do this.

A download from the NBN Atlas in March’18 revealed a total of 26,827 separate records of all species in a search of a 2km radius from the centre of Great Doward *. These records were made up of 4,443 different species. A search of a wider area gave 5,468 species in a 5km radius, but this included some records from neighbouring counties across the river. These totals are significantly higher than any other site in Herefordshire including Moccas Park, Haugh Woods and the Malverns.

It also suggests a high level of biodiversity even by national standards.

Putting all this together gives a measure of two things. Firstly, the genuine species richness of a site and secondly the extent of species recording done by visiting expert naturalists who make their observations public. In conclusion, the biodiversity Doward has been surprising well-researched over the last 150 years and is undeniably a species-rich area. However, all these species totals should only be used as a guide or indication. *In this dataset the square SO51 had only 128 records marked as being in Wales and these were mainly bryophytes in Lady Park wood NNR. On a cautionary note, the NBN Atlas data has incorporated some very old databases and some of these were not very specific about location of observations, and it is not always clear whether it was on the Doward or elsewhere in SO51.

This publication is intended to share the information and lay down a benchmark to identify any future changes in species composition. Hopefully, it will also play a role in raising awareness of the richness of the biodiversity in this small area and explain the need to protect and enhance the special habitats on this ‘patch’.

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Part 1. The Plant and Fungal Kingdoms Overview.

According to data presented in the National Biodiversity Network (NBN) Atlas in 2017 at least 1,258 species of plants and 1,556 species of fungi and lichens have been recorded within 2km of the centre of Great Doward.

This total reflects the range of terrestrial macro and micro habitats arising from the combination of both Carboniferous limestone and Devonian sandstone underlaying the site. Most of the habitats are woodland or dry open areas of grassland and rock exposures but the small marshy areas around Dropping Wells and the seasonally flooded zone along the riverbank also adds to the plant diversity.

Looking at the NBN plant data in more detail gives 784 species of flowering plants (Angiosperms and Gymnosperms) and 26 ferns (Pteridophytes) and spore-forming allies, 344 species are bryophytes (mosses and liverworts) and a further 5 are species of green algae. The data on fungi includes lichens which are a symbiotic association between and a fungal thallus and an algae or blue-green bacteria.

It should be stressed that this dataset will not be a precise figure of what is actually present on the Doward today as a good number of these records are quite old or do not have adequate information about the location. Some records may not be on the Doward side of the river however, other species will be present but have yet to be found and recorded.

To simplify the production of checklists this publication uses the following main divisions:

Vascular Plants

Non-vascular Plants

Fungi

Angiosperms – Flowering Plants

Page 7.

Gymnosperms – Conifers and allies

Page 13.

Pteridophytes – Ferns and allies

Page 14.

Bryophytes: Mosses

Page 15.

Liverworts

Page 19.

Chlorophyta – Algae

Page 20.

Larger Fungi

Page 21.

Lichen

Page 26.

Supplementary Note: Endemic Plants of the Doward

Page 27.

These plant checklists cover species considered to be ‘native’ or ‘naturalised in UK. They excluded a range of species deliberately planted in gardens, horticulture and forestry estates. They also do not include details of some aggregates of species where morphological differences are minor and hard to discern in the field. As a consequence, these lists and their species totals do not fully match all species listed in the larger NBN or BSBI databases where greater expert interpretation has been required.

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1.1 Checklist of Vascular Plants The list below is generated from a range of ‘informal’ observations by various recorders on the Doward rather than a set of formal records. Although each species listed has been reported on the Doward not all have been verified so should be treated as being for guidance purposes only.

Angiosperms - Flowering Plants Ranunclulaceae Creeping Buttercup Meadow Buttercup Lesser Celandine Goldilocks Buttercup Wood anemone Columbine Travellers Joy Stinking Hellebore Papaveraceae Opium Poppy Greater Celandine Fumiariaceae Yellow Corydalis Common Fumitory Ulmaceae Wych Elm Viscacae Mistletoe Urticaceae Common nettle Pellitory-of-the-Wall Fagaceae Sessile Oak Sweet Chestnut Beech Betulaceae Silver/Downy Birch Alder Grey Alder Hazel Chenopodiaceae Fat-hen Caryophyllaceae Red Campion White campion Bladder Campion Common Chickweed Greater Stichwort Common Mouse-ear Thyme-leaved Sandwort Annual Pearlwort Knotted/procumbent pearlwort Polygonacea Knotgrass Redshank Water Pepper Common Sorrel Broad-leaved dock Curled Dock Japanese knotweed Black Bindweed

Ranunculus repens Ranunculus acris Ranunclus ficaria Ranunclulus auricomus Anemone nemorosa Aquilega vulgaris Clematis vitalba Heleborus viridis Papavera somniferum Chelidonium majus Pseudofumaria lutea Fumaria officinalis Ulmus glabra Viscum album Urtica dioica Parietaria judaica Quercus petraea Castanea sativa Fagus sylvatica Betula pendula/pubescens Alnus glutinosa Alnus incana Corylus avellana Chenopodium album Silene dioca Silene latifolia Silene vulgaris Stellaria media Stellaria holostea Cerastium fontanum ag Arenaria serpyllifolia Sagina apetala Sagina nodosa/procumens Polygonum aviculare Percicaria maculosa Percicaria hydropiper Rumex acetosa Rumex obtusifolius Rumex crispus Fallopia japonica Fallopia convolvulus

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Clusiaceae Hairy St Johns-wort Tutsan Perforate St John's-wort Slender St John's-wort Pale St John's-wort Square-stalked St John's-wort Malvaceae Musk Mallow Common Mallow Tiliaceae Large-leaved Lime Small-leaved Lime Cistaceae Common Rock-rose Salicaceae White Poplar Sallow/Grey Willow Creeping Willow Violaceae Common Dog-violet Early Dog-violet Hairy Violet Sweet Violet Brassicaceae Hairy Bittercress Wavy Bittercress Narrow-leaved Bittercress Lady's smock Common Whitlowgrass Hutchinsia Thale Cress Treacle Mustard Swine-cress Hairy Rock-cress Honesty Shepard's Purse Hedge Mustard Garlic Mustard Charlock Rape Creeping Yellow-cress Resedaceae Wild mignonette Weld Dipsacaceae Wild Teasel Small Teasel Field Scabious Small Scabious Devil’s-bit Scabious Astercea Daisy Ox-eye Daisy Blue Fleabane Scentless Mayweed Common Fleabane Common Ragwort Hoary Ragwort Groundsel Ploughman's-spikenard Yarrow Dandelion

Hypericum hirsutum Hypericum androsaemum Hypericum perforatum Hypericum pulchrum Hypericum montanum Hypericum tetrapterum Malva moschata Malva sylvestris Tilia platyphyllos Tilia cordata Helianthemum apenninum Populus alba Salix cinerea Salix repens Viola riviana Viola riechenbachiana Viola hirta Viola odorata Cardamine hirsuta Cardamine flexuosa Cardamine impatiens Cardamine pratensis Erophila verna Hornungia petraea Arabidopsis thaliana Erysimum cheiranthoides Coronopus squamatus Arabis hirsuta Lunaria annua Capsella bursa-pastoris Sisimbryum officinale Alliaria petiolata Sinapsis arvensis Brassica napus Rorippa sylvestris Reseda lutea Reseda luteola Dipsacus fullonum Dipsacus pilosus Knautia arvensis Scabiosa columbaria Succisa pratensis Bellis perrenis Leucanthemum vulgare Erigeron acer Tripleurospermum inodorum Pulicaria dysenterica Senecio jacobaea Senecio eruciformis Senecio vulgaris Inula conyzae Achillea millefolium Taraxacum officinale

8

Canadian Goldenrod Goldenrod Pineappleweed Hemp Agrimony Mugwort Greater Burdock Nipplewort Mouse-ear Hawkweed Goat's-beard Hawkweed Oxtongue Creeping Thistle Spear Thistle Marsh Thistle Common Knapweed Cat's-ear Rough Hawksbit Autumn Hawkbit Smooth Hawks-beard ag Hawkweed ag.

Prickly Sow-thistle Smooth Sow-thistle Wall Lettuce Great Lettuce Coltsfoot Plantaginaceae Ribwort Plantain Greater Plantain Crassulacea Wall Pennywort or Navelwort English/Biting Stonecrop Reflexed Stonecrop Gentianaceae Yellowwort Common Centuary Saxifragaceae Opposite-leaved Golden Saxifrage Rosaceae Bramble Field rose Dog rose Agrimony Fragrant Agrimony Salad Burnet Meadowsweet Dropwort Wood Avens Silverweed Creeping Cinquefoil Trailing Tormentil Barren Strawberry Wild Strawberry Parsley-piert Lady's Mantle Wild Cherry Blackthorn Crab Apple Wild Service Tree Whitebeam group Rowan Cotoneaster Fabeacea Tufted Vetch Bush Vetch

Solidago canadensis Solidago virguarea Matricaria discoides Eupatorium cannabinum Artemesia vulgaris Arctium lappa Lapsana communis Philosella officinarum Tragopogon pratensis Picris hieracoides Cirsium arvense Cirsium vulgare Cirsium palustre Centaurea nigra Hypochaeris radicata Leontodon hispidus Leontodon autumnalis Crepis spp Hieraceum spp Sonchus asper Sonchus oleraceus Mycelis muralis Lactuca virosa Tussilago farfara Plantago lanceolata Plantago major Umbilicus rupestris Sedum anglicum/acre Sedum rupestre Blackstonia perfoliata Centaurium erythraea Chryosplenium alternifolium Rubus fruticosus agg.

Rosa arvensis Rosa canina agg.

Agrimonia eupatoria Agrimonia procera Sanguisorba officinalis Filipendula ulmaria Filipendula vulgaris Geum urbanum Potentilla anserina Potentilla reptens Potentilla anglica Potentilla sterilis Fragaria vesca Aphanes arvensis Alchemilla sp agg.

Prunus avium Prunus spinosa Malus sylvestris Sorbus torminalis Sorbus aria ag see text Sorbus aucuparia Cotoneaster agg.

Vicia cracca Vicia sepium

9

Hairy tare Common Vetch Wood Bitter-vetch Kidney Vetch Bitter Vetch(ling) Meadow Vetchling Spotted Medick Black Medick Red Clover Zigzag Clover White Clover Lesser Trefoil Hop Trefoil Common Birds-foot-trefoil Greater Bird's-foot-trefoil Common Restharrow Horseshoe Vetch Broom Western Gorse Geraniaceae Bloody Cranesbill Herb-Robert French Crane's-bill Cut-leaved Cranesbill Linaceae Fairy Flax Euphoriaceae Dog's Mercury Wood Spurge Petty Spurge Polygalaceae Common Milkwort Thymelaeaceae Spurge Laurel Aceracea Field Maple Sycamore Oxalidaceae Wood Sorrel Cucurbitacea White Bryony Onagraceae Enchanters Nightshade Large-flowered Evening primrose Rosebay Willowherb Great Willowherb Broad-leaved Willowherb Hoary Willowherb Cornaceae Dogwood Viscaceae Mistletoe Celastraceae Spindle Aquifoliaceae Holly Apiaceae Upright Hedge Parsley Cow Parsley Wild Angelica Burnet-saxifrage Wild Carrot Hogweed

Vicia tetrasperma Vicia sativa Vicia orobus Anthyllis vulneraria Lathyrus linifolius Lathyrus pratensis Medicago arabica Medicogo lupulina Trifolium pratense Trifolium medium Trifolium repens Trifolium dubium Trifolium campestre Lotus corniculatus Lotus pedunculatus Ononis repens Hippocrepis camosa Cytisus scoparius Ulex gallii Geranium sanguineum Geranium robertianum Geranium endressi Geranium dissectum Linium catharticum Mercurialis annua Euphorbia amygdaloides Euphorbia peplus Polygala vulgaris Daphne laurelo Acer campestre Acer pseudoplatanus Oxalis acetosella Bryonia dioica Circaea lutetiana Oenothera erythrosepala Chamerion angustifolium Epilobium hirsutum Epilobium montanum Epilobium parviflorum Cornus sanguinea Viscum album Euonymus europeaus Ilex aquifolium Torilis japonica Anthriscus sylvestris Angelica sylvestris Pimpinrella saxifraga Daucus carota Heracleum sphondylium

10

Fool's Parsley Sanicle Pignut Ericaceae Ling Heather Bilberry Primulaceae Primrose Cowslip Yellow Pimpernel Scarlet Pimpernel Apocyanaceae Greater Periwinkle Rubiaceae Field Madder Wild Madder Lady's Bedstraw Hedge bedstraw Cleavers Woodruff Crosswort Convolvulaceae Hedge bindweed Field Bindweed Boraginaceae Common Gromwell Common Comfrey Green Alkanet Field Forgetmenot Lamiaceae White Deadnettle Red Deadnettle Common Hemp-nettle Bugle Yellow Archangel Betony Wild Clary Hedge Woundwort Marsh Woundwort Ground ivy Selfheal Wild Basil Common Calamint Wild Marjoram Thyme Wood Sage Gypsywort Solanaceae Bittersweet Deadly Nightshade Scrophulariaceae Wall Speedwell Heath Speedwell Germander Speedwell Common Field speedwell Thyme-leaved Speedwell Lousewort Great Mullein Common Figwort Ivy-leaved Toadflax Common Cow-wheat Red Bartsia Foxglove

Aethusa cynapium Sanicula europaea Conopodium majus Calluna vulgaris Vaccinium myrtillus Primula vulgaris Primula veris ag Lysimachia nemorum Anagallis arvensis Vinca major Sherardia arvensis Rubia peregrina Galium verum Galium mollugo Galium aparine Galium odoratum Cruciata laevipes Calystegia sepium Convolvulus arvensis Lithospermum offinale Symthytum offininalis Pentaglottis sempervirens Myosotes arvensis Lamium album Lamium purpureum Galeopsis tetrahit Ajuga reptans Lamiastrum galeobdolon Stachys officinalis Salvia verbenacea Stachys sylvatica Stachys palustris Glechoma hederacea Prunella vularis Clinopodium vulgare Clinopodium ascendens Origanum vulgare Thymus polytrichus Teucrium scorodonia Lycopus europaeus Solanum dulcamara Atropa belladonna Veronica arvensis Veronica officinalis Veronica chamaedrys Veronia persica Veronica serpyllifolia Pedicularis sylvatica Verbascum thapsus Scropularia nodosa Cymalaria muralis Melampyrum pratense Odontites vernus Digitalis purpurea

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Yellow-rattle Eyebright Caprofoliaceae Honeysuckle Red Valerian Campanulaceae Hairbell Nettle-leaved Bellflower Monotropidae Yellow Birds-nest Araceae Lords-and-Ladies Amaryllidaceae Wild Daffodil Snowdrop Summer Snowflake Iridacea Yellow Flag Stinking Iris Montbretia Dioscoreaceae Black Bryony Liliacea Three-cornered Garlic Ramsons Bluebell Meadow Saffron Solomon's Seal Orchidacea Common Spotted Orchid Early Purple Orchid Bird's-nest Orchid Greater Butterfly Orchid Common Twayblade Bee Orchid Pyramidal Orchid Fly Orchid Broad-leaved Helleborine White Helleborine Poaceae Sweet Vernal grass Crested Dogs tail Meadow Foxtail Smooth Meadow-grass Rough Meadow-grass Annual Meadow-grass Flattened Meadow-grass Wood Meadow-grass Yorkshire Fog Creeping Soft-grass Cocksfoot Fern-grass Tufted Hair-grass Wavy Hair-grass False Brome Sheep's Fescue Red Fescue Rat's-tail Fescue Wood Melick Mountain Melick False Oat Grass Quaking Grass Perennial Rye grass

Rhinanthus minor Euphrasia ag Lonicera pericyclamenum Centranthus ruber Campanula rotundifolisa Campanula trachelium Monotropa hypopitys Arum maculatum Narcissus sp Galanthus nivalis Leucojum aestivum Iris pseudacorus Iris foetidissima Crocosmia xcrocosmiiflora Tamus communis Allium triquestrum Allium ursinum Hyacinthoides non-scripta Colchicum autumnale Polygonatum multiflorum Dactylorhiza fuchsii Orchis mascula Neottia nidus-avis Platanthera chlorantha Neottia ovata Ophrys apifera Anacamptis pyramidalis Ophrys insectifera Epipactis helleborine Cephalanthera damasonium Anthroxanthum odoratum Cynosurus cristatus Alopecurus pratensis Poa pratensis Poa trivialis Poa annua Poa compressa Poa nemoralis Holcus lanatus Holcus mollis Dactylis glomerata Catapodium rigidum Deschampsia caespitosa Deschampsia flexuosa Brachypodium sylvaticum sl.

Festuca ovina Festuca rubra Vulpia myuros Melica uniflora Melica nutans Arrhenatherum elatius Briza media Lolium perene

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Soft Brome Upright Brome Blue Moor Grass Wood Barley Timothy Heath Grass Common Reed Cyperaceae Glaucous Sedge Pendulous Sedge Soft-leaved Sedge Fingered Sedge Dwarf Sedge Spring Sedge Grey Sedge Remote Sedge Pale Sedge Wood Sedge Yellow Sedge Juncaceae Field Wood-rush Hairy Wood-rush Jointed Rush Slender Rush Hard Rush Compact rush Toad Rush

Bromus mollis Bromus erectus Sesleria caerulens Hordelymus europaeus Phleum pratense Danthonia decumbens Phragmites australis Carex flacca Carex pendula Carex montana Carex digitata Carex humilis Carex caryophyllacea Carex divulsa Carex remota Carex pallescens Carex sylvatica Carex viridula Luzula campestris Luzula pilosa Juncus articulatus Juncus tenuis Juncus inflexus Juncus conglomeratus Junus bufonius

Gynmosperms - Native or naturalised Coniferous trees and shrubs Taxaceae Yew

Taxus baccata

Pinaceae Larches Pine Spruces

Larix dedcidua agg.

Pinus sylvestris agg.

Picea abies agg.

Cupressaceae Leyland or Lawson Cypress

Cypress agg.

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Checklist of Pteridophyta - Ferns and allies Dryopteridacae Male Fern Scaly Male Fern Broad Buckler Fern Hard Shield Fern Soft Shield Fern Thelypteridaceae Lemon-Scented Fern Woodsiacae Lady Fern Dennstaedtiacae Bracken Blechnacae Hard Fern Polypodiacae Common Polypody Ophioglossacea Adder's-tongue Aspleniacae Hart's Tongue Fern Wall Rue Maidenhair Spleenwort Black Spleenwort Rustyback

Dryopteris filix-mas Dryopteris affinis Dryopteris dilatata Polystichum aculeatum Polystichum setiferum Oreopteris limbosperma Athyrium filix-femina Pteridium aquilinum Blechnum spicant Polypodium vulgare agg.

Ophioglossum vulgatum Phyllitis scolopendrium Asplenium ruta-muraria Asplenium trichomanes Asplenium adiantum-nigrum Ceterach officinarum

Checklist of Equisetaceae – Horsetails Marsh Horsetail Field Horsetail

Equisetum palustre Equisetum arvense

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1.2 Checklist of Non-vascular Plants Bryophytes – Mosses and Liverworts Taxonomy and nomenclature from Blockeel et al. A new checklist of the bryophytes of Britian and Ireland, 2020. Journal of Bryology- BBS. Local status - c= commonly found. f = frequently found. u = uncommon or rare with few sites or occurrences National status - *** Rare or uncommon - a Doward speciality. # = Difficult to identity in field.

Bryophyta - Mosses - Species recorded since 1970 on the Doward. - Listed by order in BBS sequence.

Sphagnales Blunt-leaved Bogmoss Lustrous Bog-moss

Sphagnum palustre Sphagnum subnitens

1.06 1.20

u u

Tetraphidales Pellucid Four-tooth Moss

Tetraphis pellucida

4.01

u

Polytrichales Common Smoothcap Dwarf Haircap Wood Hair-moss Juniper Haircap

Atrichum undulatum Pogonatum aloides Polytrichum formosum Polytrichum juniperinum

6.03 9.02 10.02 10.06

c u f u

Encalyptales Spiral Extinguisher-moss Common Extinguisher-moss

Encalypta streptocarpa Encalypta vulgaris

14.01 14.04

f

Funariales Bonfire Moss

Funaria hygrometrica

15.01

f

Dicranales Bendy Ditrichum Slender Ditrichum Brittle Swan-neck Moss Dwarf Swan-neck-moss Rusty Swan-neck Moss Heath Star-moss Large/Smaller White-moss Variable Forklet-moss Silky Forklet-moss Green Pocket-moss Petty Pocket-moss Narrow-leaved Pocket-moss Short-leaved Pocket-moss Lesser Pocket-moss Flatfoot Pocket-moss Slender Pocket-moss Common/great Pocket-moss Rock Pocket-moss Maidenhair Pocket-moss Broom Fork-moss Greater Fork-moss Common Pincushion Taper-leaved Earth-moss Redshank

Flexitrichum flexicaule Flexitrichum (Ditrichum) gracile Campylopus fragilis Campylopus pyriformis Campylophus flexuosus Campylophus introflexus Leucobryum glaucum Dicranella varia Dicranella heteromalla Fissidens viridulus Fissidens pusillus Fissidens gracilifolius Fissidens incurvus Fissidens bryoides Fissidens crassipes Fissidens exilis Fissidens taxifolius Fissidens dubius Fissidens adianthoides Dicranum scoparium Dicranum majus Dicranoweisia cirrata Pleuridium acuminatum sl.

Ceratodon purpureus

21.01 21.20 26.04 26.05 26.06 26.11 27.01 31.05 31.10 32.01 32.03 32.04 32.05 32.06 32.09 32.11 32.15 32.16 32.17 34.04 34.06 39.01 46.01 50.01

u f u

Pottiales Whorled Tufa Moss Small-mouthed Beardless-moss Green-tufted Stubble-moss Variable Crisp-moss Curly Crisp-moss

Eucladium verticillatum Weissia brachycarpa Weissia controversa Trichostomum brachydontium Trichostomum crispulum

53.01 54.06 54.01 55.01 55.02

f u u u f

15

u

#

u

u f

u u c

u u

u u

f u u

c f u

f u u u

f

note 1.

Neat Crisp-moss Frizzled Crisp-moss Slender Stubble-moss Luisier's Tufa-moss Blunt-leaved Tufa Moss Verdigris Tufa-moss Hornschuch's Beard-moss Revolute Beard Moss Red Beard-moss Lesser Bird's-claw Beard-moss Bird's-claw Beard-moss Wavy Beard-moss False Beard-moss Dusky Beard-moss Olive Beard-moss Nicholson's Beard-moss Soft-tufted Beard-moss Cylindrical Beard-moss Rigid Beard-moss Aloe Moss Bordered Screw-moss Wall Screw-moss Common Pottia Cuspidate Earth-moss Stanford Screw-moss Great Hairy Screw-moss Intermediate Screw-moss Small Hairy Screw-moss Marble Screw-moss Water Screw-moss Smaller Lattice-moss

Tortella nitida Tortella tortuosa Gyroweisa tenuis Gymnostomum viridulum Gymnostomum calcareum Gymnostumum aeruginosum Pseudocrossidium hornschuchianum Pseudocrossidium revolutum Bryoerythrophyllum recurvirostrum Streblotrichum (Barbula) convoluta Barbula unguiculata Didymodon sinuosus Didymodon fallax Didymodon luridus Didymodon tophaceus Didymodon nicholsonii Didymodon vinealis Didymodon insulanus Didymodon rigidulus Aloina aloides Tortula marginata Tortula muralis Tortula truncata Tortula (Phascum) acaulon Hennediella stanfordensis Syntrichia (Tortula) ruralis Syntrichia montana Syntrichia laeviphila Syntrichia papillosa Syntrichia latifolia Cinclidotus fontinaloides

57.02 57.05 61.01 62.01 62.02 62.03 67.01 67.02 68.01 70.01 71.01 72.01 72.04 72.06 72.07 72.08 72.09 72.10 72.13 76.03 77.07 77.10 77.19 77.21 79.01 82.01 82.04 82.07 82.08 82.09 83.01

u f u u

Gimmiales Thickpoint Grimmia Elegant Grimmia Grey-cushioned Grimmia Hair-pointed Grimmia

Schistidium crassipilum Schistidium elegantulum Grimmia pulvinata Grimmia trichophylla

92.17 92.18 93.16 93.20

f

Bryales Silver Moss Tall-clustered Thread-moss Bicoloured Bryum Wall Thread-moss Capillary Thread-moss Crimson-tuber thread-moss Marsh Bryum Pink-fruited Thread-moss Pale-glaucous Thread-moss Forest Star Bordered Thyme-moss Starry Thyme-moss Dotted Thyme-moss Woodsy Thyme-moss Many-fruited Thyme-moss Hart's-tongue Thyme moss Long-beaked Thyme-moss

Bryum argenteum Bryum pallens Bryum dichotomum Bryum radiculosum Bryum capillare Bryum rubens Bryum pseudotriquetrum Pohlia melanodon Pohlia wahlenbergii Mnium hornum Mnium marginatum Mnium stellare Rhizomnium punctatum Plagiomnium cuspidatum Plagiomnium affine Plagiomnium undulatum Plagiomnium rostrata

110.05 110.06 110.09 110.12 110.42 110.52 110.48 113.18 113.19 115.01 115.05 115.06 117.01 118.01 118.02 118.06 118.07

c u f u

Orthotrichales Green Yoke Moss Lesser Yoke-moss Stirton's Yoke-moss Wood Bristle-moss Lyell's Bristle moss

Zygodon viridissimus Zygodon conoideus Zygodon stirtonii Lewinskya (Orthotrichum) affinis Pulvigera (Orthothrichum) lyellii

120.01 120.04 120.02 123.03 125.01

c

16

u u

u u u c c f

f f u u

u c f

#

u

u c f u u u c u u u u

u ***

#

c u

c f u u u

c u u

u f f

c f

#

u

u c u

#

River Bristle-moss Elegant Bristle-moss Straw Bristle-moss Anomalous Bristle-moss White-tipped Bristle-moss Bruch’s Pincisshion Cape Thread-moss Drumsticks

Orthotrichum rivulare Orthotrichum pulchellum Orthotricum stramineum Orthotrichum anomalum Orthotrichum diaphanum Ulota bruchii Orthodontium lineare Aulacomnium androgynum

122.05 122.03 122.16 122.01 122.07 126.04 128.01 131.03

u

Hypnales Starry Feather-moss Dull-starry Feather-moss Greater Water-moss Dentated Silk-moss Juicy Silk-moss Woodsy Silk-moss Elegant Silk-moss Fern-leaved Hook-moss Curled Hook-moss Creeping Feather-moss Tiny Feather-moss Chalk Feather-moss Sickle-leaved Hook-moss Many-fruited Leskea Tamarisk Moss Neat Feather Moss Lesser-striate Feather-moss Common Striated Feather-moss Wall Feather-moss Clustered Feather-moss Tender Feather-moss Hair-pointed Feather-moss Beech Feather-moss Dwarf Feather Moss Swartz's Feather-moss Twist-tip Feather-moss Common Feather-moss Matted Feather-moss White Feather-moss Streaky Feather-moss Rough-stalked Feather-moss Tufted Feather-moss Velvet Feather-moss Silky Wall Feather-moss Yellow Feather-moss Cypress-leaved Plait-moss Mamillate Plait-moss Heath Plait-moss Depressed Feather-moss Flat-brochade Moss Pointed Spear-moss Lindberg’s Plait-moss Springy Turf Moss Little Shaggy Moss Big Shaggy Moss Short-beaked Wood-moss Lateral Cryphaea Flat Neckera Crisped Neckera Blunt Feather-moss Fox-tail Feather-moss Mouse-tail moss Larger Mouse-tail Moss

Campylium stellatum Campylium protensum Fontinalis antipyretica Plagiothecium denticulatum Plagiothecium succulentum Plagiothecium nemorale Pseudotaxiphyllum elegans Cratoneuron filicinum Palustriella commutata Amblystegium serpens Serpoleskea confervoides Campylophyllopsis calcarea Sanionia uncinata Leskea polycarpa Thuidium tamariscinum Pseudoscleropodium purum Plasteurhychium striatulum Eurhynchium striatum Rhynchostegium murale Rhychostegium confertum Rhychostegiella tenella Cirriphyllum piliferum Cirriphyllum crassinervium Microeurychium pumilum Oxyrrhynchium hians Oxyrhychium schleicheri Kindbergia (Eurynchium) praelonga Sciuro-hypnum populeum Brachythecium albicans Brachythecium glareosum Brachythecium rutabulum Scleropodium cespitans Brachytheciastrum velutinum Homalothecium sericeum Homalothecium lutescens Hypnum supressiforme Hypnum andoi Hypnum jutlandicum Taxiphyllum wissgrillii Platygyrium repens Calliergonella cuspidata Calliergonella lindbergii Rhytidiadelphus squarrosus Rhytidiadelphus loreus Hylocomiadelphus triquetus Loeskeobryum brevirostre Cryhaea heteromala Neckera complanata Neckera crispa Homalia trichomanoides Thamnobryum alopecurum Isothecium myosuroides Isothecium alopecuroides

134.01

u u f

17

134.02 137.01 141.03 141.08 141.09 143.01 150.01 151.01 153.01 161.01 162.01 170.01 174.01 178.01 180.01 182.02 183.01 184.03 184.04 185.01 186.01 186.02 187.01 188.01 188.02 189.01 190.04 191.01 191.03 191.06 192.01 194.01 195.01 195.02 196.01 196.02 196.04 197.01 198.01 201.01 201.02 210.01 210.03 211.01 212.01 218.01 223.04

223.03 224.01 225.01 229.01 229.03

f f f f c u u

u f

#

u f u

u c u *** u *** u u c

c u *** c

#

u f f f u

f u u ***

c u

#

u f c

#

BBS

u

u

#

c

u c f f u

u c u

c u c u c c f u f c c

# note 2.

Comb Moss Rambling Tail-moss

Ctenidium molluscum Anomodon viticulosus

230.01 233.02

c c

162 spp Notes to above list: 1. All or Leucobryum glaucum records could in fact be L. juniperoideum as field determination is not easy.

2. This now includes Hypnum resupinatum, H. lacunosum due to recent taxonomic changes

Additional Moss records – Old records prior to 1970 or not reliably seen since. Listed in alphabetic order.

AL – Rev. Augustin Ley ca. 1880 Aloina ambigua Anomodon longifolius Amphidium mougeotii Bartramia pomiformis Sciuro-hypnum plumosum Bryum cespiticium Bryum canariensis Bryum donianum Bryum kunzei Bryum torquescens Bryum turbinatum Campylophyllum chrysophyllum Climacium dendroides Dialytrichia mucronata Dichodontium pellucidum Dicranum fuscescens Dicranum spadiceum Dicranum scottianum Entosthodon fascicularis Entostodon obtusus Ephemerum stoloniferum Grimmia montana Grimmia orbicularis Hedwigia ciliata Heterocladium heteropterum Hygrohypnum luridum Hylocomium splendens Leptodictyum riparium Leucodon scuiroides Microbryum davallianum Orthotrichum tenellum Orthotrichum sprucei Physcomitrium pyriforme Philonotis fontana Platydictya jungermanniodes Rhychostegium ripariodes Pleurochaete squarrosa Pohlia proligera Pohlia nutans Polytrichum piliferum Pseudephemerum nitidum Nogopterium gracile Ptychomitrium polyphyllum Racomitrium fasciculare Racomitrium heterostichum Rhodobryum roseum Rhychostegiella teneriffae Rhytidium rugosum Schistidium apocarpum Scorpiurium circinatum Seligeria pusilla

BBS no.

76.04 216.01 28.02 106.02 190.05 110.49 110.04 110.41 110.50 110.45 110.32 152.04 147.01 66.01 29.01 34.04 34.05 34.12 16.04 16.06 65.05 93.05 93.17 95.01 228.01 159.01 214.01 155.01 221.01 78.02 122.05 122.06 17.01 107.05 139.01 184.01 64.01 113.12 113.04 10.05 47.01 22.01 88.01 94.04 94.09 111.01 185.04 216.01 92.05 181.01 87.01

AL 19thC, Symonds Yat. Status Unclear if ever on Doward side of Wye.

AL

BBS AL AL AL AL

AL AL

18

Seligeria donniana Seligeria acutifolia Trichodon cylindricus Ulota crispa 55spp

87.06 28.02 48.01 126.07

BBS status unclear cf. U bruchii

Marchantiophyta - Liverworts Species recorded since 1970 - Listed by order in BBS sequence

Lunulariales Crescent-cup Liverwort Common Liverwort

Lunularia cruciata Marchantia polymorpha

4.01 5.01

f f

Marchantiales Great Scented Liverwort Hemisphaeric Liverwort Lesser Crystalwort

Conocephalum conicum Reboulia hemisphearica Riccia subbifurca sl.

9.01 6.01 10.12

u u u

Pelliales Endive Pellia Overleaf Pellia

Pellia endiviifolia Pellia epiphylla

13.03 13.01

f f

Fossombroniales Common Frillwort

Fossombronia pusilla sl

14.05

u

Metzgeriales Bluish Veilwort Whiskered Veilwort Forked Veilwort Downy Veilwort Greasewort Jagged Germanderwort

Metzgeria violacea Metzeria consanguinea Metzgeria furcata Metzgeria pubescens Aneura pinguis Riccardia chamedryfolia sl.

19.01 19.02 19.03 19.06 20.01 21.02

f f c u *** u u

Porellales Wall Scalewort Bitter Scalewort Even Scalewort Dilated Scalewort Minute Pouncewort Micheli’s Least Pouncewort Mackay's Pouncewort Fairy Beads

Porella platyphylla Porella arboris-vitae Radula complanata Frullania dilatata Myriocoleopsis minutissima Lejeunea cavifolia Marchesinia mackaii Microlejeunea ulicina

22.01 22.03 23.01 24.05 27.01 31.01 32.01 33.01

f f f c u u u *** f

Jungermanniales Creeping Fingerwort Bifid Crestwort Variable-leaved Crestwort Greater Featherwort Lesser Featherwort Common Pouchwort Mueller's Pouchwort Notched Pouchwort Scarce Notchwort Top Notchwort Dark-green Flapwort Two-horned Pincerwort Rustwort White Earwort Grove Earwort Heath Earwort

Lepidozia reptans Lophocolea bidentata Lophocolea heterophylla Plagiochila asplenoides Plagiochila porelloides Calypogeia fissa Calypogeia muelleriana Calypogeia arguta Mesoptychia badensis Mesoptychia turbinata Jungermannia atrovirens Cephalozia bicuspidata Cephalozia curvifolia Diplophyllum albicans Scapania nemorea Scapania irrigua

40.01 44.01 44.03 48.03 48.02 54.01 54.02 54.08 57.07 57.08 60.01 71.01 71.02 92.01 94.11 94.12

f c f c c c u c u u u f u c f u

19

R. multifida

Liverworts continued – Old Records prior to 1970 or not reliably seen since. Listed in alphabetic order.

Cololejeunea rosettiana Frullania tamarisci Lophosia excisa Marsupella emmarginata Metzgeria conjugata Orthocaulis attenuatus Plagiochila Britannica Riccia glauca Riccia sorocarpa Scapania compacta

26.03 24.01 87.05 67.02 19.04 76.03 48.04 10.09 10.07 94.01

Anthocerotophyta - Hornworts Anthoceros punctatus 1.01

Algae This group is largely outside the scope of this publication as most species require advance microscopic techniques for determination. However, some species are important for various ecological reasons and can be noticeable when growing on various land and aquatic substrates.

Trentapohlia aurea, a filamentous green alga, is a common sight common on trees in The Doward and noticeable due its bright orange colouration caused by the production of carotinoid chemicals. This and the genus Trebouxia are from the class of Chlorophyta are the partner alga or photobiont of many Lichen species such as Xanthoria parietina.

Four other species of algae from Phylum Chlorophyta are also listed by NBN from 1977. Gongrosira incrustans, Paulschulzia, Cladophora glomerata, Ulva flexuosa.

Phylum Charophyta (aquatic Stoneworts) reported by NBN just outside Doward at Dixton, Monmouth with two species Chara vulgaris and C. globularis listed (1891 and 1944 respectively and again, but not to species level, in 1979).

20

1.3. Checklist of Larger Fungi and Lichens The list below is generated from a range of ‘informal’ observations by various recorders on the Doward rather than a set of formal records. It should therefore be treated as for guidance purposes only.

The family classification and sequence are taken from that used in Sterry, P & Hughes B. (2009). Collins Complete Guide to British Mushrooms & Toadstools. It has NOT been checked with the latest version from the British Mycological Society.

Most families of larger fungi are in the Phylum Basidomycota although some are in Ascomycota. Lichens are treated separately. *** denotes a nationally uncommon or notable species.

Larger Fungi: Boletacea Bay Bolete type Cep Devil's Bolete Lurid Bolete Scarletina Bolete type Bitter Beech Bolete Rooting Bolete Orange Oak Bolete Brown Birch Bolete Larch Bolete Hollow Bolete n/a Bovine Bolete n/a Matt Bolete Russulaceae Blackening Brittlegill Ochre Brittlegill Charcoal Burner Golden Brittlegill Birch Brittlegill Crowded Brittlegill Fragile Brittlegill Milk White Brittlegill Scarlet Brittlegill Bare-toothed Brittlegill Beech Milkcap Mild Milkcap Fiery Milkcap Oakbug Milkcap Tawny Milkcap Woolly Milkcap Yellowdrop Milkcap Pale Milkcap Birch Milkcap Agaricaceae George’s Mushroom Yellow Stainer Wood Mushroom n/a Freckled Dapperling Stinking Dapperling Shield Dapperling Parasol Slender Parasol n/a Hygrophoraceae Snowy Waxcap

Boletus badius Boletus edulis Boletus satanas *** Boletus luridus Boletus luridiformis/erythropus ag Boletus calopus Boletus radicans Leccinum aurantiacum Leccinum scabrum Suillus grevillea Suillus cavipes *** Suillus tridentinus Suillus bovinus Xerocomellus cisalpinus Xerocomellus pruinatus Russula nigricans Russula ochroleuca or ag Russula cyanoxantha Russula risigallina Russula betularum Russula densifolia Russula fragilis Russula delica Russula pseudointegra Russula vesca Lactarius blennius Lactarius subdulcis Lactarius pyrogalus Lactarius quietus lactarius fulvissimus Lacterius torminosus Lactarius chrysorrheus Lactarius pallidus Lactarius tabidus Calocybe gambosa Agaricus xanthodermus Agaricus silvicola Lepiota hystrix Lepiota aspera Lepiota cristata Lepiota clypeolaria Macrolepiota procera Macrolepiota mastoides Cystolepiota hetieri Hygrocybe virginea

21

Spangle Waxcap Scarlet waxcap Butter Waxcap Blackening Waxcap Honey Waxcap Parrot Waxcap Meadow Waxcap Cedarwood Waxcap Hygrophoropsidaceae Ivory Woodwax Cantharellaceae False Chanterelle Trumpet Chanterelle Chanterelle Coprinaceae Glistening Inkcap Shaggy Inkcap Pleated Inkcap Fairy Inkcap Magpie Inkcap Common Inkcap Psathyrellaceae Stump Brittlestem Conical Brittlestem Pale Brittlestem Brown Mottlegill Petticoat Brittlestem Weeping Widow Tricholomataceae Common Bonnet Drab Bonnet Burgundydrop Bonnet Saffrondrop Bonnet Lilac Bonnet Ivory Bonnet Milking Bonnet Blackedge Bonnet Pink Bonnet n/a Snapping Bonnet n/a Russet Toughshank Whitelaced Shank Sulphur Knight Blue spot Knight Soapy Knight n/a Clustered Domecap Club Foot Velvet Shank The Deceiver Amythyst Deceiver Wood Blewit Tawny Funnel Porcelain Fungus Trooping Funnel Common Funnel Ivory/Frosty Funnel Honey Fungus Plums and Custard Common Cavalier Twig Parachute Collared Parachute

Hygrocybe insipida Hygrocybe coccinea Hygrocybe ceracea Hygrocybe conica Hygrocybe reidii Hygrocybe psittacina Hygrocybe pratensis Hygrocybe russocoriacea Hygrophorus eburneus Hygrophoropsis aurantiaca Cantharellus tubaeformis Cantharellus cibarius Coprinus micacea Coprinus comatus Parasolus (Coprinus) plicatilis Coprinellus disseminatus Coprinus picacea Coprinus atramentaria Psathyrella piluliformis Psathyrella conophilus Psathyrella candoleana Panaeolina foenisecii Psathyrella artemisia Lacrymaria lacrymabunda Mycena galericulata Mycene aetites sl.

Mycena haematopus Mycena crocata Mycena pura Mycena flavoalba Mycena galopus Mycena pelianthina Mycena rosella Mycena stipata Mycena vitilis Hydropus subalpinus Collybia (Gymnopus) dryophila Megacollybia platyphylla Tricholoma sulphureum Tricholoma columbretta Tricholoma saponaceum Tricholoma ustale Lyophyllum decastes Ampulloclitocybe clavipes Flamulina velutipes Laccaria laccata Laccaria amethystina Lepista nuda Lepista flaccida Oudemansiella mucida Clitocybe geotrupa Clitocybe gibba Clitocybe dealabata/phyllophila Armillaria mellea Tricholomopsis rutilans Melanoleuca polioleuca Marasmiellus ramealis Marasmiellus rotula

22

Orange Mosscap White Domecap Clustered Domecap Strophariaceae Blue Roundhead Verdigris Roundhead Sulfur Tuft Shaggy Scalycap Golden Scalycap Petticoat Mottlegill Cortinariaceae Bulbous/Star Fibrecap type White/Lilac Fibrecap Scaly Fibrecap n/a n/a Common Rustgill n/a Variable Webcap Dappled Webcap Stocking Webcap n/a n/a Poisonpie Sweet Poisonpie Funeral Bell Moss Bell Entolomataceae Wood Pinkgill Pimple Pinkgill n/a n/a The Miller Bolbitaceae Yellow Fieldcap Velvet Toughshank Pluteaceae Deer Shield n/a Paxillaceae Brown Rollrim Auriscalpiaceae Aniseed Cockleshell Amanitaceae Snakeskin Grisette Rooting Shank Fly Agaric Grey-spotted Amanita Death cap False Deathcap Tawny Grisette Psathyrellaceae n/a n/a Lentinaceae Oyster Mushroom Pale Oyster Crepidotaceae Variable Oysterling n/a n/a Scurfy Twiglet Felted Twiglet

Rickenella fibula Lyophyllum connatum Lyophyllum decastes Stropharia caerulea Stropharia aeriginosa Hypholoma fasciculare Pholiota squarrosa Pholiota aurivella Panaeolus papilionaceus Inocybe napipes/asterospora Inocybe geophylla Inocybe hystrix Inocybe cincinnata var. major Inocybe patouillardii Gymnopilus penetrans Conocybe tenera Cortinarius anomalus Cortinarius bolaris Cortinarius torvus Cortinarius olivaceofuscus Cortinarius elegantissimus Hebeloma crustuliniforme Heboloma sacchariolens Galerina marginata (was G. autumnalis) Galerina hypnorum Entoloma rhodopolium Entoloma hebes Entoloma incanum Entoloma conferendum Clitopilus prunulus Bolbitius titubans Kuehneromyces mutabilis Pluteus cervinus Pluteus salicinus Paxillus involutus Lentinellus cochleatus Amanita ceciliae Xerula radicata Amanita muscaria Amanita excelsa Amanita phalloides Amanita citrina Amanita fulva Psathyrella candolleana Psathyrella leucotephra Pleurotus ostreatus Pleurotus pulmonarius Crepidotus variabilis Crepidotus lundellii Crepidotus cesatii Tubaria furfuracea Tuberia conspersa

23

Xylariacae Dead Man's Fingers Dead Moll's Finger Candlesnuff Beech Woodwart Hazel Woodwart Cramp Balls Heliotiales insertae sedis Green Elfcup Helotiacea Beech Jellydisc Purple Jellydisc Common Grey Disco Lemon Disco Bulgariaceae Black Bulgar Auriculariaceae Jelly (Jew's) Ear Witches Butter Leotiacea Jellybaby Clavariaceae Yellow Club Meadow Coral Crested Coral White Spindles Slender Club Coral sp Upright Coral Clavariadelphaceae Giant Club Phallaceae Stinkhorn Dacrymycetaceae Yellow Stagshorn Small Stagshorn Yellow Brain Orange Brain Helvellacea Elfin Saddle Common White Saddle Orange Peel Fungus Geoglossaceae Earthtongue Earthtongue Sarcoscyphaceae Scarlet Elfcups Geastraceae Collared Eathstar Nidulariacea Fluted Bird's Nest Lycoperdacea Common Puffball Stump Puffball Spiny Puffball Meadow/grassland Puffball Sclerodermataceae Common Earthball Polyporacaea Dryad's Saddle Blackfoot Polypore Tuberous Polypore

Xylaria polymorpha Xylaria longipes Xylaria hypoxxylon Hypoxylon fragiforme Hypoxylon fuscum Daldinia concentrica Chlorociboria aeruginascens Neobulgaria pura Ascocoryne sarcoides Mollisia cinerea Bisporella citrina Bulgaria inquinans Auricularia auricula-judae Exidia glandulosa Leotia lubrica Clavulinopsis helvola/ luteoalba Clavulinopsis corniculata Clavulina coralloides Clavaria fragilis Macrotyphula juncea Ramaria spp poss. curta or decurrens Ramaria stricta Clavariadelphus pistillaris Phallus impudicus Calocera viscosa Calocera cornea Tremella mesenterica Tremella aurantia Helvella lacunosa Helvella crispa Aleuria aurantia Geoglossum cookeanum ag Geoglossum viride or olivacum Sarcoscypha austraca Geastrum triplex Cyathus olla Lycoperdon perlatum Lycoperdon pyriforme Lycoperdon echinatum Lycoperdon pratense/lividum Scleroderma citrinum Polyporus squamosus Polyporus leptocephalus/varius Polyporus tuberaster

24

Winter Polypore Ganodermateae Artists Bracket n/a Turkeytail Lacquered Bracket Purplepore Bracket Hazel Bracket Fistulinaceae Beefsteak Fungus Hymenochaetaceae Oak Bracket Hydnaceae Wood hedgehog Terracota Hedgehog Coriolaceae Birch Polypore Fomitopsidaceae Powderpuff Bracket Resupinates Wrinkled Crust Cobalt Crust Bleeding Broadleaf Crust Bleeding Oak Crust n/a Silverleaf Fungus Morchellacea Bleach Cup Semifree Morel

Polyporus brumalis

Rust on Pendulous Sedge

Puccinia caricina var ribesii-pendulae

Tar spot

Rhytisma acerinum

Ganoderma applanatum Ganoderma australe Trametes versicola Ganoderma lucidum Trichaptum abietinum Skeletocutis nivea Fistulina hepatica Inonotus dryadeus Hydnum repandum Hydnum rufescens Piptoporus betulinus Postia ptychogaster Phlebia radiata Terana caerulea Stereum rugosum Stereum gausapatum Stereum hirsutum Chondrostereum purpureum Disciota venosa Morchella semilibera

25

A checklist of Lichens A substantial number of species are listed on NBN and in other sources but due to recent taxonomic changes a detailed review would be required to provide an updated list.

The following list is a set of common species known on the Doward using old nomenclature and are all unverified records so should be treated as such.

Arthonia radiata Graphis scripta ag.

Verrucaria nigrescens ag.

Caloplaca citrina ag.

Caloplaca flavescens Lecidella elaeochroma Lecanora expallens ag.

Lecanora chlarotera Lecanora albella sl.

Lecanora sp prob dispersa sl.

Xanthoria parietina Physcia tennella-adscendens Physcia aipolia Parmelia sulcata Punctelia subrudecta Hypnogymia physodes Evernia prunastri Ramalina fastigiata Ramalina farinacea Usnea subfloridana Usnea florida Peltigera membranacea ag.

Cladonia coniocraea ag.

Hypotrachyna revoluta Collema auriforme Protoblastenia rupestris

26

Supplementary Notes 1.

Endemics plants of the Doward

The two taxa groups where the local speciation is deemed to have occurred are: 1. Whitebeams: In a contribution to Herefordshire Wildlife Trust’s March 2015 Flycatcher magazine Dave Green asserted that the Doward now has 14 species of Sorbus of which 5 are endemic to the Lower Wye Valley with two being found only on the Doward. He also described the Doward is the second most important site in the British Isles for its Sorbus diversity after the Avon Gorge. Studies by Tim Rich of Cardiff University aided by Dave Green and others describe an ongoing evolutionary process for this genus in this area of the Wye Valley. Hybridisation and then back-crossing has taken place as revealed by genetic study techniques such as flow cytometry and with virtually all Whitebeam trees on the Doward having now been examined and mapped. It seems that subtly different habitats in closed canopy woodland and woodland edges on rocky ground have led to a range of new species being able to survive.

The full explanation of whitebeams on the Doward is given in scientific publications. In summary, there are three well known and well distributed species of Sorbus in UK, the Common Whitebeam Sorbus aria, the Rowan Sorbus aucuparia and the Wild Service Tree Sorbus torminalis and all occur on the Doward in reasonable numbers.

Two species are considered endemic to the Doward: Sorbus herefordensis - Herefordshire Whitebeam, a woodland canopy species found in Miner’s Rest reserve and other central areas of Great Doward.

Sorbus greenii - Green’s Whitebeam, a woodland edge species found particularly around old quarries in the west and north of Great Doward.

Nine further species occurring on the Doward are considered near-endemic as they also occur in nearby areas of the Lower Wye Valley: Sorbus evansii Evans Whitebeam Cliff edge or woodland fringe Sorbus eminentiformis Doward Whitebeam Open high woodland Sorbus rupicola Rock Whitebeam Cliff edge Sorbus saxicola Symonds Yat Whitebeam.

Cliff edge Sorbus porrigentiformis Grey-leaved Whitebeam Rock faces Sorbus anglica English Whitebeam Rock faces Sorbus eminens Round-leaved Whitebeam Cliff face and woodland fringe Sorbus thuringiacia unnamed Cliff face (= S. aria x S. aucuparia) Sorbus x decipiens Wye Whitebeam (old name) Open high woodland (= S. aria x S.torminalis previously called S. x tomentella) Also not recorded on Doward but found nearby on Coldhams Rocks: Sorbus parviloba Ship Rock Whitebeam.

27

One of the rare cliff-edge Sorbus species at Seven Sisters.

2. Hawkweeds – Hieracium species: To most people Hawkweed is just another yellow Dandelion-like plant but to people who like to dig deeper into the taxonomy and morphology of the family Aceracea (or Compositae to the old school) there is a range of interesting micro-species. One of them, Hieracium subplanifolium, is endemic to the Doward but known only from a small patch. It is unclear how many other similar Hawkweed species occur here, but this this aggregate can be numerous especially around rocky areas such as in Lords Wood Quarry NR and on certain other thin-soiled track verges.

A Hawkweed Hieracium spp in Lord’s Wood Quarry

28

Part 2. – Invertebrates: This part of the publication presents checklists of all the main terrestrial invertebrate groups covering Insects, Arachnids, Molluscs, Crustaceans, Springtails and Myriapods. Microscopic invertebrates are omitted and some groups such as annelids, nematodes and mites are lacking any significant information from this site at the time of writing.

Contents: Page

Class Insecta – The Insects 3.1 Order Odonata 3.2 Order Orthoptera 3.3 Order Hemiptera 3.4 Order Coleoptera 3.5 Order Diptera 3.6 Order Lepidoptera

Dragonflies and Damselflies Grasshoppers and Crickets Plant Bugs Beetles True Flies Butterflies and Moths Moths 3.7 Order Hymenoptera Bees, Wasps and Ants 3.8 Orders Ephemeroptera (Mayflies), Plecoptera (Stoneflies) and Order Tricoptera (Caddisflies) 3.9 Other Insect Orders

31.

32.

33.

36.

44.

54.

55.

70.

73.

74.

76.

Class Arachnida 3.10 Order Aranea – Spiders 3.11 Orders Opiliones (Harvestmen), Psuedoscorpionida and Acari (mites)

77 79

Phylum Mollusca 3.12 Classes Gastropoda and Bivalva – Slugs, Snails and Mussels

80

Other Invertebrates 3.13 Class Crustacea – Woodlice and freshwater shrimps 3.14 Sub-phylum Myriapoda – Diploda – Millipedes - Chilopoda – Centipedes 3.15 Phylum Entognatha - Collebola – Springtails 3.16 Annelids, Nematodes, Platyhelminths and allies

29

83 84 85 86 87

2.1. Class Insecta – The insects There are around 24,000 known insect species in UK, which are usually divided into 24 taxonomic groups loosely called ‘Orders’. Checklists covering 18 of these with representatives on the Doward are presented here.

Each checklist is based around species in their taxonomic family within that order. However, the format and information presented is not uniform and in some cases information about local and national status is included and may or may not include the source of the record.

Coleoptera

A Burying Beetle Nicrophilus vespillioides

Diptera

The Downland Villa Villa cingulata

Lepidoptera

Grizzled Skipper – Pyrgus malvae

Orthoptera

Rufous Grasshopper – Gomphocerippus rufus

30

3.1. Checklist of Order Odonata: Dragonflies and Damselflies

Last update Aug’23

Suborder Zygoptera - Damselflies Calopterygidae Calopteryx virgo Calopteryx splendens

Beautiful Demoiselle Banded Demoisele

Status unclear Common (from River Wye)

Coenagrionidae Coenagrion puella Ishnura elegans Pyrrhosoma nymphula Platycnemis pennipes

Azure Damselfly Blue-tailed Damselfly Large Red Damselfly White-legged Damselfly

Common Occasional Occasional Common (from River Wye)

Suborder Anisoptera - Dragonflies Libellulidae Libellula depressa Sympetrum striolatum Libellula depressa Orthetrum cancellatum

Broad-bodied Chaser Common Darter Broad-bodied Chaser Black-tailed Skimmer

Regular Regular Regular Occasional

Aeshnidae Aeshna cyanea Aeshna mixta Anax imperator

Southern Hawker Migrant Hawker Emperor Dragonfly

Common Occasional Regular

Corduligasteridae Cordulegastor boltonii Gomphus vulgartissimus

Golden Ringed Dragonfly Club-tailed Dragonfly

Scarce visitor Regular (from River Wye)

31

3.2 Checklist of Order Orthoptera – Grasshoppers and Bushcrickets Acrididae - Grasshoppers Meadow Grasshopper Field Grasshopper Common Green Grasshopper Rufous Grasshopper Mottled Grasshopper

Chlorthippus parallelus Chlorthippus brunneus Omocestus viridulus Gomphocerippus rufus Myrmeleotettix maculatus

Common Common Reported but status unclear.

Little Doward 2023 Suspected but status unclear.

Pholidoptera griseoaptera Leptophyes punctatissima Meconema thalassinum Metrioptera roeselii Conocephalus discolor

Frequent Frequent Frequent Now frequent since 2013.

Regular small numbers.

Tetrix undulata

Frequent

Tettigonidae – Bushcrickets Dark Bushcricket Speckled Bushcricket Oak Bushcricket Roesel's Bushcricket Long-winged Conehead

Tetrigidae Common Groundhopper

32

3.3 Checklist of Order Hemiptera – Plant Bugs National Status codes C = Common L = Local Nb = Notably Scarce group B, present in 30-100 squares of 10km Record Source code ID = authors own record (if in italics if not verified by a national recording scheme) WT = Woodland Trust commissioned expert survey NBN* = National Biodiversity Network data. Asterix * denotes spatial resolution of SO51 only

Family

Species

National status

Source

Suborder - Heteroptera Acanthosomatidae Acanthosoma haemorrhiodale Elasmucha grisea Elasmostethus interstinctus

Hawthorn Shieldbug Parent Bug Birch Shieldbug

m WT m

Aradidae Aneurus avenius Aradus depressus

n/a n/a

L

Pentatomidae Dolycoris baccarum Palomena prassina Pentatoma ruficeps Troilus luridus Eurydema oleracea Elasmostethus interstinctus Aelia acuminata

Hairy or Sloe Shieldbug Green Shieldbug Forest (Red-legged) Shieldbug Bronze Shieldbug Crucifer Shieldbug Birch Shieldbug Bishop's Mitre

Scutelleridae Eurygaster testudinaria

Tortoise Shieldbug

Stenocephalidae Dicranocephalus medius

Spurgebug

L

m m

m m m ID ID ID NBN*

ID

Nb

ID/WT

Cynidae Legnotus limbosus Sehirus luctuosus Rhopalidae Corizus hyoscyami Rhophalus subrufus Myrmus miriformis

n/a Forget-me-not Shieldbug

NBN NBN

n/a n/a n/a

ID ID NBN*

Nabidae Himacerus apterus Nabis rugosus Coreidae Coreus marginatus Coriomeris denticulatus

Tree Damsel Bug n/a

c

Dock Bug Denticulated Leatherbug

c

ID WT m m

Lygaeidae Heterogaster urticae Drymus brunneus Drymus sylvaticus Drymus ryei Eremocoris podagricus Kleidocerys resedae Lasiosomus enervis Peritrechus lundi Scolopostethus affinis Scolopostethus thomsoni Stygnocoris rusticus Stygnocoris sabulosus n/a

Nettle Groundbug n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a

c c c L

33

ID m m WT WT WT NBN* WT m WT NBN* NBN*

Trapezonotus desertus Taphropeltus contractus

n/a n/a

m WT

Gerris lacustris ag.

Pond Skater

ID

Common Backswimmer

m

Water Boatman

ID

n/a n/a

NBN* NBN*

Gerridae

Notonectidae Nonecta glauca Corixidae Corixa sp Microphysidae Loricula elegantula Loricula pselaphiformis Miridae

Adelphocoris lineolatus Lucerne Bug Acetropis gimmerthalii n/a Blepharidopterus angulatus Black-kneed Apple Capsid Bryocoris pteridis n/a Capsus ater n/a Calocoris alpestris n/a Calocoris quadrimaculatus n/a Closterotomus norwegicus Potato Capsid Grypocoris stysi n/a Dicyphus epilobii n/a Dicyphus pallidicornis n/a Dicyphus globulifer n/a Deraeocoris flavilinea n/a Deraeocoris ruber n/a Deraeocoris lutescens n/a Dryophilocoris flavoquadrimaculatus n/a Harpocera thoracica n/a Heterocordylus genistae n/a Heterotoma planicornis n/a Leptopterna dolabrata Meadow Plant Bug Lygocoris pabulinus Common Green Capsid Liocoris tripustulatus n/a Monalocoris filicis Bracken Bug Oncotylus viridiflavus Knapweed Plant Bug Orthotylus (Orthotylus)marginalis Dark Green Apple Capsid Orthocephalus coriaceus n/a Orthonotus rufifrons n/a Notostira elongata Elongated Grass Bug Pachytomella parallela n/a Phytocoris varipes n/a Psallus perrisi n/a Psallus varians n/a Psallus wagneri n/a Phylus coryli n/a Phytocoris reuteri n/a Pinalitus cervinus n/a Pinalitus viscicola n/a Pithanus maerkeli n/a Plagiognathus arbustorum n/a Stenodema laevigata n/a Stenodema calcarata n/a Stenotus binotatus Timothy Grassbug Naucoridae Ilyocoris cimicoides

Saucer bugs

L

c c

c c

c

c c c

c c c

L

m NBN* NBN* NBN* m WT WT NBN* m WT WT NBN* NBN* m NBN* NBN* WT NBN* ID NBN* ID NBN* NBN* m NBN* NBN* NBN* m NBN* ID WT m WT NBN* NBN* WT NBN* m NBN* m NBN* NBN*

NBN

Reduviidae Empicoris vagabundus

n/a

Acalypta parvula Agramma laetum

n/a n/a

Velia caprai

Water Cricket

NBN*

Tingidae c L

WT WT

Veliidae NBN*

34

Suborder - Auchenorrhyncha Cercopidae Cercopis vulnerata

Red-and-black Froghopper

ID

Aphrophoridae Philaenus spumarius Aphrophora ulni

Common Froghopper n/a

ID m

Lacehopper sp n/a n/a

ID NBN* m

Delphacidae Dicranotropis hamata Ditropis pteridis Muellerianella fairmairei

n/a n/a n/a

NBN* NBN* NBN*

Membracidae Centrotus cornutus

Horned Treehopper

ID

Cicadellidae Agallia consobrina Alnetoidea alneti Anoscopus flavostriatus Aphrodes makarovi Cicadella viridis Eupterycyba jucunda Eupteryx aurata Eupteryx urticae Euscelis incisus Evacanthus interruptus Iassio lanio Kybos smaragdulus Ledra aurita Megophthalmus scanicus Oncopsis flavicollis Thamnotettix dilutior

n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a Oak Leafhopper n/a Eared Leafhopper n/a n/a n/a

NBN* NBN* NBN* NBN* ID NBN* NBN* NBN* NBN* m ID NBN* ID NBN* NBN* ID

n/a

NBN*

Hazel Whitefly

ID

Microlophium carnosum Aphis fabae prob.

Common Nettle Aphid Black bean aphid

ID ID

Arytainilla spartiophila Cacopsylla visci Psyllopsis fraxinicola

n/a n/a n/a

NBN NBN NBN

Cixiidae Cixius nervosus Cixius cunicularius Tachycixius pilosus

Issidae Issus coleoptratus

Suborder Sternorrhycha Aleyrodidae Aleyrodes proletella Aphididae

Psyllidae

35

3.4 Checklist of Order Coleoptera – Beetles Last update Aug’22 Source Codes: WT Woodland Trust commissioned Survey by NBN National Biodiversity Network (or Atlas) KAS Keith Alexandra Survey report for Buglife 2022 ID Ian Draycott (own records) either verified through iRecord or pending (italics). iRecord Records in iRecord (other than I Draycott) M Multiple sources i.e. >2 HBRC Herefordshire Biological Records Centre data (and not in NBN) Status Codes: NR-RDB1 Nationally Rare – Endangered. <5 hectads NR-RDB2 Nationally Rare – Vulnerable. 6-10 hectads NR-RDB3 Natioanlly Rare – Threatened. 11-15 hectads Na Nationally Scarce or Notable – 16-30 hectads Nb Nationally Scarce or Notable – 31-100 hectads L Local – 101-300 hectads

Family

Species

Aderidae

Euglenes oculatus

Anthicidae

Omonadus floralis

NBN

Anobiidae

Anobium punctatum Dorcatoma dresdensis Dorcatoma substriata Ptinus subpilosus Ptilinus pectinicornis Grynobius planus

KAS KAS KAS NBN WT,KAS WT, KAS

Anthribidae

SO51 only 1

1

Source KAS

Anthribus fasciatus Platyrhinus resinosus

1

Aphodiidae

Aphodius (Limarus) zenkeri Aphodius (Planolinus) fasciatus Aphodius (Bodilus) ictericus Aphodius (Chilothorax) conspurcatus Aphodius fossor Aphodius rufipes

1 1 1 1

NBN NBN NBN NBN iRecord iRecord

Apionidae

Aspidapion (Aspidapion) radiolus Perapion (Perapion) hydrolapathi Betulapion simile Protapion fulvipes

1 1

NBN NBN WT WT

Attelabidae

Attelabus nitens

1

NBN

Biphyllidae

Diplocoelus fagi

Buprestidae

Agrilus angustulus Agrilus biguttatus Agrilus sinuatus

Cantharidae 18

Jewel Beetle

Malthinus balteatus Malthodes minimus Malthodes flavoguttatus Malthodes marginatus Malthodes mysticus Malthinus flaveolus Malthodes fibulatus Malthodes pumilus Cantharis rustica Cantharis fusca Cantharis pellucida Cantharis decipiens Cantharis livida

1

1 1

36

NBN KAS, ID

Nat. Status pre-1970 Nb

1 L

1 Nb 1 1 1 1

1

KAS

Nb

M WT KAS

Na

NBN, KAS M NBN NBN, KAS M NBN KAS WT WT M M NBN M NBN

1

Nb Nb

1 1

Cantharis lateralis Rhagonycha fulva Rhagonycha lignosa Rhagonycha translucida Rhagonycha limbata Carabidae 26

Cerambycidae 27

1 Common Red Soldier Beetle 1

Cicindela campestris Green Tiger Beetle Badister bullatus Badister sodalis 1 Platynus assimilis 1 Agonum scitulum 1 Carabus arvensis 1 Carabus monilis 1 Carabus violaceus Violet Ground Beetle Calosoma inquisitor 1 Calodromius spilotus Abax parallelepipedus Synuchus vivalis 1 Bembidion varium Dromius agilis Dromius quadrimaculatus Notiophilus biguttatus Notiophilus palustris Syntomus foveatus Macronychus quadrituberculatus 1 Blemus discus Pterostichus oblongopunctatus Pterostichus nigrita ag Pterostichus madidus Poecilus cupreus Copper Greenclock Amara aenea Abax paralellepipidus Cychrus caraboides Alosterna tabacicolor Anaglyptus mysticus Anoplodera sexguttata Arhopalus rusticus Aromia moschata Clytus arietis Grammoptera ruficornis Leiopus nebulosus Leptura quadrifasciata Lepturobosca virens Mesosa nebulosa Molorchus minor Pachytodes cerambyciformis Phytoecia cylindrica Phymatodes testaceus Pogonocherus hispidus Prionus coriarius Pyrrhidium sanguineum Rhagium bifasciatum Rhagium mordax Ruptella maculata Saperda carcharias Saperda populnea Stenocorus meridianus Stenurella melanura Tetropium gabrieli Tetrops praeusta

NBN M M NBN M M M NBN NBN NBN NBN HBRC ID NBN KAS M NBN WT WT WT, KAS WT ID WT HBRC HBRC HBRC/NBN ID ID, KAS ID ID ID ID

Tobacco-coloured Longhorn Rufous-shouldered Longhorn 1 Dusky Longhorn 1 Wasp Beete Common Grammoptera

M M NBN ID NBN M M M Four-banded Longhorn NBN 1 NBN M Spruce Shortwing ID Speckled Longhorn M Umbelifer Longhorn 1 NBN Tanbark Borer ID Greater Thorn-tipped Longhorn M Tanner Beetle NBN Welsh Oak Longhorn M Two-banded Longhorn M Black-spotted Longhorn M Black and Yellow Longhorn M Large Poplar Borer 1 NBN Small Poplar Borer 1 NBN Variable Longhorn M Black-striped Longhorn M 1 NBN Plum Longhorn WT

Cerylonidae

Cerylon fagi Cerylon ferrugineum Cerylon histeroides

1

NBN M KAS

Cetoniidae

Trichius fasciatus Cetonia aurata

1 1

NBN NBN

Ciidae

Cis bilamellatus Cis boleti

WT, KAS WT, KAS

37

1

1

1 ?

1 1 1

1

1

Nb Nb

1 1

Nb 1 1

1 1 NR-RDB3

1

1 Na/RDB2

1 1

1

Cis castaneus Cis fagi Cis nitidus Octotemnus glabriculus

KAS KAS WT WT

Cleridae

Tillus elongatus

WT, KAS

Nb

Chrysomelidae 28

Apteropeda orbiculata Chrysolina polita Chrysolina americana Rosemary Beetle Chrysolina herbacea Mint Leaf Beetle Chrysolina brunsvicensis Chrysolina varians Chaetocnema hortensis Cryptocephalus moraei Cryptocephalus labiatus Cryptocephalus oreoles ag Crepidodera plutus Clytra quadripunctata Epitrix atropae Gastrophysa viridula Green Dock Beetle Mniophila muscorum 1 Pyrrhalta viburni 1 Phratora vulgatissima Phyllotreta consobrina Phyllotreta ochripes Phyllotreta nigripes Phyllotreta undulata Orsodacne lineola Oomorphus concolor Ouelema melanopus or rufocyanea Cereal Leaf Beetle Longitarsus membranaceus 1 Hermaeophaga mercurialis Lilioceris lilii Lily Beetle Sermylassa halensis

WT NBN ID M NBN WT WT M WT ID iRecord WT M NBN NBN NBN iRecord WT WT WT WT WT NBN ID NBN M iRecord ID

L

Adalia bipunctata 2-spot Ladybird Adalia decempunctata 10-spot Ladybird Anatis ocellata Eyed Ladybird 1 Aphidecta obliterata Calvia quattuordecimguttata Cream-spot Ladybird Chilocorus renipustulatus Kidney-spot Ladybird Coccidula rufa 1 Coccinella hieroglyphica Hieroglyphic Ladybird 1 Coccinella septempunctata 7-spot Ladybird Coccinella undecimpunctata 11-spot Ladybird 1 Exochomus quadripustulatus Pine Ladybird Halyzia sedecimguttata Orange Ladybird Harmonia axyridis Harlequin Ladybird Nephus redtenbacheri 1 Propylea quattuordecimpunctata 14-spot Ladybird Psyllobora vigintiduopunctata Rhyzobius litura Subcoccinella vigintiquattuorpunctata 24-spot Ladybird Tytthaspis sedecimpunctata 16-spot Ladybird

M M NBN M M M NBN NBN M NBN M M M NBN M M NBN NBN M

Anthophagus pallens Atomaria rubida Atomaria fuscata Atomaria linearis Atomaria lewisi Paramecosoma melanocephalum Cryptophagus angustus Cryptophagus dentatus Cryptophagus scanicus

WT NBN NBN NBN NBN NBN NBN KAS KAS

Coccinellidae 19

Cryptophagidae

Cucujidae

1 1 1 1 1

Pediacus dermestoides

1

38

NBN, KAS

L

1

Nb Nb 1 1

Nb

L

1

?

L 1

1 1 1 1

Curculionidae 66

Acalles misellus Weevils Acalles ptinoides Acalles roboris Acrantus vittatus Anthonomus ulmi Anthonomus bituberculatus Anthonomus pedicularius Anthonomus rubi Archarias pyrrhoceras Bagous tempestivus Barypeithes pellucidus Brachysomus hirtus Ceutorhynchus atomus Ceutorhynchus assimilis Ceutorhynchus cochleariae Ceutorhynchus typhae Cionus hortulanus Cionus scrophulariae Cleopus pulchellus Caenopsis waltoni Coelositona cambricus Curculio venosus Curculio glandium Dorytomus rufatus Euophryum confine Ellescus bipunctatus Gymnetron melanarium Gymnetron veronicae Hypera nigrirostris Hypera plantaginis Hylastinus obscurus Isochnus foliorum Ips typographus Ips sexdentatus Leiosoma deflexum Liophloeus tessulatus Magdalis barbicornis Nedyus quadrimaculatus Neliocarus nebulosus Orchestes fagi Orchestes pilosus Orchestes signifer Orchestes quercus Orchestes rusci Orobites cyaneus Orthotomicus erosus Otiorhynchus singularis Pelenomus quadrituberculatus Phloeophagus gracilis Phyllobius argentatus Phyllobius glaucus Phyllobius pomaceus Polydrusus mollis Polydrusus cervinus Polydrusus pterygomalis Ramphus oxyacanthae Rhinoncus pericarpius Sitona macularius Sitona lineatus Sitona sulcifrons Sciaphilus asperatus Scolytus scolytus Strophosoma melanogrammum Trachodes hispidus Trypodendron domesticum Trypodendron signatum

1

1

1 1 1 1 1 1 1

1

1

1 1 1 1

1 1 1

1

1

1 1 1

M NBN WT WT NBN WT WT WT WT HBRC WT WT WT WT WT WT WT WT WT NBN NBN M NBN NBN WT, KAS NBN NBN NBN WT/ID WT NBN HBRC WT NBN WT ID NBN WT NBN M NBN NBN WT WT M NBN M NBN NBN KAS WT M NBN WT WT WT WT NBN M iRecord WT WT WT NBN NBN NBN

Dasytidae

Dasytes aeratus

M

Dermestidae

Anthrenus verbasci Attagenus pellio

ID ID

39

1 Nb L 1

Nb

1

NR-RDB3 Na L

1 1 1

L

1 1 1

1 Na 1

1 1 1 1

L 1 1 1

1

1

L

L L

1 1 1

Ctesias serra Megatoma undata Dytiscidae

Elateridae

Elmidae

KAS KAS

Agabus bipustulatus Hydroporus planus Nebrioporus elegans Oreodytes sanmarkii Oreodytes septentrionalis Stictotarsus duodecimpustulatus Acilius sulcatus Ampedus cinnabarinus Ampedus pomorum Anostirus castaneus Agriotes obscurus Agriotes pallidulus Agriotes sputator Athous haemorrhoidalis Athous bicolor Calambus bipustulatus Ctenicera pectinicornis Dalopius marginatus Denticollis linearis Melanotus castanipes Melanotus villosus Paraphotistus nigricornis Prosternon tessellatum Selatosomus incanus Stenagostus rhombeus

1 1 1 1

Click Beetles

1 1

1 1

Limnius volckmari Oulimnius tuberculatus Riolus cupreus Normandia nitens Macronychus quadrituberculatus Normandia nitens

Erotylidae

Triplax aenea Dacne rufifrons

Eucnemidae

Melasis buprestoides Eucnemis capucina Microrhagus pygmaeus

Geotrupidae

Geotrupes mutator Geotrupes stercorosus Anoplotrupes stercorosus Typhaeus tyhoeus

Gyrinidae

1 1 1

1

NBN NBN NBN NBN NBN NBN ID

1 1

NBN NBN NBN WT WT, KAS WT M ID NBN NBN WT M KAS WT NBN NBN WT iRecord, ID

1 1 1

NBN KAS KAS

Violet Dor Beetle 1 Dor Beetle Woodland Dor Beetle Minotaur Beetle

NBN WT ID ID

Orectochilus villosus Gyrinus substriatus

1

NBN ID

Haliplidae

Haliplus lineatocollis

1

NBN

Histeridae

Paromalus flavicornis Gnathoncus buyssoni Gnathoncus nannetensis Paromalus flavicornis Plegaderus dissectus

iRec KAS KAS KAS KAS

Hydraenidae

Hydraena gracilis

1

NBN

Hydrophilidae

Helophorus brevipalpis Cercyon melanocephalus Cryptopleurum minutum

1

NBN NBN NBN

Lampyridae

Lampyris noctiluca

Lathridiidae

Aridius bifasciatus Aridius nodifer Cartodere nodifer

Glow-worm

M WT WT KAS

40

1

1 1

1 1

NBN NBN NBN NBN M NBN M KAS

1

Nb Nb

L

1 NR- RDB3 Nb 1

Nb

Dienerella elongata Enicmus brevicornis Enicmus rugosus Enicmus testaceus Latridius minutus Leiodidae

WT KAS KAS KAS KAS

Anisotoma orbicularis Anisotoma humeralis Agathidium atrum Agathidium nigripenne Agathidium seminulum Amphicyllis globus Colon zebei Leiodes cinnamomea

1 1 1 1 1

Lucanidae

Dorcus parallelipipedus Sinodendron cylindricum

Lycidae

Erotides cosnardi

NBN, KAS

Malachiidae

Malachius bipustulatus

M

Melandryidae

Orchesia undulata

M

Meloidae

Meloe rugosus

Monotomidae

Rhizophagus nitidulus Rhizophagus cribratus Rhizophagus bipustulatus Rhizophagus parallelocollis Rhizophagus depressus Rhizophagus picipes

1 1 1 1 1 1

NBN NBN NBN, KAS NBN NBN NBN

Mordella holomelaena Mordellochroa abdominalis Mordellistena neuwaldeggiana

1

NBN M KAS

Mordellidae

Mycetophagidae

Nitidulidae

Oedemeridae

Lesser Stag Beetle Rhinoceros Beetle

NBN WT, KAS NBN NBN NBN M NBN KAS

1 1 1 1

Nb

M WT, KAS

Rugged Oil Beetle

NBN

Mycetophagus atomarius Mycetophagus quadripustulatus Triphyllus bicolor

WT, KAS WT, KAS KAS

Brachypterus urticae Carpophilus sexpustulatus Cryptarcha strigata Cychramus luteus Epuraea biguttata Epuraea guttata Epuraea pallescens Epuraea rufomarginata Epuraea silacea Glischrochilus hortensis Glischrochilus quadriguttatus Meligethes atramentarius Meligethes kunzei Meligethes aeneus Pocadius ferrugineus Soronia punctatissima

L Nb Nb

1 1 1 1 1

1 1 1

1

Oedemera nobilis Oedemera femoralis Oedemera lurida Ischnomera cyanea Ischnomera sanguinicollis

NR-RDBI

Nb

1 1 1 1 1 L Nb L Nb

WT NBN NBN WT NBN NBN KAS NBN KAS M NBN NBN NBN ID KAS NBN M ID, KAS WT M, KAS KAS

1

1 1

1 1

1

Nb Nb Nb

Phloiophilidae

Phloiophilus edwardsii

1

NBN

1

Platypodidae

Platypus cylindrus

1

NBN

1

Ptiliidae

Ptinella limbata

1

NBN

1

Pyrochroidae

Pyrochroa coccinea

M

41

Pyrochroa serraticornis

M

Rhynchitidae

Involvulus cupreus Lasiorhynchites cavifrons Neocoenorrhinus minutus Neocoenorrhinus interpunctatus

Rhizophagidae

Rhizophagus dispar

Ripiphoridae

Metoecus paradoxus

1

NBN

1

Salpingidae

Salpingus planirostris Salpingus ruficollis Scaphisoma agaricinum Vincenzellus ruficollis

1

NBN KAS KAS NBN

1

Scarabaeidae

Aphodius ater Aphodius borealis Aphodius prodromus Aphodius sticticus Cetonia aurata Melolontha melolontha Onthophagus coenobita Onthophagus joannae Onthophagus similis Phyllopertha horticola Serica brunnea Trichius fasciatus

1 1 1 1

NBN NBN NBN NBN WT

1 Scarab Beetles

M WT ID WT M M M M M M ID HBRC

Rose Chafer Cockchafer

Garden Chafer Bee Chafer

Scirtidae

Prionocyphon serricornis Hylesinus crenatus Xyleborus monographus

WT KAS KAS

Scraptiidae

Anaspis frontalis Anaspis garneysi Anaspis lurida Anaspis maculata Anaspis pulicaria Anaspis regimbarti

WT KAS KAS WT, KAS WT, KAS WT, KAS

Scydmaenidae

Cephennium gallicum Stenichnus bicolor

Sphindidae

Aspidiphorus orbiculatus Sphindus dubius

Silphidae

Silpha atrata Silpha obscura Dendroxena quadrimaculata Nicrophorus vespilloides Nicrophorus investigator Nicrophorus humator Necrodes littoralis

Black Snail Beetles

Atheta hypnorum Atrecus affinis Bisnius puella Bledius subterraneus Bolitobius cingulatus Bryaxis curtisii Coryphium angusticolle Dasygnypeta velata Dropephylla gracilicornis Euplectus mutator Euplectus tholini Gyrohypnus atratus Gabrius splendidulus Hapalaraea pygmaea Hydrosmecta longula Ischnopoda leucopus Lamprinodes saginatus

Rove Beetles

Staphylinidae 52

1 1 1 1

1

Black Sexton Beetle

1 1 1 1 1 1 1

1

1 1 1

42

L L L L L L Nb

1

L Nb

M NBN NBN M ID ID ID NBN KAS NBN NBN NBN NBN NBN NBN KAS KAS NBN HBRC iRecord KAS NBN NBN NBN

1

Nb

NBN KAS WT, KAS WT

1 1 Common Burying Beetle

L

1 1

1 1 1 1 1 Nb Nb Nb

1 1

1 1 1

Lathrobium longulum Leptacinus formicetorum Lesteva sicula sl.

Medon brunneus Metopsia gallica Mniusa incrassata Micropeplus staphylinoides Neobisnius prolixus Ocypus bruniceps ag Ocypus olens Oxypoda soror Phloeostiba plana Phyllodrepa floralis Philonthus politor Platydracus fulvipes Platydracus stercorarius Quedius boops Quedius cruentus Quedius mesomelinus Quedius nigriceps Quedius scitus Quedius truncicola Silusa rubiginosa Scaphisoma agaricinum Sepedophilus testaceus Staphylinus olens Stenus brunnipes Stenus canaliculatus Stenus guttula Stenus impressus Stenus nitidulus Tachinus humeralis Tachyporus hypnorum Tachyporus solutus Thamiaraea hospita Tenebrionidae

1 1 1 Devil’s Coach Horse 1 1 1 1

1 1

1 1

1

Corticeus fraxini Gonodera luperus Isomira murina Lagria hirta Pseudocistela ceramboides Prionychus ater

Throscidae

Trixagus obtusus

Trogositidae

Thymalus limbatus

Zopheridae

Synchita variegata

65 families

1

1

NBN WT ID NBN WT NBN M NBN ID M NBN NBN NBN ID NBN ID WT WT, KAS KAS M NBN KAS NBN WT HBRC ID WT NBN NBN WT WT WT WT ID NBN NBN KAS WT ID ID, KAS KAS

1 L 1 1 1

1 1

1

L 1 Nb 1 L

1

1 1 Nb

Nb

WT 1

NBN

1

KAS

Species Total = 452

Site ref only SO51 = 151

43

Only pre 1970 = 122

3.5 - Checklist of Order Diptera – True Flies status at September 2021 Nematophora

Lower Diptera

Tipulidae

(21-23 of 87)

Recorder ref.

Status

Nephrotoma appendiculata Spotted Cranefly NBN C Nephrotoma flavipalpis Tiger Cranefly ID C Nephrotoma flavescens Primrose Cranefly ID U Nephrotoma quadrifaria n/a M C Tipula paludosa Common Daddy-long-legs ID A Tipula oleracea n/a ID A Tipula lunata n/a ID C Tipula fulvipennis n/a ID C Tipula flavolineata n/a ID U Tipula rufina n/a ID C Tipula pabulina / pruinosa n/a ID/PB U Tipula confusa n/a ID C Tipula maxima Giant Cranefly ID U Tipula submarmorata n/a ID U Tipula staegeri n/a ID U Tipula vernalis n/a ID U Tipula vittata n/a ID C Tipula lateralis n/a ID C Tipula pagana n/a ID C Ctenophora pectinicornis n/a ID U Note: Tipula couckei, T. alpina reported in SO51 by NBN. Also Tanyptera atrata in 1944.

Cylindrotomidae

(0-1 of 4) Note: NBN reports Cylindrotoma distinctissima from England part of SO51

Pediciidae (1-2 of 20) Ula mollissima n/a Note: NBN reports Pedicia rivosa from England part of SO51.

NBN

U

Limoniidae (14-22 of 224) Limonia nubeculosa n/a Epiphragma ocellare n/a Austrolimnophila ochracea n/a Elliopteroides alboscutellatus n/a Rhipidia maculata n/a Molophilus sp n/a Dicranomyia chorea n/a Dicranophragma adjuctum n/a Antocha vitripennis n/a Dicranomyia modesta ag. n/a Achyrolimonia decemmaculata n/a Rhypholophus bifurcates n/a Tasiocera murina n/a Lonchoptera lutea n/a Note: A further 8 species reported by NBN in SO51.

M ID ID ID ID ID ID ID ID ID NBN NBN other 1 other 1

C U A U U C C U U U U U

Biblionidae (4 of 18) Biblio marci Biblio johannis Biblio pomonae Dilophus febrilis

St Marks fly n/a n/a Feverfly

ID ID ID ID

C C U U

Bolitophilidae (2 of 16) Bolitophila saundersii Bolitophila occlusal

n/a n/a

NBN NBN

U U

44

Diadocidiidae (2 of 3) Diadocidia ferruginosa Diadocidia spinosula

n/a n/a

NBN NBN

Ditomyiidae (1 of 3) Symmerus annulatus

fungus gnat

M

Keroplatidae (10-11 of 51) Macrocera angulata fungus gnat sp.

Macrocera fasciata n/a Macrocera stigmoides n/a Macrocera vitatta n/a Macrocera centralis n/a Macrocera inversa n/a Macrocera phalaerata n/a Macrocera vittate n/a Platyura marginata n/a Keroplatus testaceus n/a Note: Pyratula zonata also reported for SO51 by NBN Mycetophilidae. (45 of 500) Allodia lundstroemi Allodia ornaticollis Anatella minuta Boletina sciarina Brevicornu griseicolle Cordyla crassicornis Cordyla fissa Cordyla insons Exechia bicincta Exechia cincta Exechia contaminate Exechia dizona Exechia dorsalis Exechia festiva Exechia fusca Exechiopsis intersecta Exechiopsis jenkinsoni Exechiopsis pollicata Exechiopsis subulate Mycetophila Britannica Mycetophila curviseta Mycetophila edwardsi Mycetophila fungorum Mycetophila hetschkoi Mycetophila ichneumonea Mycetophila marginata Mycetophila ocellus Mycetophila perpallida Mycetophila pumila Mycetophila ruficollis Mycetophila spectabilis Mycetophila trinotata Mycomya flavicollis Mycomya prominens Phronia biarcuata Phthinia winnertzi Pseudobrachypeza Helvetica Rymosia spinipes Sceptonia cryptocauda Stigmatomeria crassicornis

fungus gnat n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a

45

ID NBN NBN NBN/other 1 other 1 other 1 other 1 other 1 M M

NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN

A U

Synplasta gracilis Zygomyia pictipennis Zygomyia semifusca Zygomyia valida Zygomyia vara

n/a n/a n/a n/a n/a

NBN NBN NBN NBN NBN

(1 of 267) Sciara hemerobiodes ag.

Black Fungus Gnat

ID

U

Cecidomyiidae.

(1-2 of 635) Hartigiola annulipes Hairy Beech Gall Note: Jaapiella veronicae reported in SO51 England by NBN

ID

C

Psychodidae (0-3 of 99) Pericoma sp n/a Note: at least 2 other undetermined species

ID

A

M ID M M M

C U U U U

Anisopodidae (1-2 of 4) Sylvicola fenestralis Window Gnat Note: Sylvicola cinctus reported by NBN in SO51

M

A

Ptychopteridae (3-4 of 7) Ptychoptera contaminata A Fold-winged Cranefly Ptychoptera albimana n/a Ptychoptera lacustris n/a Note: Ptychoptera Longicauda reported in SO51 by NBN.

ID ID ID

U U U

mosquito n/a Tee-hole Mosquito

ID ID ID

C C C

Simuliidae Simulium posticatum sl.

Blandford Bomber Note: at least 1 other undetermined species

ID

C

n/a

ID

U

Xylophagidae (0-1 of 3) Xylophagus ater

Common Awl-fly

M

C

Rhagonidae (5 of 15) Chrysopilus asiliformis Chrysophilus cristatus Rhagio scolopacea Rhagio tringarius Rhagio annulatus

Little Snipefly Black Snipefly Downlooker Snipefly Marsh Snipefly n/a

M ID M M NBN

Tabanidae (7-8 of 30) Haematopota pluvialis Haematopota crassicornis Tabanus bromius

Notch-horned Cleg n/a Band-eyed Brown Horsefly

M NBN M

Sciaridae

Trichoceridae (5 of 12) Trichocera annulata Trichocera parva Trichocera saltator Trichocera regelationis Trichocera major

Winter Gnat n/a n/a n/a n/a

Culicidae.

Culex pipiens Culiceta annulata Dahliana geniculata

Chironomidae.

(1 of 630) Microtendipes pedellus

Brachycera

46

A

Tabanus sudeticus Dark Giant Horsefly Tabanus cordiger Hybomitra distinguenda Bright Horsefly Chrysops caecutiens Splayed Deerfly Note: Chrysops viduatus reported by NBN for SO51 Atheracidae

(0-2 of 3) Note: Atherix ibis and Atricops crassipes reported by NBN from SO51

Stratiomyidae (7-8 of 48) Chloromyia formosa Broad Centurion Sargus bipunctatus Twin-spot Centurion Sargus iridatus n/a Chorisops nagatomii Bright Four-spined Centurion Chorisops tibialis Dull Four-spined Centurion Beris chalybata (poss B. morrissii) n/a Pachygater atra n/a Microchrysa polita n/a Note: Oxycera pygmaea reported by NBN in SO51 England Bombyliidae (4 of 10) Bombylius major Bombylius discolor Bombylius canescens Villa cingulata Therevidae

Asilidae

ID NBN anon ID ID

Dark-edged Bee-fly Dotted Bee-fly Western Bee-fly Downland Villa

M ID ID ID ID M ID ID

ID ID ID ID

(0-3 of 14) Note: NBN reports 3 species from SO51

(3-4 of 29) Machimus atricapillus Kite-tailed Robberfly Leptogaster cylindrica Striped Slender Robberfly Diorctria linearis Small Yellow-legged Robberfly Note: Dioctria oelandica reported by NBN in SO51 England

Hybotidae (18-21 of 182) Hybos grossipes n/a Hybos femorata n/a Hybos culiciformis n/a Bicellaria nigra n/a Ocydromia glabricula n/a Euthyneura gyllenhali n/a Leptopeza flavipes n/a Ocydromia glabricula n/a Oedalea holmgreni n/a Oedalea stigmatella n/a Oropezella sphenoptera n/a Platypalpus incertus n/a Platypalpus nigritarsis n/a Platypalpus major n/a Platypalpus pallidiventris n/a Platypalpus luteolus n/a Platypalpus subtilis n/a Trichinomyia flavipes n/a Note: Another 3 are reported by NBN in SO51 Empididae (7-18 of 208) Empis tessellata Emphis stercorea Emphis lutea Empis nigritarsis Emphis livida Phyllodromia melanocephala

A Dance Fly n/a n/a n/a n/a n/a

47

M M M

ID NBN NBN other 1 NBN other 1 other 1 other 1 other 1 other 1 other 1 other 1 other 1 other 1 NBN NBN NBN NBN

M M ID other 1 NBN other 1

R C C

Rhamphomyia dentipes n/a Note NBN reports a further 11 species in England part of SO51.

Dolichopodidae (10-21 of 308) Xanthochlorus tenellus (tent).

Long-legged Fly sp Microphor sp n/a Poecilobothrus nobilitatus n/a Neurigona pallida n/a Neurigona quadrifasciata n/a Rhaphium crassipes n/a Campsicnemus curvipes n/a Xanthochlorus ornatus n/a Dolichopus ungulates n/a Sybistroma obscurellum n/a Note: NBN reports an addition 11 species in SO51 Platypezidae (2 of 34) Agathomyia sp Platypeza consobrina sl Phoridae

(1 of 356) possible Megaselia sp

ID ?

ID ?

ID M M M NBN NBN NBN NBN

U U C U C C

Flat-footed Fly sp n/a

ID ID

U U

n/a

ID

Lonchopteridae (1-2 of 7) Lonchoptera bifurcate n/a Note: NBN reports Lonchoptera lutea in SO51 Syphidae

(83 of 283) Baccha elongata Melanostoma mellinum Melanostoma scalare Platycheirus albimanus ag Platycheirus angustatus ag Platycheirus rosarum Platycheirus scutatus ag Paragus haemorrhous Syrphus ribesii Syrphus torvus Sphaerophoria interrupta ag Sphaerophoria scripta Meliscaeva auricollis Meliscaeva cinctella Melangyna umbellatarum Melangyna compositarum/labiatum Melanygna lasiophthalma Didea fasciata Chrysotoxum bicinctum Chrysotoxum festivum Chrysotoxum cautum Leucozona lucorum Leucozona glaucia Epistrophe eligans Epistrophe nitidicollis Epistrophe grossulariae Episyrphus balteatus Eupeodes luniger Eupeodes latifasciatus Eupeodes corollae Dasysyrphus albostriatus Dasysyrphus venustus Scaeva selentica Scaeva pyrastri Parasyphus punctulatus

other 1

n/a n/a Chequered Hoverfly n/a n/a n/a n/a n/a Common Hoverfly Common Hoverfly n/a Long Hoverfly n/a n/a n/a n/a n/a n/a n/a Dark Wasp Hoverfly n/a n/a n/a n/a n/a n/a Marmalade Hoverfly n/a n/a n/a n/a n/a n/a n/a n/a

48

ID

ID M M M ID ID ID ID ID ID ID M ID ID ID ID ID ID ID ID ID M M ID ID ID M ID ID ID ID other 1 ID M other 1

Xanthogramma citrofasciatum Xanthogramma pedissequum Rhingia rostrata Rhingia campestris Ferdinandea cuprea Cheilosia scutellata ag Cheilosia pagana ag Cheilosia variabilis Cheilosia vulpina Cheilosia impressa Cheilosia illustrata Cheilosia albitarsus Portevinia maculata Chrysogaster solstitialis Orthonerva nobilis Melanogaster hirtella Brachypoda scutellaris Helophilus pendulus Helophilus trivittatus Eristalis pertinax Eristalis tenax Eristalis arbustorum Eristalis nemorum Eristalis horticola Eristalis intricarius Myathropa florea Eumerus striganus ag Merodon equestris Pipiza noctiluca ag Pipizella viduata sl Sericomyia silentis Arctophila superbiens Volucella pellucens Volucella bombylans Volucella inflata Volucella inanis Volucella zonaria Syritta pipiens Criorhina berberina ssp oxyacanthi Criorhina berberina typical form Criorhina floccosa Criorhina asilica Chalcosyrphus nemorum Brachypalpoides lentus Xylota jukatorum sl.

Xylota sylvarum Xylota segnis Sphegina clunipes Pipunculidae (1 of 99) Pipunculus sp

n/a Ornate Hoverfly (Heiniken Hoverfly) n/a n/a n/a n/a n/a n/a n/a n/a n/a Ramsons Hoverfly n/a n/a n/a n/a Brindled Hoverfly n/a Tapered Drone fly Dronefly Dwarf Drone fly n/a n/a Dronefly sp False Dronefly n/a n/a n/a n/a n/a n/a Pellucid Hoverfly Bumblebee Hoverfly n/a Lesser Hornet Hoverfly Hornet Hoverfly Thick-legged Hoverfly n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a

Big-headed Fly sp

.

ID ID ID ID ID ID ID M ID M NBN NBN M ID ID ID ID M M ID M ID ID ID ID ID ID ID ID ID ID ID M M M ID ID M ID ID ID ID ID M ID ID ID NBN

ID

.

U

Schizophora - Acalyptratae Micropezidae (0-2 of 10) Note: two species, Calobata petronella and Neria cibaria, are reported by NBN from the England part of SO51.

Psilidae

(1-3 of 27) Note: Three species, Psila fimetaria, Psila fimetaria, Chyliza leptogaster, are reported by NBN from the England part of SO51 and one unidentified species found by ID.

Conopidae (5 of 25) Sicus ferrugineus Physocephala rufipes

n/a n/a

ID ID

49

C U

Conops quadrifasciatus Conops vesicularis Myopa testacea Myopa pellucida Thecophora atra Lonchaeaidae (1) Lonchaea sp

n/a n/a n/a n/a n/a

ID ID ID ID ID

n/a

ID

Pallopteridae (4-5 of 13) Palloptera muliebris Looped Flutterfly ID Palloptera quinquemaculata Five-spotted Flutterfly ID Palloptera ustulata ag.

Dark-tipped Flutterfly ID Palloptera saltuum Hogweed Flutterfly ID Note: NBN reports one additional species, Palloptera modesta, in SO51

Piophilidae (1 of 14) Neottiophilum praeustum

Ulidiidae

Nest Skipper

ID

(3-4 of 20) Herina nigrina Three-spot Wingwaver ID Herina lugubris Four-spot Wingwaver ID Seioptera vibrans n/a Note:One species, Physiphora alceae, reported by NBN in England part of SO51

U U U U U

C C C U

R

C C

Tephritidae (11-12 of 77) Anomoia purmunda n/a ID Tephritis formosa n/a ID Tephritis neesii n/a ID Tephritis bardanae n/a ID Terellia tussilaginis n/a ID Trypeta zoe n/a ID Urophora quadrifasciata n/a ID Ragoletis alternata n/a ID Chaetostomella cylindrica n/a ID Euleia heraclei Celery Fly ID Oxyna parietina n/a NBN Note: One species Urophora jaceana reported by NBN from England part of SO51 Lauxaniidae (9-13 of 56) Peplomyza litura n/a M Minettia inusta n/a M Minettia longipennis n/a M Meiosimyza platycephala n/a NBN Sapromyza albiceps n/a NBN Meiosimyza rorida n/a NBN Tricholauxania praeusta n/a NBN Minettia rivosa n/a NBN Sapromyza basalis n/a NBN Sapromyza halidayi n/a NBN Sapromyza quadripunctata n/a ID Note: the following 4 additional species are reported by NBN from the England part of SO51. Calliopum simillimum, Pseudolyciella pallidiventris, Meiosimyza laeta.

Dryomyzidae (1 of 3) Dryomyza anilis n/a Note: previously known as Neuroctena anilis Sciomyzidae (9-12 of 72) Pherbellia albocostata Pherbellia cinerella

n/a n/a

M

M other 1

50

Pherbellia dubia n/a M Pherbellia scutellaris n/a M Trypetoptera punctulata Picture-winged Snail-killer ID M Coremacera marginata n/a M Dichetophora obliterata Chalk Snail-killer ID Limnia unguicornis Stripe-backed Snail-killer ID Pelidnoptera fuscipennis n/a other 1 Note: NBN reports 3 additional species in the England part of SO51. They are Pherbellia annulipes, Renocera pallida and Tetanura pallidiventris. Includes all Phaeomyiidae.

Sepsidae

(3 of 29) Nemopoda nitidula Sepsis fulgens Sepsis punctum ag

n/a n/a Black scavenger fly

NBN NBN ID

Clusiodes albimanus

a Druid fly

M

n/a n/a

ID ID

Clusiidae

Agromyzidae (2 of 420) Phytomyza ranuncluli Agromyza lithospermi

Opomyzidae (2-3 of 16) Geomyza tripunctata n/a NBN Opomyza florum n/a NBN Note: NBN reported Opomyza germinationis in England part of SO51 Chloropidae (1 of 178) Chlorops sp

n/a

ID

Heleomyzidae (1 of 56) Suillia variegata sl.

Tephrochlamys tarsalis Scoliocentra villosa

n/a n/a n/a

ID ID ID

Drosophilidae (4-5 of 65) Drosophila andalusiaca n/a Drosophila subobscura n/a Drosophila fenestrarum n/a Scaptomyza pallida n/a Note: NBN reports Scaptomyza flava from SO51

NBN NBN NBN NBN

Campichoetidae (1 of 2) Campichoeta punctum

n/a

NBN

Diastatidae (1 of 6) Diastata fuscula

n/a

ID

Ephyhydridae (1-3 of 151) Hydrellia maura n/a NBN Note: NBN reported Ditrichophora calceata and Parydra coarctata in England part of SO51 Schizophora - Calyptratae Scathophagidae (2 of 55) Scathophaga stercoraria ag.

Cordilura pudica

Yellow Dung Fly n/a

M ID

Anthomyiidae (9 of 246) Pegomya hyoscyami Pegomya bicolor Anthomyia procellaris Anthomyia pluvialis

n/a n/a n/a n/a

ID ID ID ID

51

Hylemya vegans Hylemya variata Hylemya nigrimana Adia cinerella Delia platura

n/a n/a n/a n/a n/a

M ID NBN ID ID

(5 of 61) Fannia coracina Fannia fuscula sl Fannia canicularis Fannia mollisima Fannia lustrator sl

n/a n/a Lesser Housefly n/a n/a

NBN ID ID ID ID

(26 of 293) Musca domestica Musca autumnalis Muscina prolapsa Mesembrina meridiana Graphomya maculata Phaonia errans Phaonia valida Phaonia rufiventris Phaonia subventa Phaonia pallida Phaonia tugoriorum Phaonia palpata Helina reversio Helina impuncta Helina evecta Thricops diaphanus Thricops semicinereus Eudasyphora cyanella Polietes lardarius Neomyia viridescens Neomyia cornicina Morellia simplex Coenosia agromyzina Coenosia mollicula Hebecnema nigra Hydrotaea irritans

Common Housefly Face Fly n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a (n/a tentative) n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a

ID ID ID M M ID ID M M M ID NBN ID M NBN M ID ID M ID ID ID NBN NBN NBN NBN

Calliphoridae (8 of 30) Calliphora vomitoria Calliphora vicina Lucilia caesar ag.

Protocalliphora azurea Eurychaeta palpalis Melinda viridicyanea Melinda gentilis Bellardia vulgaris

Orange-bearded Bluebottle Common Bluebottle Greenbottle Bird Blowfly False Fleshfly Dark-palped Melinda Pale-palped Melina Pale-veined Emerald-bottle

ID ID M ID ID ID ID ID

Pollenidae (4 of 8) Pollenia pediculata Pollenia armentaria Pollenia griseotomentosa Pollenia rudis

Tufted Clusterfly Black-bellied Clusterfly Little Clusterfly Common Clusterfly

ID ID ID ID

Rhinophoridae (1 of 8) Paykullia maculata

Woodlouse Fly

ID

Sarcophagidae (3 of 60) Scathophaga carnaria ag.

Scathophaga stercoraria

Fleshfly n/a

M NBN

Fanniidae

Muscidae

52

Ravinia pernix Tachinidae (17 of 270) Tachina fera Tachina ursina Nemorilla floralis Nowickia ferox Eriothrix rufomaculata Lophosia fasciata Phasia hemiptera Eurithia anthopila Epicampocera succinata Exorista larvarum Gymnochaeta viridis Hemyda vittata Phania fenestra sl Siphona geniculata Dexiosoma caninum Chrysosomopsis auratus Winthemia quadripustulata

n/a

ID

Tachinid fly n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a

ID ID ID ID ID ID M ID ID ID ID ID ID ID M ID NBN

53

3.6 Order Lepidoptera – Butterflies and Moths Checklist of Butterflies (Order Lepidoptera) of the Doward. April 2022.

Local Status* Pieridae Large White Small White Green-veined White Wood White Orange tip Brimstone Clouded Yellow

Pieris brassicae Pieris rapae Pieris napi Leptidea sinapis Anthocharis cardamines Gonepteryx rhamni Colias croceus

Common Common Common Present in small numbers. (last seen 2022) Common Present in small numbers.

Occasional

Nymphalidae Peacock Small Tortoiseshell Red Admiral White Admiral Comma Painted Lady Silver-washed Fritillary Dark-green Fritillary

Inachis io Aglais urticae Vanessa atalanta Limentitis camilla Polygonia c-album Cynthia cardui Argynnis paphia Argynnis aglaja

Common Common Common Present in very small numbers. (last seen 2020) Common Common visitor Common Occasional visitor

Satyridae Marbled White Ringlet Gatekeeper Speckled Wood Meadow Brown

Melanargia galathea Aphantopus hyperantus Pyronia Tithonus Pararge aegeria Manila jurtina

Common Common Common Common Common

Lycaenidae Common Blue Holly Blue Brown Argus Small Copper Purple Hairstreak White-letter Hairstreak

Polyommatus icarus Celastrina argiolus Aricia agestis Lycaena phlaeas Quercusia quercus Satyrium w-album

Common Common Present in very small numbers. (Present 2023) Frequent Present in small numbers.

Present in small numbers. (Present 2023)

Hesperiidae Large Skipper Small Skipper Essex Skipper Dingy Skipper Grizzled Skipper

Thymelicus sylvestris Thymelicus flavus Thymelicus lineola Erynnis tages Pyrgus malvae

Common Common Common Present in very small numbers. (Last seen 2022) Frequent (late April – mid June)

31 species * Note on Local Status: The comment refers to status during the flight period (which may be short) and in suitable habitat.

Other Historical or uncertain records Pearl-bordered Fritillary Boloria euphrosyne Small Pearl-bordered Fritillary Boloria selene Wall Brown Lasiommata megera Green Hairstreak Callophrys rubi Small Heath Coenonympha pamphilus Grayling Hipparchia semele Purple Emperor Apatura iris High-brown Fritillary Argynnis adippe

Formerly present. last report 2014. Could reappear.

Past status unclear. Could reappear.

Past status unclear. Could reappear.

Past status unclear. Could reappear.

Past status unclear. Could reappear.

Past status unclear. Now rare inland.

Last report 1976 Last report 1976

54

Checklist of Moths. Last update August’22 The basis of this list is an extract from the county database held by the County Recorder, Peter Hall and provided in Jan.’21.

Additionally, a few species (marked with an *) have been added to this list as they have been accepted by PH since that date or their identification is not in doubt. This list does not include records from Whitchurch.

Family

Code

Taxon

Vernacular

National Status

1.001 1.003 1.004 1.005

Micropterix tunbergella Micropterix aureatella Micropterix aruncella Micropterix calthella

n/a n/a n/a n/a

L L C C

2.001 2.005 2.006

Dyseriocrania subpurpurella Eriocrania salopiella Eriocrania cicatricella

n/a n/a n/a

C L L

3.001 3.002 3.004 3.005

Triodia sylvina Korscheltellus lupulina Phymatopus hecta Hepialus humuli

Orange Swift Common Swift Gold Swift Ghost Moth

C C C C

4.002 4.003 4.004 4.011 4.013 4.014 4.018 4.019 4.026 4.031 4.034 4.040 4.042 4.045 4.056 4.058 4.062 4.065 4.071 4.075 4.079 4.089 4.095 4.096

Stigmella lapponica Stigmella confusella Stigmella tiliae Stigmella prunetorum Stigmella malella Stigmella catharticella Stigmella ulmivora Stigmella viscerella Stigmella oxyacanthella Stigmella mespilicola Stigmella tityrella Stigmella assimilella Stigmella plagicolella Stigmella aurella Stigmella speciosa Stigmella basiguttella Stigmella samiatella Trifurcula cryptella Bohemannia pulverosella Ectoedemia louisella Ectoedemia atrifrontella Ectoedemia albifasciella Ectoedemia atricollis Ectoedemia arcuatella

n/a n/a n/a n/a Apple Pygmy n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a

C C L Nb L L L L C pRDB2 C L C C C C L Na C L Unkn C C Na

5.004

Pseudopostega crepusculella

n/a

Nb

6.001 6.002

Antispila metallella Antispila petryi

n/a n/a

L L (A. treitschkeilla)

7.001 7.002 7.006 7.008 7.009 7.012 7.014 7.015

Nemophora degeerella n/a Nemophora metallica n/a Adela reaumurella n/a Adela croesella n/a Cauchas fibulella n/a Nematopogon schwarziellus n/a Nematopogon metaxella n/a Nematopogon swammerdamella n/a

C Nb C L C C L C

8.001 8.002 8.004 8.005

Incurvaria pectinea Incurvaria masculella Incurvaria praelatella Phylloporia bistrigella

n/a n/a n/a n/a

L C L L

10.001 10.002 10.003

Tischeria ekebladella Tischeria dodonaea Coptotriche marginea

n/a n/a n/a

C L C

Micropteridae

Eriocranidae

Hepalidae - Swifts

Nepticulidae

Opostegidae Heliozelidae

Adelidae

Incurvariidae

Tischeriidae

55

10.006

Coptotriche angusticollella

n/a

Nb (Emmetia a.)

11.004 11.006 11.010 11.012

Dahlica inconspicuella Taleporia tubulosa Bacotia claustrella Psyche casta

Lesser Lichen Case-bearer n/a n/a n/a

Na L pRDB3 C

12.010 12.012 12.016 12.017 12.019 12.021 12.027 12.032 12.033 12.035 12.036 12.037 12.038 12.048

Morophaga choragella Triaxomera parasitella Nemapogon cloacella Nemapogon koenigi Nemapogon ruricolella Nemapogon clematella Tinea pellionella Tinea semifulvella Tinea trinotella Niditinea striolella Monopis laevigella Monopis weaverella Monopis obviella Psychoides filicivora

n/a n/a Cork Moth n/a n/a n/a Case-bearing Clothes Moth n/a n/a n/a Skin Moth n/a n/a n/a

L L C Nb (N. wolfiella) Nb L L C C Nb C C L L

13.002

Roeslerstammia erxlebella

n/a

L

14.002 14.006 14.009 14.010 14.013

Bucculatrix nigricomella Bucculatrix frangutella Bucculatrix thoracella Bucculatrix ulmella Bucculatrix demaryella

n/a n/a n/a n/a n/a

L L C C L

15.002 15.003 15.005 15.006 15.008 15.009 15.01 15.012 15.014 15.015 15.016 15.022 15.025 15.026 15.028 15.029 15.032 15.034 15.035 15.036 15.039 15.04 15.043 15.045 15.047 15.049 15.051 15.052 15.058 15.059 15.063 15.064 15.073 15.074 15.075 15.076 15.078 15.08 15.081 15.082

Caloptilia cuculipennella n/a Caloptilia populetorum n/a Caloptilia betulicola n/a Caloptilia rufipennella n/a Caloptilia alchimiella n/a Caloptilia robustella n/a Caloptilia stigmatella n/a Caloptilia semifascia n/a Gracillaria syringella n/a Aspilapteryx tringipennella n/a Euspilapteryx auroguttella n/a Callisto denticulella n/a Parornix betulae n/a Parornix fagivora n/a Parornix anglicella n/a Parornix devoniella n/a Parornix finitimella n/a Phyllonorycter harrisella n/a Phyllonorycter roboris n/a Phyllonorycter heegeriella n/a Phyllonorycter quercifoliella n/a Phyllonorycter messaniella n/a Phyllonorycter oxyacanthae n/a Phyllonorycter mespilella n/a Phyllonorycter hostis n/a Phyllonorycter spinicolella n/a Phyllonorycter lantanella n/a Phyllonorycter corylifoliella n/a Phyllonorycter hilarella n/a Phyllonorycter cavella n/a Phyllonorycter maestingella n/a Phyllonorycter coryli Nut Leaf Blister Moth Phyllonorycter lautella n/a Phyllonorycter schreberella n/a Phyllonorycter ulmifoliella n/a Phyllonorycter emberizaepenella n/a Phyllonorycter tristrigella n/a Phyllonorycter froelichiella n/a Phyllonorycter nicellii n/a Phyllonorycter klemannella n/a

Psychidae

Tineidae

Roeslerstammiidae Bucculatricidae

Gracillariidae

56

Na L C C C C C L C C C C C L C C C C Nb C C C C Nb L C L C C L C C L C C L C C C C

15.084 15.089 15.092

Phyllonorycter acerifoliella Cameraria ohridella Phyllocnistis unipunctella

n/a n/a n/a

C C C

16.001 16.002 16.005 16.007 16.01 16.014 16.015 16.017 16.018 16.02 16.021 16.022 16.024

Yponomeuta evonymella Bird-cherry Ermine Yponomeuta padella Orchard Ermine Yponomeuta rorrella Willow Ermine Yponomeuta plumbella n/a Zelleria hepariella n/a Pseudoswammerdamia combinella n/a Swammerdamia caesiella n/a Swammerdamia pyrella n/a Swammerdamia compunctella n/a Paraswammerdamia nebulella n/a Cedestis gysseleniella n/a Cedestis subfasciella n/a Ocnerostoma friesei n/a

C C L L L C C C Na C L L L

17.001 17.002 17.003 17.005 17.01 17.011 17.012

Ypsolopha mucronella Ypsolopha nemorella Ypsolopha dentella Ypsolopha scabrella Ypsolopha parenthesella Ypsolopha ustella Ypsolopha sequella

n/a n/a Honeysuckle Moth n/a n/a n/a n/a

L L C C C C C

18.001 18.003 18.007

Plutella xylostella Plutella porrectella Eidophasia messingiella

Diamond-back Moth n/a n/a

M C L

19.002 19.003 19.006 19.007 19.009 19.014

Glyphipterix thrasonella Glyphipterix fuscoviridella Glyphipterix forsterella Glyphipterix simpliciella Digitivalva perlepidella Acrolepia autumnitella

n/a n/a n/a Cocksfoot Moth n/a n/a

C C L C Na C

20.001 20.004 20.01 20.011 20.012 20.013 20.014 20.016 20.018 20.019 20.020 20.022 20.023 20.024

Argyresthia laevigatella Argyresthia arceuthina Argyresthia ivella Argyresthia brockeella Argyresthia goedartella Argyresthia pygmaeella Argyresthia sorbiella Argyresthia retinella Argyresthia spinosella Argyresthia conjugella Argyresthia semifusca Argyresthia bonnetella Argyresthia albistria Argyresthia semitestacella

n/a n/a n/a n/a n/a n/a n/a n/a n/a Apple Fruit Moth n/a n/a n/a n/a

L Nb Nb C C C L C C C C C C L

21.001 21.004 21.008

Lyonetia clerkella Leucoptera laburnella Leucoptera malifoliella

Apple Leaf Miner Laburnum Leaf Miner Pear Leaf Blister Moth

C C C

22.002

Prays fraxinella

Ash Bud Moth

C

27.001

Oegoconia quadripuncta

n/a

C

28.008 28.009 28.010 28.012 28.013 28.014 28.015 28.019 28.020 28.022 28.024

Metalampra italica n/a Endrosis sarcitrella White-shouldered House-moth Hofmannophila pseudospretella Brown House-moth Borkhausenia fuscescens n/a Crassa tinctella n/a Crassa unitella n/a Batia lunaris n/a Esperia sulphurella n/a Dasycera oliviella n/a Alabonia geoffrella n/a Tachystola acroxantha* Ruddy Streak

Yponomeutidae

Ypsolophidae

Plutellidae

Glyphipterigidae

Argyresthiidae

Lyonetidae

Praydidae Autostichidae Oecophoridae

57

Mc C C C Nb C (Batia) C C Na (Esperia) C L

Chimbachidae 29.001

Diurnea fagella

n/a

C

30.002 30.003

Agnoea subochreella Agnoea josephinae

n/a n/a

L (Pseudatemelia) L

31.001

Carcina quercana

n/a

C

32.001 32.002 32.007 32.008 32.009 32.013 32.015 32.017 32.018 32.019 32.024 32.026 32.029 32.031 32.032 32.038 32.042 32.044 32.045 32.05

Semioscopis avellanella Semioscopis steinkellneriana Agonopterix ocellana Agonopterix liturosa Agonopterix purpurea Agonopterix carduella Agonopterix subpropinquella Agonopterix arenella Agonopterix heracliana Agonopterix ciliella Agonopterix assimilella Agonopterix kaekeritziana Agonopterix umbellana Agonopterix alstromeriana Agonopterix angelicella Depressaria badiella Depressaria pulcherrimella Depressaria douglasella Depressaria albipunctella Telechrysis tripuncta

n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a

L L C L L Nb C C C C L L L C L L L Nb Na Nb

34.001 34.004

Pancalia leuwenhoekella Limnaecia phragmitella

n/a n/a

L C

35.003 35.004 35.011 35.012 35.017 35.018 35.02 35.025 35.026 35.027 35.028 35.031 35.040 35.046 35.047 35.058 35.060 35.065 35.080 35.081 35.085 35.093 35.107 35.109 35.123 35.141 35.143 35.145 35.147 35.148 35.151 35.153 35.157 35.159 35.160 35.161

Aproaerema larseniella Aproaerema taeniolella Anacampsis populella Anacampsis blattariella Neofaculta ericetella Hypatima rhomboidella Anarsia spartiella Dichomeris alacella Acompsia cinerella Acompsia schmidtiellus Brachmia blandella Helcystogramma rufescens Bryotropha terrella Bryotropha senectella Bryotropha affinis Metzneria metzneriella Apodia bifractella Monochroa cytisella Eulamprotes unicolorella Eulamprotes atrella Athrips mouffetella Mirificarma mulinella Psoricoptera gibbosella Scrobipalpa acuminatella Scrobipalpa costella Teleiodes vulgella Teleiodes luculella Teleiodes sequax Carpatolechia decorella Carpatolechia fugitivella Carpatolechia proximella Pseudotelphusa paripunctella Recurvaria leucatella Exoteleia dodecella Stenolechia gemmella Parachronistis albiceps

n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a

L (Syncopacma) L “ C L C C L Nb L Nb C C C L C C L L Nb L C C L C C C C L Nb L C Nb L L L L

36.001

Batrachedra praeangusta

n/a

C

Lypusidae

Peliopodidae Depressariidae

Cosmopterigidae

Gelechiidae

Batrachedridae

58

Coleophoridae 37.006 37.007 37.01 37.011 37.012 37.015 37.022 37.028 37.032 37.044 37.046 37.048 37.049 37.051 37.052 37.053 37.061 37.063 37.066 37.069 37.072 37.073 37.080 37.102 37.103 37.106

Coleophora gryphipennella Coleophora flavipennella Coleophora alnifoliae Coleophora badiipennella Coleophora limosipennella Coleophora serratella Coleophora lusciniaepennella Coleophora juncicolella Coleophora albitarsella Coleophora discordella Coleophora deauratella Coleophora mayrella Coleophora anatipennella Coleophora kuehnella Coleophora ibipennella Coleophora betulella Coleophora lixella Coleophora albicosta Coleophora laricella Coleophora caespititiella Coleophora otidipennella Coleophora alticolella Coleophora virgaureae Coleophora argentula Coleophora follicularis Coleophora paripennella

n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a Pistol Case-bearer n/a n/a n/a n/a n/a Larch Case-bearer n/a n/a n/a n/a n/a n/a n/a

C C Nb L L C C L L C L C L L L L Nb C C C L C Nb C L L

38.003 38.004 38.005 38.008 38.012 38.015 38.017 38.022 38.023 38.025 38.029 38.030 38.033 38.036 38.037 38.038 38.041

Stephensia brunnichella Elachista argentella Elachista triatomea Elachista triseriatella Elachista obliquella Elachista gangabella Elachista adscitella Elachista gleichenella Elachista biatomella Elachista atricomella Elachista luticomella Elachista albifrontella Elachista subnigrella Elachista humilis Elachista canapennella Elachista rufocinerea Elachista cinereopunctella

n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a

Nb C L Na Nb L Nb L Nb C L C Nb C C C Nb

39.001 39.002 39.003

Blastodacna hellerella Blastodacna atra Spuleria flavicaput

n/a Apple Pith Moth n/a

C Nb L

40.002 40.004 40.005 40.008 40.011 40.012 40.015

Mompha ochraceella Mompha propinquella Mompha divisella Mompha subbistrigella Mompha langiella Mompha miscella Mompha raschkiella

n/a n/a n/a n/a n/a n/a n/a

C L Na C L L C

41.002 41.003

Blastobasis adustella Blastobasis lacticolella

n/a n/a

C C

43.002 43.004

Scythris grandipennis Scythris picaepennis

n/a n/a

Nb Nb

44.001

Alucita hexadactyla

Many-plumed Moth

C

45.004 45.010 45.011 45.012 45.013

Platyptilia gonodactyla Amblyptilia acanthadactyla Amblyptilia punctidactyla Stenoptilia pterodactyla Stenoptilia bipunctidactyla

Triangle Plume Beautiful Plume Brindled Plume Brown Plume Twin-spot Plume

C C L C C

Elachistidae

Parametriotidae

Momphidae

Blastobasidae

Scythridae

Alucidae Pterophoridae

59

45.021 45.023 45.025 45.028 45.030 45.031 45.033 45.034 45.037 45.038 45.039 45.041 45.043 45.044

Stenoptilia zophodactylus Marasmarcha lunaedactyla Oxyptilus parvidactyla Capperia britanniodactylus Pterophorus pentadactyla Porrittia galactodactyla Merrifieldia leucodactyla Merrifieldia baliodactylus Oidaematophorus lithodactyla Hellinsia tephradactyla Hellinsia carphodactyla Hellinsia osteodactylus Adaina microdactyla Emmelina monodactyla

Dowdy Plume Crescent Plume Small Plume Wood Sage Plume White Plume Spotted-white Plume Thyme Plume Dingy White Plume Dusky Plume Plain Plume Citron Plume Small Goldenrod Plume Hemp Agrimony Plume Common Plume

L L Nb Nb C Nb (Pterophorus) L Nb L Nb (Euleioptilus) L (Euleioptilus) L C C

46.001

Schreckensteinia festaliella

n/a

C

47.001 47.005

Phaulernis fulviguttella Epermenia chaerophyllella

n/a n/a

C C

48.001 48.007

Anthophila fabriciana Choreutis pariana

Nettle-tap Apple Leaf Skeletonizer

C L

49.001 49.002 49.004 49.005 49.009 49.013 49.015 49.018 49.021 49.022 49.023 49.024 49.025 49.026 49.028 49.031 49.038 49.039 49.040 49.044 49.047 49.049 49.050 49.051 49.058 49.059 49.060 49.062 49.063 49.066 49.067 49.069 49.070 49.071 49.073 49.076 49.077 49.078 49.079 49.080 49.082 49.083 49.084 49.085 49.087 49.090 49.091 49.095

Olindia schumacherana Isotrias rectifasciana Ditula angustiorana Epagoge grotiana Capua vulgana Archips podana Archips xylosteana Choristoneura hebenstreitella Ptycholomoides aeriferana Ptycholoma lecheana Pandemis cinnamomeana Pandemis corylana Pandemis cerasana Pandemis heparana Syndemis musculana Zelotherses paleana Clepsis consimilana Epiphyas postvittana Lozotaeniodes formosana Tortricodes alternella Eana incanana Cnephasia incertana Cnephasia stephensiana Cnephasia asseclana Spatalistis bifasciana Tortrix viridana Aleimma loeflingiana Acleris forsskaleana Acleris bergmanniana Acleris laterana Acleris abietana Acleris sparsana Acleris rhombana Acleris emargana Acleris schalleriana Acleris cristana Acleris variegana Acleris aspersana Acleris umbrana* Acleris hastiana Acleris hyemana Acleris ferrugana Acleris notana Acleris kochiella Acleris literana Eulia ministrana Pseudargyrotoza conwagana Phtheochroa rugosana

n/a n/a Red-barred Tortrix n/a n/a Large Fruit-tree Tortrix Variegated Golden Tortrix n/a n/a n/a n/a Chequered Fruit-tree Tortrix Barred Fruit-tree Tortrix Dark Fruit-tree Tortrix n/a Timothy Tortrix n/a Light Brown Apple Moth n/a n/a n/a Light Grey Tortrix Grey Tortrix Flax Tortrix n/a Green Oak Tortrix n/a n/a n/a n/a n/a n/a Rhomboid Tortrix n/a n/a n/a Garden Rose Tortrix n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a

L L C C C C C L C C C C C C C C C C C C L C C C L C C C C C Na C C C L C C C Na C L C C L L C C C

Schreckensteiniidae Epermeniidae

Choreutidae

Tortricidae

60

49.096 49.097 49.099 49.109 49.110 49.120 49.121 49.124 49.127 49.128 49.129 49.133 49.134 49.137 49.144 49.149 49.150 49.151 49.154 49.156 49.157 49.158 49.161 49.166 49.180 49.181 49.183 49.184 49.186 49.187 49.188 49.190 49.191 49.193 49.194 49.199 49.203 49.204 49.206 49.207 49.209 49.211 49.214 49.215 49.216 49.223 49.224 49.225 49.230 49.231 49.233 49.234 49.237 49.238 49.240 49.243 49.244 49.245 49.246 49.248 49.249 49.251 49.252 49.253 49.254 49.255 49.257 49.260 49.264 49.265 49.266

Hysterophora maculosana Cochylimorpha straminea Phalonidia gilvicomana Agapeta hamana Agapeta zoegana Aethes smeathmanniana Aethes tesserana Aethes francillana Aethes cnicana Aethes rubigana Cochylidia rupicola Cochylis nana Cochylis roseana Cochylis dubitana Eudemis profundana Apotomis turbidana Apotomis betuletana Apotomis capreana Orthotaenia undulana Hedya nubiferana Hedya pruniana Hedya ochroleucana Celypha striana Celypha lacunana Piniphila bifasciana Lobesia occidentis Lobesia abscisana Lobesia reliquana Endothenia gentianaeana Endothenia oblongana Endothenia marginana Endothenia ustulana Endothenia nigricostana Endothenia quadrimaculana Bactra lancealana Eucosmomorpha albersana Ancylis laetana Ancylis obtusana Ancylis upupana Ancylis geminana Ancylis diminutana Ancylis myrtillana Ancylis badiana Ancylis achatana Ancylis mitterbacheriana Rhopobota naevana Spilonota ocellana Spilonota laricana Epinotia trigonella Epinotia brunnichana Epinotia solandriana Epinotia abbreviana Epinotia signatana Epinotia cruciana Epinotia immundana Epinotia demarniana Epinotia subocellana Epinotia tetraquetrana Epinotia pygmaeana Epinotia tenerana Epinotia ramella Epinotia rubiginosana Epinotia tedella Epinotia fraternana Epinotia bilunana Epinotia nisella Zeiraphera griseana Zeiraphera isertana Eucosma obumbratana Eucosma cana Eucosma hohenwartiana

n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a Marbled Orchard Tortrix Plum Tortrix n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a Holly Tortrix Bud Moth n/a n/a n/a n/a n/a n/a Willow Tortrix n/a n/a n/a n/a n/a Nut Bud Moth n/a n/a n/a n/a n/a n/a Larch Tortrix n/a n/a n/a n/a

61

L C L C C C L L C C L C L C C C C L C C C C C C L pRDB2 C L C L C Nb L L C L L Nb Nb L L L C C C C C L C C C C L L C L C C Nb C C L C Nb C C L C L C C

49.269 49.270 49.276 49.279 49.280 49.281 49.284 49.285 49.286 49.289 49.292 49.293 49.294 49.295 49.296 49.297 49.298 49.306 49.307 49.309 49.311 49.313 49.316 49.318 49.320 49.321 49.322 49.324 49.325 49.332 49.338 49.341 49.342 49.345 49.347 49.354 49.360 49.363 49.365 49.367 49.371 49.376 49.377 49.381 49.382

Eucosma campoliliana Eucosma aemulana Eucosma aspidiscana Gypsonoma dealbana Gypsonoma oppressana Gypsonoma sociana Epiblema sticticana Epiblema scutulana Epiblema cirsiana Epiblema costipunctana Notocelia cynosbatella Notocelia tetragonana Notocelia uddmanniana Notocelia roborana Notocelia incarnatana Notocelia rosaecolana Notocelia trimaculana Rhyacionia pinicolana Rhyacionia pinivorana Dichrorampha plumbana Dichrorampha aeratana Dichrorampha acuminatana Dichrorampha sequana Dichrorampha vancouverana Dichrorampha alpinana Dichrorampha petiverella Dichrorampha plumbagana Cydia nigricana Cydia ulicetana Cydia coniferana Cydia pomonella Cydia splendana Cydia fagiglandana Lathronympha strigana Grapholita compositella Grapholita jungiella Pammene splendidulana Pammene argyrana Pammene albuginana Pammene fasciana Pammene rhediella Pammene aurita Pammene germmana Strophedra weirana Strophedra nitidana

n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a Bramble Shoot Moth n/a n/a n/a n/a n/a Spotted Shoot Moth n/a n/a n/a n/a n/a n/a n/a n/a Pea Moth n/a n/a Codling Moth n/a n/a n/a n/a n/a n/a n/a n/a n/a Fruitlet Mining Tortrix n/a n/a n/a n/a

C pRDB2 Nb C Nb C L C C C C Na C C Nb C C C C L L L L L L C L C C Nb C C C C C C Nb L L C L L L L L

50.002

Zeuzera pyrina

Leopard Moth

C

52.003 52.010 52.012 52.014

Sesia bembeciformis Synanthedon andrenaeformis Synanthedon vespiformis Bembecia ichneumoniformis

Lunar Hornet Moth Orange-tailed Clearwing Yellow-legged Clearwing Six-belted Clearwing

L L L L

54.008 54.009

Zygaena filipendulae Zygaena lonicerae

Six-spot Burnet Narrow-bordered Five-spot Burnet

C C

62.001 62.005 62.006 62.007 62.015 62.024 62.025 62.028 62.029 62.032 62.034 62.035 62.038 62.042 62.048

Aphomia sociella Achroia grisella Galleria mellonella Cryptoblabes bistriga Delplanqueia inscriptella Rhodophaea formosa Dioryctria sylvestrella Dioryctria abietella Phycita roborella Nephopterix angustella Acrobasis repandana Acrobasis advenella Acrobasis consociella Myelois circumvoluta Euzophera pinguis

Bee Moth Lesser Wax Moth Wax Moth n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a Thistle Ermine n/a

C L C L L L (Pempella) Nb C C L C C (Trachycera) L C C

Cossidae Sesiidae

Zygaenidae

Pyralidae

62

62.054 62.058 62.065 62.072 62.074 62.075 62.076 62.077

Homoeosoma sinuella Phycitodes binaevella Ephestia woodiella Pyralis farinalis Aglossa pinguinalis Hypsopygia costalis Hypsopygia glaucinalis Endotricha flammealis

n/a n/a n/a Meal Moth Large Tabby Gold Triangle n/a n/a

C C L C L C C (Orthopygia) C

63.005 63.006 63.007 63.009 63.014 63.017 63.018 63.020 63.022 63.024 63.025 63.028 63.029 63.030 63.031 63.033 63.034 63.037 63.038 63.040 63.052 63.054 63.057 63.060 63.062 63.063 63.064 63.066 63.067 63.069 63.073 63.074 63.075 63.080 63.081 63.086 63.088 63.089 63.090 63.091 63.093 63.095 63.099 63.102 63.114 63.115 63.116

Pyrausta despicata Pyrausta aurata Pyrausta purpuralis Pyrausta nigrata Sitochroa palealis Anania lancealis Anania coronata Anania perlucidalis Anania crocealis Anania funebris Anania hortulata Ostrinia nubilalis Paratalanta pandalis Paratalanta hyalinalis Udea ferrugalis Udea lutealis Udea prunalis Udea olivalis Patania ruralis Mecyna asinalis Nomophila noctuella Cydalima perspectalis* Evergestis forficalis Evergestis pallidata Scoparia subfusca Scoparia basistrigalis Scoparia ambigualis Scoparia pyralella Eudonia lacustrata Eudonia angustea Eudonia truncicolella Eudonia mercurella Eudonia pallida Chrysoteuchia culmella Crambus pascuella Crambus lathoniellus Crambus perlella Agriphila tristella Agriphila inquinatella Agriphila latistria Agriphila straminella Agriphila geniculea Catoptria pinella Catoptria falsella Elophila nymphaeata Acentria ephemerella Cataclysta lemnata

n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a Small Magpie European Corn-borer Bordered Pearl n/a Rusty-dot Pearl n/a n/a n/a Mother of Pearl n/a Rush Veneer Box-tree Moth Garden Pebble n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a Garden Grass-veneer n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a Brown China-mark Water Veneer Small China-mark

C C C L L C (Periephela) C (Phylctaenia) C (Phylctaenia) L (Ebulea) Nb C (Eurrypara) L Na Nb M C C C C (Pleuroptya) Nb M M- A C C C L C C C C C C L C C C C C C L C C C C C C C

65.001 65.002 65.003 65.005 65.006 65.007 65.008 65.009 65.010 65.011 65.012 65.013 65.014 65.015

Falcaria lacertinaria Watsonalla binaria Watsonalla cultraria Drepana falcataria Sabra harpagula Cilix glaucata Thyatira batis Habrosyne pyritoides Tethea ocularis Tethea or Tetheella fluctuosa Ochropacha duplaris Cymatophorina diluta Polyploca ridens

Scalloped Hook-tip Oak Hook-tip Barred Hook-tip Pebble Hook-tip Scarce Hook-tip Chinese Character Peach Blossom Buff Arches Figure of Eighty Poplar Lutestring Satin Lutestring Common Lutestring Oak Lutestring Frosted Green

C C - VU C C Na - NR C C C C L L C L C

Crambidae

Drepanidae

63

65.016

Achlya flavicornis

Yellow Horned

C

66.001 66.002 66.003 66.007 66.008 66.010

Poecilocampa populi Trichiura crataegi Malacosoma neustria Lasiocampa quercus* Macrothylacia rubi Euthrix potatoria

December Moth Pale Eggar Lackey Oak Eggar Fox Moth Drinker

C C - VU C C C C

69.001 69.002 69.003 69.010 69.016

Mimas tiliae Smerinthus ocellata Laothoe populi Macroglossum stellatarum Deilephila elpenor

Lime Hawk-moth Eyed Hawk-moth Poplar Hawk-moth Humming-bird Hawk-moth Elephant Hawk-moth

C C C M C

70.006 70.008 70.009 70.011 70.012 70.013 70.016 70.027 70.029 70.031 70.036 70.037 70.038 70.041 70.043 70.045 70.049 70.051 70.052 70.053 70.054 70.055 70.057 70.059 70.061 70.062 70.064 70.065 70.066 70.067 70.068 70.074 70.075 70.077 70.079 70.081 70.084 70.085 70.086 70.087 70.089 70.091 70.093 70.094 70.095 70.097 70.098 70.099 70.100 70.101 70.103 70.104 70.105 70.106 70.107 70.108

Idaea fuscovenosa Idaea seriata Idaea subsericeata Idaea dimidiata Idaea trigeminata Idaea biselata Idaea aversata Scopula floslactata Timandra comae Cyclophora annularia Cyclophora punctaria Cyclophora linearia Rhodometra sacraria Scotopteryx luridata Scotopteryx bipunctaria Scotopteryx chenopodiata Xanthorhoe fluctuata Xanthorhoe spadicearia Xanthorhoe ferrugata Xanthorhoe designata Xanthorhoe montanata Xanthorhoe quadrifasiata Catarhoe rubidata Camptogramma bilineata Epirrhoe alternata Epirrhoe rivata Euphyia biangulata Euphyia unangulata Earophila badiata Anticlea derivata Mesoleuca albicillata Hydriomena furcata Hydriomena impluviata Pennithera firmata Thera britannica Thera obeliscata Plemyria rubiginata Cidaria fulvata Electrophaes corylata Cosmorhoe ocellata Eulithis prunata Eulithis populata Gandaritis pyraliata Ecliptopera silaceata Chloroclysta siterata Dysstroma truncata Dysstroma citrata Colostygia olivata Colostygia pectinataria Colostygia multistrigaria Lampropteryx suffumata Lampropteryx otregiata Operophtera fagata Operophtera brumata Epirrita dilutata Epirrita christyi

Dwarf Cream Wave Small Dusty Wave Satin Wave Single-dotted Wave Treble Brown Spot Small Fan-footed Wave Riband Wave Cream Wave Blood-Vein Mocha Maiden's Blush Clay Triple-lines Vestal July Belle Chalk Carpet Shaded Broad-bar Garden Carpet Red Twin-spot Carpet Dark-barred Twin-spot Carpet Flame Carpet Silver-ground Carpet Large Twin-spot Carpet Ruddy Carpet Yellow Shell Common Carpet Wood Carpet Cloaked Carpet Sharp-angled Carpet Shoulder Stripe Streamer Beautiful Carpet July Highflyer May Highflyer Pine Carpet Spruce Carpet Grey Pine Carpet Blue-bordered Carpet Barred Yellow Broken-barred Carpet Purple Bar Phoenix Northern Spinach Barred Straw Small Phoenix Red-Green Carpet Common Marbled Carpet Dark Marbled Carpet Beech-green Carpet Green Carpet Mottled Grey Water Carpet Devon Carpet Northern Winter Moth Winter Moth November Moth Pale November Moth

C C C C C C C C C L C C M C L C C C C - NT C C C L C C C L C C C C C C C C C C C C C C C C C C C C L C C C L C C C C

Lasiocampidae

Sphingidae

Geometridae

64

70.110 70.111 70.112 70.113 70.114 70.116 70.117 70.118 70.119 70.121 70.123 70.126 70.127 70.128 70.132 70.133 70.134 70.137 70.138 70.141 70.142 70.144 70.145 70.146 70.147 70.148 70.150 70.151 70.155 70.156 70.157 70.158 70.159 70.160 70.161 70.162 70.163 70.164 70.171 70.173 70.175 70.177 70.179 70.180 70.181 70.182 70.183 70.184 70.187 70.190 70.192 70.193 70.195 70.196 70.198 70.200 70.202 70.203 70.205 70.206 70.207 70.208 70.211 70.214 70.217 70.221 70.222 70.223 70.224 70.226 70.227

Epirrita filigrammaria Asthena albulata Euchoeca nebulata Hydrelia sylvata Hydrelia flammeolaria Venusia blomeri Minoa murinata Philereme vetulata Philereme transversata Rheumaptera undulata Triphosa dubitata Horisme vitalbata Horisme tersata Melanthia procellata Perizoma affinitata Perizoma alchemillata Perizoma bifaciata Perizoma albulata Perizoma flavofasciata Gymnoscelis rufifasciata Chloroclystis v-ata Pasiphila rectangulata Pasiphila debiliata Eupithecia haworthiata Eupithecia tenuiata Eupithecia inturbata Eupithecia linariata Eupithecia pulchellata Eupithecia venosata Eupithecia abbreviata Eupithecia dodoneata Eupithecia pusillata Eupithecia phoeniceata Eupithecia tripunctaria Eupithecia virgaureata Eupithecia tantillaria Eupithecia lariciata Eupithecia egenaria Eupithecia indigata Eupithecia centaureata Eupithecia trisignaria Eupithecia satyrata Eupithecia absinthiata Eupithecia expallidata Eupithecia valerianata Eupithecia assimilata Eupithecia vulgata Eupithecia exiguata Eupithecia icterata Eupithecia subfuscata Aplocera plagiata Aplocera efformata Chesias legatella Chesias rufata Lobophora halterata Acasis viretata Trichopteryx carpinata Archiearis parthenias Abraxas grossulariata Abraxas sylvata Lomaspilis marginata Ligdia adustata Macaria notata Macaria liturata Macaria brunneata Cepphis advenaria Petrophora chlorosata Plagodis pulveraria Plagodis dolabraria Opisthograptis luteolata Epione repandaria

Small Autumnal Moth Small White Wave Dingy Shell Waved Carpet Small Yellow Wave Blomer's Rivulet Drab Looper Brown Scallop Dark Umber Scallop Shell Tissue Small Waved Umber Fern Pretty Chalk Carpet Rivulet Small Rivulet Barred Rivulet Grass Rivulet Sandy Carpet Double-striped Pug V-Pug Green Pug Bilberry Pug Haworth's Pug Slender Pug Maple Pug Toadflax Pug Foxglove Pug Netted Pug Brindled Pug Oak-tree Pug Juniper Pug Cypress Pug White-spotted Pug Golden-rod Pug Dwarf Pug Larch Pug Fletcher's Pug or Pauper Pug Ochreous Pug Lime-speck Pug Triple-spotted Pug Satyr Pug Wormwood Pug Bleached Pug Valerian Pug Currant Pug Common Pug Mottled Pug Tawny Speckled Pug Grey Pug Treble-bar Lesser Treble-bar Streak Broom-tip Seraphim Yellow-barred Brindle Early Tooth-striped Orange Underwing Magpie Moth Clouded Magpie Clouded Border Scorched Carpet Peacock Moth Tawny-barred Angle Rannoch Looper Little Thorn Brown Silver-line Barred Umber Scorched Wing Brimstone Moth Bordered Beauty

65

L C C L C L Nb - NS C C C L C C C C C C C C C C C Nb - NS C C C - EN C - NT C L C C C L C C C C - NT Nb - NS C C C C C L L C C C C C C C C L - NT C C C C C C C C C C Nb - NS Nb - NS C C C C C

70.229 70.231 70.233 70.234 70.235 70.236 70.237 70.238 70.239 70.240 70.241 70.243 70.244 70.245 70.246 70.247 70.248 70.251 70.252 70.253 70.254 70.255 70.256 70.257 70.258 70.264 70.265 70.266 70.267 70.270 70.272 70.273 70.274 70.276 70.277 70.278 70.279 70.280 70.282 70.283 70.284 70.287 70.288 70.295 70.297 70.299 70.300 70.302 70.303 70.305

Pseudopanthera macularia Apeira syringaria Ennomos quercinaria Ennomos alniaria Ennomos fuscantaria Ennomos erosaria Selenia dentaria Selenia lunularia Selenia tetralunaria Odontopera bidentata Crocallis elinguaria Ourapteryx sambucaria Colotois pennaria Alsophila aescularia Apocheima hispidaria Phigalia pilosaria Lycia hirtaria Biston strataria Biston betularia Agriopis leucophaearia Agriopis aurantiaria Agriopis marginaria Erannis defoliaria Menophra abruptaria Peribatodes rhomboidaria Deileptenia ribeata Alcis repandata Alcis jubata Hypomecis roboraria Ectropis crepuscularia Paradarisa consonaria Parectropis similaria Aethalura punctulata Bupalus piniaria Cabera pusaria Cabera exanthemata Lomographa bimaculata Lomographa temerata Theria primaria Campaea margaritaria Hylaea fasciaria Charissa obscurata Cleorodes lichenaria Perconia strigillaria Pseudoterpna pruinata Geometra papilionaria Comibaena bajularia Hemistola chrysoprasaria Jodis lactearia Hemithea aestivaria

Speckled Yellow Lilac Beauty August Thorn Canary-shouldered Thorn Dusky Thorn September Thorn Early Thorn Lunar Thorn Purple Thorn Scalloped Hazel Scalloped Oak Swallow-tailed Moth Feathered Thorn March Moth Small Brindled Beauty Pale Brindled Beauty Brindled Beauty Oak Beauty Peppered Moth Spring Usher Scarce Umber Dotted Border Mottled Umber Waved Umber Willow Beauty Satin Beauty Mottled Beauty Dotted Carpet Great Oak Beauty Engrailed Square Spot Brindled White-spot Grey Birch Bordered White Common White Wave Common Wave White-pinion Spotted Clouded Silver Early Moth Light Emerald Barred Red Annulet Brussels Lace Grass Wave Grass Emerald Large Emerald Blotched Emerald Small Emerald Little Emerald Common Emerald

C C C C C C C C C C C C C C C C - VU C C C C C C C - VU C C C C C L C C C C C C C C C C C C L C L C C C C C C

71.003 71.005 71.006 71.007 71.009 71.010 71.011 71.012 71.013 71.016 71.017 71.018 71.020 71.021 71.025 71.027

Cerura vinula Furcula furcula Furcula bicuspis Furcula bifida Stauropus fagi Drymonia dodonaea Drymonia ruficornis Notodonta dromedarius Notodonta ziczac Peridea anceps Pheosia tremula Pheosia gnoma Pterostoma palpina Ptilodon capucina Phalera bucephala Clostera curtula

Puss Moth Sallow Kitten Alder Kitten Poplar Kitten Lobster Moth Marbled Brown Lunar Marbled Brown Iron Prominent Pebble Prominent Great Prominent Swallow Prominent Lesser Swallow Prominent Pale Prominent Coxcomb Prominent Buff-tip Chocolate-tip

C C L C C C C C C C C C C C C C

72.001 72.002 72.003

Scoliopteryx libatrix Rivula sericealis Hypena proboscidalis

Herald Straw Dot Snout

C C C

Notodontidae

Erebidae

66

72.004 72.007 72.010 72.013 72.015 72.017 72.019 72.020 72.022 72.024 72.029 72.031 72.034 72.035 72.036 72.037 72.038 72.041 72.042 72.043 72.044 72.045 72.046 72.049 72.053 72.055 72.061 72.063 72.067 72.069 72.076 72.078 72.083 72.084

Hypena rostralis Hypena crassalis Lymantria monacha Euproctis similis Calliteara pudibunda Orgyia antiqua Spilosoma lutea Spilosoma lubricipeda Diaphora mendica Phragmatobia fuliginosa Callimorpha dominula Tyria jacobaeae Utetheisa pulchella Miltochrista miniata Nudaria mundana Thumatha senex Cybosia mesomella Lithosia quadra* Atolmis rubricollis Eilema depressa Eilema griseola Eilema lurideola Eilema complana Eilema sororcula Herminia tarsipennalis Herminia grisealis Schrankia costaestrigalis Lygephila pastinum Phytometra viridaria Laspeyria flexula Catocala fraxini Catocala nupta Euclidia glyphica Euclidia mi

Buttoned Snout Beautiful Snout Black Arches Yellow-tail Pale Tussock Vapourer Buff Ermine White Ermine Muslin Moth Ruby Tiger Scarlet Tiger Cinnabar Crimson Speckled Rosy Footman Muslin Footman Round-winged Muslin Four-dotted Footman Four-spotted Footman Red-necked Footman Buff Footman Dingy Footman Common Footman Scarce Footman Orange Footman Fan-foot Small Fan-foot Pinion-streaked Snout Blackneck Small Purple-barred Beautiful Hook-tip Clifden Nonpareil Red Underwing Burnet Companion Mother Shipton

L C C C C C C C C C C C M C C C C L C C C C C C C C C C - NT L C M - NS C C C

73.001 73.002 73.012 73.015 73.016 73.017 73.022 73.024 73.032 73.033 73.036 73.037 73.038 73.039 73.040 73.045 73.046 73.047 73.048 73.052 73.053 73.058 73.062 73.063 73.064 73.068 73.069 73.084 73.092 73.095 73.096 73.097 73.101 73.102 73.105 73.107

Abrostola tripartita Abrostola triplasia Diachrysia chrysitis Autographa gamma Autographa pulchrina Autographa jota Plusia festucae Deltote pygarga Colocasia coryli Diloba caeruleocephala Acronicta alni Acronicta tridens Acronicta psi Acronicta aceris Acronicta leporina Acronicta rumicis Subacronicta megacephala Craniophora ligustri Panemeria tenebrata Cucullia umbratica Cucullia chamomillae Cucullia verbasci Amphipyra pyramidea Amphipyra berbera Amphipyra tragopoginis Allophyes oxyacanthae Xylocampa areola Bryophila domestica Caradrina morpheus Caradrina clavipalpis Hoplodrina octogenaria Hoplodrina blanda Charanyca trigrammica Rusina ferruginea Dypterygia scabriuscula Mormo maura

Spectacle Dark Spectacle Burnished Brass Silver Y Beautiful Golden Y Plain Golden Y Gold Spot Marbled White Spot Nut-tree Tussock Figure of Eight Alder Moth Dark Dagger Grey Dagger Sycamore Miller Knot Grass Poplar Grey Coronet Small Yellow Underwing Shark Chamomile Shark Mullein Copper Underwing Svensson's Copper Underwing Mouse Moth Green-brindled Crescent Early Grey Marbled Beauty Mottled Rustic Pale Mottled Willow Uncertain Rustic Treble Lines Brown Rustic Bird's Wing Old Lady

C C C M C C C C C L - EN C C C C C C C C C C C C C C C - VU C C C C C C C C C C C

Noctuidae

67

73.109 73.113 73.114 73.120 73.121 73.123 73.128 73.131 73.134 73.144 73.147 73.154 73.155 73.156 73.158 73.16 73.162 73.163 73.168 73.169 73.17 73.171 73.172 73.173 73.174 73.175 73.176 73.179 73.18 73.181 73.182 73.186 73.187 73.188 73.189 73.19 73.192 73.193 73.194 73.195 73.197 73.200 73.201 73.202 73.206 73.21 73.212 73.215 73.216 73.219 73.22 73.222 73.224 73.225 73.228 73.233 73.238 73.241 73.242 73.243 73.244 73.245 73.246 73.247 73.249 73.250 73.251 73.253 73.254 73.261 73.264

Thalpophila matura Phlogophora meticulosa Euplexia lucipara Eremobia ochroleuca Gortyna flavago Hydraecia micacea Amphipoea oculea Luperina testacea Rhizedra lutosa Denticucullus pygmina Photedes minima Apamea remissa Apamea epomidion Apamea crenata Apamea sordens Apamea scolopacina Apamea monoglypha Apamea lithoxylaea Lateroligia ophiogramma Mesapamea secalis Mesapamea didyma Litoligia literosa Mesoligia furuncula Oligia strigilis Oligia latruncula Oligia versicolor Oligia fasciuncula Tiliacea citrago Tiliacea aurago Xanthia togata Cirrhia icteritia Agrochola lychnidis Agrochola litura Agrochola helvola Agrochola lota Agrochola macilenta Agrochola circellaris Omphaloscelis lunosa Conistra vaccinii Conistra ligula Conistra rubiginea Lithophane semibrunnea Lithophane socia Lithophane ornitopus Lithophane leautieri Eupsilia transversa Ipimorpha retusa Cosmia affinis Cosmia trapezina Atethmia centrago Brachylomia viminalis Apterogenum ypsillon Griposia aprilina Dryobotodes eremita Antitype chi Aporophyla nigra Mniotype adusta Panolis flammea Orthosia incerta Orthosia miniosa Orthosia cerasi Orthosia cruda Orthosia populeti Orthosia gracilis Orthosia gothica Anorthoa munda Egira conspicillaris Tholera decimalis Cerapteryx graminis Polia nebulosa Lacanobia thalassina

Straw Underwing Angle Shades Small Angle Shades Dusky Sallow Frosted Orange Rosy Rustic Ear Moth Flounced Rustic Large Wainscot Small Wainscot Small Dotted Buff Dusky Brocade Clouded Brindle Clouded-bordered Brindle Rustic Shoulder-knot Slender Brindle Dark Arches Light Arches Double Lobed Common Rustic Lesser Common Rustic Rosy Minor Cloaked Minor Marbled Minor Tawny Marbled Minor Rufous Minor Middle-barred Minor Orange Sallow Barred Sallow Pink-barred Sallow Sallow Beaded Chestnut Brown-spot Pinion Flounced Chestnut Red-line Quaker Yellow-line Quaker Brick Lunar Underwing Chestnut Dark Chestnut Dotted Chestnut Tawny Pinion Pale Pinion Grey Shoulder-knot Blair's Shoulder-knot Satellite Double Kidney Lesser-spotted Pinion Dun-bar Centre-barred Sallow Minor Shoulder-knot Dingy Shears Merveille du Jour Brindled Green Grey Chi Black Rustic Dark Brocade Pine Beauty Clouded Drab Blossom Underwing Common Quaker Small Quaker Lead-coloured Drab Powdered Quaker Hebrew Character Twin-spotted Quaker Silver Cloud Feathered Gothic Antler Moth Grey Arches Pale-shouldered Brocade

68

C C C C C C C C C C C C C C C C C C C C C C - NT C C C C C C C C C C - NT C C - NT C C C C C C C C C C C C L L C C C C C C C C C C C L C C C C C C Nb - NS C C C C

73.265 73.267 73.270 73.273 73.274 73.279 73.281 73.290 73.291 73.293 73.298 73.300 73.301 73.317 73.319 73.320 73.325 73.327 73.328 73.329 73.332 73.333 73.334 73.336 73.337 73.338 73.342 73.343 73.345 73.346 73.348 73.351 73.352 73.353 73.357 73.358 73.359 73.36 73.361 73.365

Lacanobia contigua Lacanobia oleracea Melanchra persicariae Hada plebeja Mamestra brassicae Hecatera bicolorata Hadena bicruris Mythimna conigera Mythimna pallens Mythimna impura Mythimna ferrago Mythimna l-album* Leucania comma Agrotis exclamationis Agrotis segetum Agrotis clavis Agrotis puta Agrotis ipsilon Axylia putris Ochropleura plecta Diarsia brunnea Diarsia mendica Diarsia rubi Cerastis rubricosa Cerastis leucographa Lycophotia porphyrea Noctua pronuba Noctua fimbriata Noctua comes Noctua interjecta Noctua janthe Graphiphora augur Anaplectoides prasina Xestia baja Xestia xanthographa Xestia sexstrigata Xestia c-nigrum Xestia ditrapezium Xestia triangulum Eugnorisma glareosa

Beautiful Brocade Bright-line Brown-eye Dot Moth Shears Cabbage Moth Broad-barred White Lychnis Brown-line Bright-eye Common Wainscot Smoky Wainscot Clay L-album Wainscot Shoulder-striped Wainscot Heart and Dart Turnip Moth Heart and Club Shuttle-shaped Dart Dark Sword-grass Flame Flame Shoulder Purple Clay Ingrailed Clay Small Square-spot Red Chestnut White-marked True Lover's Knot Large Yellow Underwing Broad-bordered Yellow Underwing Lesser Yellow Underwing Least Yellow Underwing Lesser Broad-bordered Yellow Underwing Double Dart Green Arches Dotted Clay Square-spot Rustic Six-striped Rustic Setaceous Hebrew Character Triple-spotted Clay Double Square-spot Autumnal Rustic

L C C C C C - NT C C C C C

74.003 74.004 74.007 74.008 74.009

Nola cucullatella Nola confusalis Bena bicolorana Pseudoips prasinana Nycteola revayana

Short-cloaked Moth Least Black Arches Scarce Silver-lines Green Silver-lines Oak Nycteoline

C C C C C

L C C C C C M C C C C C C L C - VU C C C C C C C C C C C C C C

Nolidae

Undetermined to species: Oegoconia sp.

An Oegoconia species Parornix sp.

Parornix species agg.

Yponomeuta padella/malinellus/cagnagella sp.

Grand Total 972 without agg The following are not yet accepted or awaiting determination: Stigmella floslactella Stigmella hemargyrella 15.086 Phyllonorycter geniculella 15.083 Phyllonorcter trifasciella Argyrestia curvella Cauchas rufimitrella Agonopterix scopariella Syncopacma larseniella ag. Monochroa tenebrella chk cystella Cnephasia communana ’22 with PH Clepsis spectrana Acleris comariana Acleris umbrana Eucosma conterminana Jul’17, Aug’15

69

3.7 Checklist of order Hymenoptera – Bees, Wasps & Ants Family

Species

Status

Source

Suborder APOCRITA - ACULEATA. Ants, Bees and Wasps Apoidea

Apidae - Honey Bee, Bumblebees etc Apinae Aphis mellifera Western Honey Bee Bombus terrestris Buff-tailed Bumblebee Bombus hortorum Garden Bumblebee Bombus pratorum Early-nesting Bumblebee Bombus leucorum White-tailed Bumblebee Bombus lapidarius Red-tailed Bumblebee Bombus pascuorum Common Carder Bumblebee Bombus vestalis Vestal Cuckoo-Bee Bombus hypnorum Tree Bumblebee Bombus rupestris Red-tailed Cuckoo Bumblebee Bombus ruderatus Large Garden Bumblebee Bombus sylvestris Forest Bumblebee Anthophorinae Nomada goodeniana Gooden's Nomad Bee Nomada flava sl.

Flavous Nomad Bee Nomada fabriciana Fabricius Nomad Bee Nomada marshamella Marsham's Nomad Bee Nomada lathburiana Lathbury's Nomad Bee Anthophora plumipes Hairy-footed Flower Bee Anthophora furcata Fork-tailed Flower Bee Eucera longicornis Long-horned Bee Andreninae Andrena bicolor Gwynne's Mining Bee Andrena bucephala Big-headed Mining Bee Andrena cineraria Ashy (Grey-haired) Mining Bee Andrena chrysosceles Hawthorn Mining Bee Andrena dorsata Short-fringed Mining Bee Andrena fucata Painted Mining Bee Andrena fulva Tawny Mining Bee Andrena haemorrhoa Orange-tailed Mining Bee Andrena helvola Coppice Mining Bee Andrena minutula Common Mini-miner Andrena nitida Grey-patched Mining Bee Andrena nigroaenea Buffish Mining Bee Andrena praecox Small Sallow Mining Bee Andrena semilaevis Shiny-margined Mini-miner Andrena scotica Chocolate Mining Bee Andrena wilkella Wilke's Mining Bee Melitta leporina Clover Melitta Halictinae Halictus rubicundus Orange-legged Furrow-bee Halictis tumulorum Bronze Mining Bee Sphecodes monilicornis Box-headed Blood-bee Sphecodes ephippius Bare-saddled Blood-bee Lasioglossum laevigatum Red-backed Furrow-bee Lasioglossum calceatum Common Furrow Bee Lasioglossum morio Green Furrow Bee Lasioglossum smeathmanellum Smeathman's Furrow Bee Lasioglossum leucopus White-footed Furrow Bee Lasioglossum fulvicorne Chalk Furrow Bee Megachilinae Stelis phaeoptera

Plain Dark Bee

70

Nb

m m ID m m m m m m WT m ID ID ID ID ID ID ID ID ID ID/LO ID ID/NBN ID ID/LO ID ID ID ID ID ID NBN* NBN* ID NBN* ID ID ID ID ID ID ID NBN NBN ID ID

ID

Osmia bicornis Osmia leaiana Osmia caerulescens Megachile willughbiella Megachile centuncularis Chelostoma florisomne Anthidium maniculatum Melecta albifrons Coelioxys elongate sl Colletinae Hylaeus communis Hylaeus confusus Colletes daviesanus Colletes hederae Vespoidea Vespidae – Social Wasps Vespula vulgaris Vespula germanica Vespa crabro Dolichovespula sylvestris Dolichovespula norwegica Eumeninae Ancistrocerus nigricornis Crabronidae - Digger Wasps Mellinus arvensis Ectemnius continuus Ectemnius cavifrons Ectemnius cephalotes Pemphredon lugubris Philanthus triangulum Lindenius albilabris Trypoxylon attenuatum sl Chrysidoidea Chrysididae - Jewel Wasps Chrysis ignita ag Pompiloidea Pompilidae - Spider-hunting Wasps Dipogon varigatus Anoplius nigerrimus Tiphiodea Tiphiidae - Tiphiid Wasps Methocha articulata

Red Mining Bee Orange-vented Mason Bee Blue Mason Bee Willughby’s Leafcutter Bee Patchwork Leafcutter Sleepy Carpenter Bee Wool Carder Bee Common Mourning Bee Sharp-tailed Bee

ID m ID m ID NBN ID ID ID

Common Yellow-faced Bee White-jawed Yellow-faced Bee Davies’ Colletes Ivy Bee

m ID ID ID

Common Wasp German Wasp Hornet Tree Wasp Norwegian Wasp

M ID M ID ID

Early Mason Wasp

ID

Field Digger Wasp Common Ectemnius Garden Ectemnius Large Ectemnius Mournful Wasp Beewolf White-lipped Digger Wasp Slender Wood-borer Wsp

ID ID ID ID ID ID ID ID

Ruby-tailed Wasp

I

n/a Black Spider Wasp

ID ID

Tiger Beetle-hunting Wasp

NBN*

Red Ant n/a n/a n/a Black Garden Ant Yellow Meadow Ant n/a Southern Wood Ant Negro Ant Shining Guest Ant n/a n/a n/a

ID NBN* WT WT m m ID m m NBN* WT WT WT

Formicoidea Formicidae - Ants Mymica ruginodis Myrmica rubra Myrmica lobicornis Myrmica scabrinodis Lasius niger Lasius flavus Lasius alienus Formica rufa Formica fusca Formicoxenus nitidulus Leptothorax acervorum Leptothorax nylanderi Stenamma westwoodi

71

l c

1 c l l

Suborder APOCRITA PARASITICA Parasitic Wasps Ichneumonidae- Ichneumon or Darwin Wasps Ophion luteus ag. n/a Ophion scutellaris sl n/a Ichneumon suspiciosus ag. n/a Ichneumon stramentarius/stramentor n/a Ichneumon deliratorius Black Ichneumon Stenichneumon culpator n/a Ambylteles armatorius Toothed Tiger Ichneumon Heteropelma amictum ag.

Black and red.

Ephialtes manifestator n/a Rhyssa persuasoria Sabre Wasp Exetastes adpressorius a white-footed ichneumon

ID ID ID ID ID ID ID ID ID ID ID

Gasteruptiidae Gasteruption assectator Gasteruption jaculator

n/a n/a

ID ID

Braconidae unidentified to species.

ID

Cynipidae - Gall Wasps Diplolepis rosea Robin's Pin Cushion Andricus kollari Oak Apple or Marble Gall Neuroterus quercusbaccarum Common Spangle Gall Wasp Andricus quercuscalicis Acorn Knopper Gall

m m ID ID

Chalcididae - Chalcicid Wasps Brachymeria tibialis

Black and Yellow chalcid

I

Cimbicidae - Club-anntennaed Sawflies Cimbex luteus

n/a

NBN*

Tenthredinidae - Common Sawflies Athalia rosae Pachyprotasis rapae Tenthredo zona ag.

Tenthredo atra ag.

Tenthredo mesomela Tenthredo temula Tenthredopsis litterata ag.

Tenthredopsis nassata Macrophylla annulata Eutomostethus ephippium Rhogogaster viridis Nematus septentrionalis Neurotoma saltuum Allantus calceatus

Turnip Sawfly n/a n/a n/a n/a n/a n/a n/a n/a Lawn Sawfly n/a Birch or Flat-legged Sawfly Social Pear Sawfly n/a

ID ID ID ID NBN ID ID NBN ID ID ID ID ID ID

Cephidae - Stem-boring Sawflies Calameuta filiformis Cephus spinipes

Reed Stem Borer n/a

ID ID

Large Rose Sawfly Rose Sawfly

ID ID

Giant Woodwasp

ID

Suborder SYMPHYTA. Sawflies and Woodwasps

Argidae - Argid Sawflies Arge pagana Arge ochropus Siricidae - WoodWasps Urocerus gigas

72

3.8 Checklist of Orders Ephemeroptera (Mayflies), Plecoptera (Stoneflies), and Tricoptera (Caddisflies) Source

Status

Ephemeroptera - Mayflies Baetidae Pale Watery Southern Iron Blue Pond Olive Pale Evening Dun Ephemeridae Green Drake Drake Mackerel Striped Mayfly Caenidae Large Broadwings Potamanthidae Yellow Mayfly Emphemerellidae Blue-winged Olive

Baetis fuscatus Baetis niger/digitatus Cloeon dipterum ag Procloeon bifidum

ID ID ID ID

Ephemera danica Ephemera vulgata Ephemera lineata

ID ID NBN

NS-VU

Brachycercus harrisella

NBN

NS -VU

Potamanthus luteus

M

NS - EN

Serratella ignita

M

Amphinemura standfussi

NBN

Brachyptera putata

NBN

Perlodes microcephalus Isoperla grammatica

NBN NBN

Leuctra fusca Leuctra geniculate

NBN NBN

Rhabdiopteryx acuminata Taeniopteryx nebulosa

NBN NBN

Plecoptera – Stoneflies Nemouridae Taeniopterygidae Perlodidae

Leuctridae

Taeniopterygidae

73

Order Tricoptera – Caddisflies Beraeidae

Beraeodes minutus

NBN

Brachycentridae

Brachycentrus subnubilus

NBN

Halesus radiatus ag Limnephilus rhombicus Limnephilus hirsutus Limnephilus lunatus Limnephilus marmoratus Limnephilus sparsus Potamophylax latipennis Stenophylax permistus

ID ID NBN NBN NBN NBN NBN NBN

Lepidostoma hirtum

NBN

Athripsodes albifrons Athripsodes aterrimus Athripsodes bilineatus Athripsodes cinereus Adicella reducta Ceraclea albimacula Ceraclea annulicornis Ceraclea dissimilis Ceraclea nigronervosa Leptocerus interruptus Mystacides azurea Mystacides nigra Oecetis notata Oecetis Testacea Setodes punctatus Ylodes conspersus

M NBN NBN NBN NBN NBN NBN NBN NBN NBN M NBN NBN NBN NBN NBN

Hydropsyche siltalai Hydropsyche contubernalis Hydropsyche pellucidula Diplectrona felix Cheumatopsyche lepida

M NBN NBN NBN NBN

Allotrichia pallicornis Hydroptila lotensis Hydroptila sparsa Ithytrichia lamellaris

NBN NBN NBN NBN

Agapetus fuscipes ag Glossosoma boltoni

NBN NBN

Goeridae

Goera pilosa Silo pallipes

M NBN

Molannidae

Molanna angustata

NBN

Philopotamidae

Chimarra marginata

NBN

Limnephidae Caperer Cinnamon Sedge

Lepidostomatidae

Leptoceridae Brown Silverhorn

Black Silverhorn

Hydropsychidae Grey Flag etc

Hydroptilidae

Gossosomatidae n/a

74

Philopotamus montanus

NBN

Polycentropodidae

Polycentropus flavomaculatus

NBN

Psychomyiidae

Psychomyia pusilla Tinodes waeneri

NBN NBN

Rhyacophilidae

Rhyacophila dorsalis Rhyacophila munda

NBN NBN

Sericostoma personatum

M

Sericostomatidae Welshman's Button

75

3.9 Checklist of other insect orders Source

Status

Order Dermaptera - Earwigs Common Earwig Lesser Earwing Lesne’s Earwig

Forficula auricularia Labia minor Forficula lesnei

Ectobius pallidus

SO51 only

ID ID NBN

NS

SO51 only

NBN

NS

Order Dictyoptera – Cockroaches Tawny Cockroach

Order Mecoptera - Scorpionflies Panorpidae Scorpionfly

Panorpa communis

M

Atlantorhaphidia maculicollis Phaeostigma notata

M M

Chrysoperla carnea

ID

Order Raphidioptera - Snakeflies Raphididae Pine Snakefly Oak Snakefly

Order Neuroptera - Lacewings Chrysopidae Green lacewing Hemerobiidae Brown Lacewing

Osmylidae Sisyridae

Wesmaelius subnebulosus Sympherobius elegans Hemerobius stigma ag.

Hemerobius humulinus ag.

Micromus variegatus ag.

Osmylus fulvicephalus Sisyra terminalis

SO51

SO51

ID NBN ID ID ID NBN NBN

Thysanoptera - Thrips None reported.

Psocoptera - Barkflies Ectopsocidae n/a

Ectopsocus petersi ag

ID

Dilta hibernica

ID

Lepisma saccharina

ID

Aperygota – Wingless Insects Archaeognatha (formerly Thysanura) Machilidae Southern Bristletail Zygentoma (formerly in Thysanura) Lepismatidae Silverfish

76

Checklists of Class Arachnida.

3.10. Order Aranea - Spiders

WT

Agelenidae Funnelweb Spiders Agelena labyrithica Labyrinth Spider Tegenaria gigantea ag House spider Tegenaria silvestris n/a Amaurobidae Coelotes atropos Amaurobius similis Amaurabius ferox

Laceweb Spiders n/a Lace-web spider n/a

Anyphaenidae Buzzing Spiders Anyphaea accentuata Buzzing Spider Araneidae

Orbweb Spiders Araneus diadematus Garden Spider Araneus quadratus Four-spot Orbweb Spider Araneus marmoreus var pyramidus n/a Agalenatea redii n/a Araniella cucurbitina Cucumber Spider Cyclosa conica n/a Larinioides cornutus n/a Mangora acalypha Cricket-bat Spider Nuctenea umbratica Walnut Orbweb Spider Zygiella x-notata n/a Zilla diodia n/a

1

Lathys humilis

1 1

1

1 1

1 1 1

1

1 1 1 1 1 1

1

1 1 1

Meshweb Spiders n/a

NR

1

1

Gnaphosidae Ground Spiders Drassodes cupreus n/a Drassyllus pusilla ag. n/a Zelotes latreillei ag. n/a

1 1 ?

Linyphiidae Money Spiders Ceratinella brevis Drapetisca socialis n/a Dismodicus bifrons Hypomma cornutum Lepthyphantes tenuis Linyphia triangularis Sheet-web Spider Linyphia hortensis Meioneta saxatilis Microneta viaria Neriene clathrata Neriene montana Pelecopsis radicicola Porrhomma egeria Tenuiphantes flavipes Trematocephalus cristatus Walckenaeria nodosa

Agroeca proxima

1

1 1 1

Dysderidae Woodlouse Spiders Harpactea hombergi n/a

Liocranidae

1 1 1

1

Clubionidae Sac Spiders Clubiona corticalis Silk cell (Sac) Spider Clubiona terrestris (or lutescens) ag. n/a Clubiona caerulescens n/a Dictynidae

Record source ID NBN Status

1 1 1 1 1 1 1 1

1 1

NR NS NS NS

1

77

1 1

1 1 1

Running Foliage Spiders n/a

1

Lycosidae

Wolf Spiders Pardosa amentata Pardosa pullata Trochosa terricola ag Alopecosa pulverulenta

Wolf spider

1 1 1 1

n/a n/a

Philodromidae Running Crab Spiders Philodromus dispar n/a Philodromus aureolus Pholcidae

1 1

1

Pholcus phalangiodes

Cellar Spiders Daddylonglegs spider

1

1

Pisaura miribalis

Nursury-web Spiders Nursery-web spider

1

1

1 1 1 1 1 1

1 1

Pisauridae

Salticidae Salticus scenicus Heliophanus cupreus Ballus chalybeius Euophrys fronalis Evarcha falcata Neon reticulatus

Jumping Spiders Zebra Spider n/a

Sparassidae Huntsman Spiders Micrommata viresens Green Huntsman Tetragnathidae

NS 1

1

NS

Long-jawed Orbweb Spiders

Meta (Metellina) segmentata n/a Metellina mengei Meta menardi Cave spider Tetragnatha extensa/montana ag n/a Pachygnatha degeeri n/a

1 1 1 1

Theridiidae Comb-footed Spiders Achearanea lunata Parasteatoda lunata ?

Anelosimus vittatus Enoplognatha ovata Episinus angulatus Neottiura bimaculata Paidiscura pallens Robertus lividus Robertus neglectus Steatoda bipunctata Comb-footed Spider Theridiom mystaceum Theridion sisyphium

Thomasidae Xysticus cristatus Xysticus kochi Diaea dorsata Misumena vatia

Crab Spiders Crab spider

Uloboridae

Cribellate Orbweb Spiders

Pale crab spider

Hyptiotes paradoxus Zoridae

1

1 1 1

1 1 1 1 1 1 1 1 1

Status NR - Nationally Rare – Vulnerable (=RDB2) - 2 NS - Nationally Scarce -7

78

NS

1 1

1 1 1 1

1 1 1

1

1 81 species

<5%

1

Triangle Spider

Ghost Spiders Zora spinimana

1

NS

3.11. Checklist of Other Arachnida.

Record source Order Opiliones - Harvestmen Phalangiidae Dicranopalpus ramosus Leibunum rotundatum Leiobunum blackwalli Opilio saxatilis Oligolophus tridens Lacinius ephippiatus Lophopilio palpinalis Megabunus diadema Mitopus morio Paroligolophus agrestris Phalangium opilio Platybunus triangularis Rilaena triangularis Homalenotus quadridentatus

WT

NBN

n/a Round Harvestman n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a n/a

1 1

1 1 1 1 1

n/a

1

1

n/a

1

1

1 1

1

1 ?

1 1

1 1 1 1

1 1

Trogulidae Anelasmocephalus cambridgei Nemastomatidae Nemastoma bimaculatum

Order Pseudoscorpionida Pselaphochernes scorpiodes Chthonius ischnoncheles ag.

Compost Chernes Common Chthonid

Order Acariformes – Mites and Ticks Penthaleidae

Penthaleus major

Red-legged Earth-mite

Gall mites

Aceria myriadeum

on Field Maple

Aceria macrochela

on Field Maple

Deer Tick

Ixodes ricinus

(from dog)

79

Status

3.12 Checklist of phylum Mollusca, classes Gastropoda and Bivalva – Slugs, Snails and Mussels Class Gastropoda SO51 only

Source Last record Status

Terrestrial Snails Aciculidae Acicula fusca

1

NBN

Azeca goodalli

1

NBN

Azecidae Carychiidae Carychium tridentatum

NBN

Abida secale

WT/NBN

1968

Chondrinidae Local

Clausiliidae Clausilia bidentata Cochlodina laminata

1

WT/NBN NBN

1974

Cochlicopa lubricella Cochlicopa lubrica

1 1

WT/NBN NBN

2015

Cochlicopidae

Discidae Discus rotundatus

NBN

Merdigera obscura Ena obscura

NBN WT

Carychium tridentatum

WT

cf. Patulidae

Enidae

Ellobiidae Euconulidae Euconulus fulvus

1

NBN

1965

Cecilioides acicula

1

NBN

1885

Zonitoides excavatus

1

NBN

1965

Ferussaciidae Gastrodontidae

Helicoidea Cornu (Helix) aspersum

Common Garden Snail

NBN

Cepaea hortensis Cepaea nemoralis Arianta arbustorum Cochlodina laminata Helicigona lapicida Candidula gigaxii Candidula intersecta Monacha cantiana Trochulus striolatus Trochulus hispidus Zenobiella subrufescens

White-lipped Snail Black-lipped snail Copse Snail Plaited Door Snail Lapidary Snail Eccentric Snail Wrinked Snail

NBN WT/NBN NBN ID WT/NBN ID WT/NBN NBN NBN NBN NBN

Hygromiidae

1 1 1

Local

2015

Lauriidae Lauria cylindracea

WT/NBN

Aegopinella pura Aegopinella nitidula Nesovitrea hammonis Oxychilus alliarius Oxychilus navarricus Oxychilus cellarius

NBN NBN NBN NBN NBN NBN

Oxychilidae

1

1968 2015

2015 2015

Pomatiasidae Pomatias elegans

WT/NBN

Pristilomatidae

80

Local

Vitrea contracta Vitrea crystallina

NBN NBN

1968 1968

Punctum pygmaeum

NBN

1968

Pupilla muscorum

NBN

1895

Pyramidula rupestris Pyramidula pusilla

WT NBN

1899

Punctidae Pupillidae Pyramidulidae Local

Succineidae Oxyloma elegans Succinea putris

1

NBN NBN

1965 1895

Anodonta anatina Pseudanodonta complanata

1

NBN NBN

1909 2005

NBN WT NBN

1968

Unionidae

Valloniidae Acanthinula aculeata Vallonia costata Vallonia excentrica

1968

Vertiginidae Vertigo pygmaea Columella edentula

1

WT/NBN NBN

1965

Vitrinidae Vitrina pellucida Trochulus hispidus Phenacolimax major

n/a Hairy Snail Greater Pellucid Snail

WT/NBN ID HBRC/NBN 1976 Local

Zonitidae Vitrea contracta Total Terrestrial Snails

WT 54 species

Doward 39

plus

SO51

15

Bithyniidae Bithynia tentaculata

1

NBN

2015

Lymnaeidae Galba truncatula Radix balthica

1 1

NBN NBN

1965 2015

1

NBN

2015

1 1 1 1 1

NBN NBN NBN NBN NBN NBN

1895 1965 1910 1895 1965 1895

1

NBN

2015

NBN

1965

Aquatic Snails

Neritidae Theodoxus fluviatilis Planorbidae Ancylus fluviatilis Anisus vortex Anisus leucostoma Gyraulus crista Hippeutis complanatus Planorbis planorbis Tateidae Potamopyrgus antipodarum Valvatidae Valvata piscinalis

81

Slugs Agriolimacidae Deroceras invadens Deroceras reticulatum

Tramp Slug 1 Field or Netted Slug

NBN 1

1965 NBN

1965

ID NBN ID NBN NBN NBN NBN NBN

1965 1965 1965 1965

Tree Slug Leopard Slug Ash-black Slug Green Cellar or Irish Yellow Slug Lemon Slug

NBN NBN NBN ID NBN

1965

Tandonia sowerbyi Tandonia (Milax) budapestensis Milax gagates

Sowerby’s Keeled Slug Budapest Slug Smooth Jet Slug NBN

NBN NBN NBN

Selenochlamys ysbryda

The Ghost Slug

Arionidae Arion ater Arion flagellus Arion hortensis Arion circumscriptus silvaticus Arion subfuscus Arion intermedius Arion distinctus Arion fasciatus Limacidae Lehmannia marginata Limax maximus Limax cinereoniger Limacus maculatus Malacolimax tenellus

Large Black Slug Green-soled Slug Garden Slug Silver False-keeled Slug Dusky Slug Hedgehog Slug Brown Soil Slug Rusty False-keeled Slug

1 1

Milacidae

1

iRec

AQUATIC Bivalves Sub-Total 12 10 in SO51 Sphaeriidae Pisidium supinum Pisidium casertanum Pisidium amnicum Pisidium subtruncatum Pisidium personatum Pisidium pulchellum Pisidium moitessierianum Pisidium henslowanum Pisidium nitidum Sphaerium rivicola Sphaerium corneum

1 1

1

NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN NBN

1920 1920 1965 1920 1965 1920 1920 1920 1965 1930 1965

1

NBN

?

1 1 1 1

Unionidae Unio tumidus

Mollusc species Total = 97

82

3.13 Checklist of subphylum Crustacea – Woodlice and freshwater shrimps Class Malacostraca Sub-order Isopoda - Woodlice and Waterlice Oniscidea – Terestrial Woodlice Trichoniscidae Androniscus dentiger Haplophthalmus danicus Trichoniscus pusillus agg Trichoniscus pygmaeus.

Rosy Woodlouse Spurred Ridgeback Pygmy Woodlouse Least Pygmy Woodlouse

M M local M NBN

Oniscidae Oniscus asellus

Common Shiny Woodlouse

Philoscidae Philoscia muscorum

Common Striped Woodlouse

M

Armadillidiidae Armidillidium vulgare Armidillidium depressum Armadillidium pictum

Common Pill Woodlouse Southern Pill Woodlouse Mottled Pill Woodlouse

M M NBN Rare

Porcellionidae Porcellio scaber Porcellio spinicornis

Common Rough Woodlouse Painted Woodlouse

M M

Platyarthidae Platyarthus hoffmannseggi

Ant Woodlouse

M

M

local

Total 12 species

Sub-order Amphipoda Gammaeridae Gammarus pulex agg.

Crangonyctidae Crangonyx pseudogracilis

Sub-order Decapoda Astacidae (Austropotamobius pallipes

Freshwater Shrimp

ID

Freshwater Shrimp

HBRC

White-clawed Crayfish EN

Plus other microscopic groups such as Copepoda e.g Daphniidae

83

HBRC. local status unclear)

3.14 Sub-phylum Myriapoda – Diploda (Millipedes) and Chilopoda (Centipedes) Millipedes - Diplopoda Glomeridae Glomeris marginata Pill Millipede Brachychaeteumatidae Brachychaeteuma melanops Craspedosomatidae Craspedosoma rawlinsii Nanogona polydesmoides Chordeumatidae Chordeuma proximum Melogona gallica Melogona scutellaris Polydesmidae Brachydesmus superus Polydesmus angustus Ophiodesmus albonanus Blaniulidae Proteroiulus fuscus Blaniulus guttulatus Nemasomatidae Nemasoma varicorne Julidae Julus scandinavius Ophyiulus pilosus Cylindroiulus caeruleocinctus Cylindroiulus punctatus Tachypodoiulus niger White-legged Millipede

UK Status

Source

LC

ID, WT

LC. NS

NBN

LC. NS LC

NBN NBN

LC LC LC

NBN NBN, WT NBN

LC LC LC

NBN NBN NBN

LC LC

NBN NBN

LC

NBN

LC LC LC LC LC

NBN NBN NBN NBN NBN

LC

ID

LC

ID

LC

ID

LC LC LC LC LC

NBN ID ID ID NBN

Note: Some records refer to SO51 and not specifically the Doward.

Centipedes – Chilopoda Himantariidae Haplophilus (Stigmatogaster) subterraneus Geophilidae Geophilus flavus Schendylidae Schendyla nemorensis Cyptopsidae Cryptops hortensis Lithobius forficatus Lithobius melanops Lithobius microps Lithobius variegatus

84

3.15. Checklist of phylum Entognatha: Non-insect Hexapods Note: All records should be considered unverified.

Class (Order) Collembola – Springtails

Source

(sub)Order Poduromorpha Entomobryidae Entomobrya albocinta Orchesella cincta Orchesella villosa Tetradontophora bielanensis prov.

Tomoceridae Tomocerus minor Tomecerus longicornis Cyphoderidae Cyphodoerus albinus Neanuridae Neamura muscorum Monobella grassei

ID ID ID ID ID ID ID ID ID

Onychiuridae Deuteraphora inermis Isotomidae Folsomia candida

ID ID

(sub)Order Neelipleona Dicyrtomidae Dicyrtomina saundersi ag.

Katiaanidae Sminthurinus aureus ag possibles

Class (Order) Diplura n/a

taxonomic status unclear

ID ID

Lepidocyrtus lanuginosus Schaefferia emucronata

SO51

NBN

Two-pronged Bristletails Campodea sp

85

ID

3.16 Annelids, Nematodes, Platyhelminths and allies

Phylum Annelida

Earthworms

Oligochaeta Lobworm

Lumbricus terrestris

Red worm

Phylum Nemotoda

Lumbricus rubella

Nematode Worms

Seen but none determined to family or species

Phylum Nematomorpha Gordiacea

Phylum Platyhelminths Freshwater Planarian

Gordian or Horsehair Worms probably Gordius aquaticus

Planarians, Leaches and Flukes Polycelis nigra sl

86

found in garden pond.

ID

Section 4. Vertebrates – Mammals, Birds Reptiles & Amphibians and Fish.

4.1 Checklist of Mammal – Page 88.

4.2 Checklist of Birds – Page 90.

4.3 Checklist of Reptiles & Amphibians – Page 93.

4.4 Checklist of Fish – Page 94.

87

4.1. Checklist of Mammals Order Insectivora Soricidae Common Shrew Sorex eraneus Pygmy shrew Sorex minutus Talpidae Common Mole Talpa europeus Erinaceidae European Hedgehog Erineceus europaeus

Frequent Suspected but unconfirmed based on size.

Frequent lower Ground Regular but not common.

Order Lagomorpha – Rabbits and Hares Rabbit

Oryctolegus cuniculus

Frequent

Order Chiroptera - Bats Sporano pipistrelle Pipistrellus pygmaeus Common pipistrelle Pipistrellus pipistrellus Western Barbastelle Barbastella barbastella Daubenton's Bat Leuconoe daubentonii Lesser Horseshoe Bat Rhinolophus hipposideros Greater Horseshoe Bat Rhinolophus ferrum-equinum Whiskered Bat Myotis mystacinus Natterer's Bat Myotis nattereri Brown Long-eared Bat Plecotus auritus Noctule Nyctalus noctule Serotine Eptisecus serotinus Order Rodentia - Rodents Bank Vole Clethrionomys glareolus Field Vole Microtus agrestis Yellow-necked Mouse Sylvaemus flavicollis Wood Mouse Sylvaemus sylvaticus House Mouse Mus musculus Brown rat Rattus norvegicus Hazel Dormouse Muscardinus avellanarius Grey squirrel Neosciuiurus carolinensis

Common Common Reported by Bat Group Near River Frequent Regularly encountered.

Reported NBN Reported NBN Regular Regular Regular

Near Threatenened Near Threatenened

Common Status unclear Common Believed common. Local status confused with above.

Occasional. Synanthropic.

Regular formerly abundant. Largely synanthropic.

Resident. Population unknown Abundant.

Order Artiodactyla – Even-toed Ungulates Fallow Deer Dama dama Roe Deer Capreolus capreolus Reeve’s Muntjac Deer Muntiacus reevesi Wild Boar Sus scrofa

Resident Resident Now resident Occasional visitor from over the river

Order Carnivora - Carnivores Red Fox Polecat (European) Badger (European) Otter Stoat (Least) Weasel American Mink

Vulpes vulpes Mustela putorius Meles meles Lutra lutra Mustela erminea Mustela nivealis Neovison vison

Resident Present status unknown. (Trailcam Miner's Rest NR) Resident Occasional, maybe regular on River Wye. Regular Regular Present on river (at least until 2021)

88

Notes: Radio tracked Pine Marten believed to have crossed the Doward from their release site in Forest of Dean. Also, there are other unconfirmed sightings.

Polecat also found dead on A40 road near Ganarew.

Muntjac Deer were first noted in 2014 and now seem to have a significant settled population across the Doward.

Water Voles may have occurred along parts of the Wye in the past and were present in Whitchurch parish until 1964.

Brown Hare has doubtless occurred in the parish and perhaps still does but is unlikely to visit the higher wooded terrain of the Doward.

There is a bizarre record of an Atlantic Grey Seal seen and photographed at Symonds Yat following winter floods in 2014. It apparently spent a few weeks in Monmouth. (Ref. Lower Wye Valley AONB website).

89

4.2. Checklist of Birds British vernacular names, scientific names and status as given in BOU British List (2020), however, official taxonomic sequence is not followed here.

Species that are resident, regular or occasional visitors since 2000.

Vernacular and scientific name Mute Swan Cygnus olor Canada Goose Branta canadensis Greylag Goose Anser anser Mallard Anas platyrhynchos

UK Status

Teal Anas crecca Mandarin Duck Aix galericulata Goosander Mergus merganser

A

Pheasant Phasianus colchicus Red-legged Partridge Alectoris rufa

C1E*

Cormorant Phalacrocorax carbo Grey Heron Ardea cinerea Little Egret Egretta garzetta

A

Common Buzzard Buteo buteo Red Kite Milvus milvus Sparrowhawk Accipiter nisus Goshawk Accipiter gentilis Peregrine Falcon Falco peregrinus (Common) Kestrel Falco tinnunculus (Eurasian) Hobby Falco subbuteo

AE*

A

Resident and commonly seen.

Regular sightings Frequent Regular Regular overhead or in quarries Occasional Occasional in summer

Moorhen Gallinula chloropus

A

Occasional on river

Woodcock Scolopax rusticola Curlew Numenius arquata Common Sandpiper Actitis hypoleucos Lapwing Vanellus vanellus

A

Former breeder now winter occasional.

Occasional overhead.

Occasional along river Formerly regular now scarce flying over

Lesser Black Backed Gull Larus fuscus Greater Black backed gull Larus marinus Common Gull Larus canus Herring Gull Larus argentatus Black-headed gull Chroicocephalus ridibundus

A

Wood Pigeon Columba palumbus Stock Dove Columba oenas Collared Dove Streptopelia decaocto

A

Cuckoo Cuculus canorus Tawny Owl Strix aluco

A A

Now occasional non-breeder.

Frequent resident

Common Swift Apus apus

A

Regular visitor in variable numbers

AC2 AC2E* AC2C4E* AC2C4E* C1E* A

C1E*

A A

AC3E* A AC3E* AE A

A A A

A A A A

A A

90

Comment

Resident along river Resident non-native along river Occasional with Canada Geese Resident along river Occasional on river or fly-over Regular up to 50 On river esp. late winter Frequent. Feral - released stock.

Occasional. Feral - released stock.

Frequent on river and as fly-over.

Regular around River Now regular along river

Commonly seen by river/overhead Occasional fly-over (hard to confirm).

Regular fly-over or on lower fields.

Frequent flyover Regular flyover Common resident Frequent resident Frequent near lower dwellings

Green Woodpecker Picus viridis Great Spotted Woodpecker Dendrocopos major Lesser Spotted Woodpecker Dryobates minor

A A

Frequent resident Common resident.

Elusive resident in very small numbers.

Kingfisher Alcedo atthis

A

Frequent along River

(Barn) Swallow Hirundo rustica House Martin Delichon urbicum Sand Martin Riparia riparia

AE

Frequent in summer Frequent in summer Regular visitor along river

Pied Wagtail Grey Wagtail Tree Pipit Meadow Pipit Skylark

Motacilla alba

A

Motacilla cinerea Anthus trivialis Anthus pratensis Alauda arvensis

A

A

A A

A

Resident Resident Summer visitor in small numbers Regular migrant Regular overhead migrant

Dipper Cinclus cinclus

A

Occasional from river

Dunnock Prunella modularis Robin Erithacus rubecula (Common) Redstart Phoenicurus phoenicurus Stonechat Saxicola rubicola (Northern) Wheatear Oenanthe oenanthe Blackbird Turdus merula Song thrush Turdus philomelos Mistle thrush Turdus viscivorus Redwing Turdus iliacus Fieldfare Turdus pilaris

A

A

Common resident Common resident A few pairs breed on Little Doward Occasional visitor. 2022 a pair on Little Doward Occasional on passage esp. Little Doward Common resident Common resident Resident in small numbers Winter visitor with notable spring gatherings.

Winter visitor in variable numbers

Cetti's Warbler Cettia cetti Garden warbler Sylvia borin Willow warbler Phylloscopus trochilus Wood warbler Phylloscopus sibilatrix Chiffchaff Phylloscopus collybita Blackcap Sylvia atricapilla Sedge Warbler Acrocephalus schoenobaenus Common Whitethroat Curruca communis Lesser Whitethroat Curruca curruca Goldcrest Regulus regulus Firecrest Regulus ignicapilla Wren Troglodytes troglodytes Spotted Flycatcher Muscicapa striata Pied flycatcher Ficedula hypoleuca

A

Occasional on riverbank

A

Regular breeder in small numbers esp. by river Regular breeder in small numbers Declining summer breeder in Lords Wood Common summer visitor Common summer visitor Occasional visitor Summer visitor in small numbers Occasional visitor Common resident Uncommon visitor.

Common resident Regular breeder in small numbers Visitor & scarce breeder (2+ pairs in 2017)

Blue tit Cyanistes caeruleus Great tit Parus major Marsh tit Poecile palustris Coal tit Periparus ater Long-tailed Tit Aegithalos caudatus Tree creeper Certhia familiaris Nuthatch Sitta europaea

A

A A

A A A A A A A A

A A A A A A A A A A A A

A A A A A A

91

Common resident Common resident.

Fairly common resident Common resident Common resident Fairly common resident Fairly common resident

Jay Garrulus glandarius Magpie Pica pica Jackdaw Coloeus monedula Carrion Crow Corvus corone Rook Corvus frugilegus Raven Corvus corax

A

A

Fairly common resident Resident in lower areas uncommon on top.

Resident Common resident Visitor to lower fields and flyover Regular visitor in small numbers

Starling Sturnus vulgaris

A

Frequent, mainly in lower areas.

House Sparrow Passer domesticus Crossbill Loxia curvirostra Hawfinch Coccothraustes coccothraustes Bullfinch Pyrrhula pyrrhula Greenfinch Chloris chloris Chaffinch Fringilla coelebs Goldfinch Carduelis carduelis Brambling Fringilla montifringilla Siskin Spinus spinus Lesser Redpoll Acanthis cabaret Linnet Linaria cannabina Reed Bunting Emberiza schoeniclus

A

Frequent in small groups.

Regular in small numbers esp. winter.

Occasional in small groups. May have bred.

Frequent Frequent Common in variable numbers Common Resident Regular in small numbers usually late winter Frequent resident Regular Regular Regular in riverside scrub

Doward Total

A A A A

A A A A A A A A A A A

97 species

Additional species reported either as rare visitors or regulars just outside the Doward Whooper Swan Cygnus cygnus Ae Near Goodrich bridge (2016) Common Snipe Gallinago gallinago A Lewstone Feb'22 Gargany Spatula querquedula A Reported. Lewstone (ca. 2017 pers com) Little Owl Athene noctua C1E* Farmland reported. Huntsham, Lewstone Yellow Wagtail Motacilia flava C Flyover open areas Hoopoe Upupa epops AE Reported 2003 HBRC Grasshopper Warbler Locustella naevia A One Report.

Yellowhammer Emberiza citrinella A Scarce flyover Additional species reported prior to year 2000.

Nightjar Caprimulgus europaeus A Nightingale Luscinia megarhynchos A

92

Reported in 1970s Reported to be present until 1970’s

4.3 Checklist of Reptiles and Amphibians Class Amphibia.

Order Arodela - Tailless Amphibians Bufonidae – True Toads Common Toad

Bufo bufo

Common throughout.

Ranidae – Typical Frogs Common Frog

Rana temporaria

Frequently found.

Order Anura – Tailed Amphibians Salamandridae - Newts and salamanders Smooth (Common) Newt Triturus vulgaris Palmate Newt Triturus heveticus

Frequent ponds but true status unclear Common ponds.

Great (Northern) Crested Newt

Status unclear. 1 old Whitchurch record

Triturus cristatus

Class Reptilia Order Squamata – Snakes and Lizards Lacertidae – Typical Lizards Common (Viviparous) Lizard

Zootoca vivipara

Status unclear. Occurs nearby.

Anguidae – Slow Worms Slow Worm

Anguis fragilis

Common

Columbridae – Typical Snakes Grass Snake

Natrix natrix

Frequent

Viperidae - Vipers Adder

Viperus berus

Status unclear. Occurs nearby.

Taxonomy and nomenclature reference Kwet, A. (2009). European Reptile and Amphibian Guide.

New Holland Publishers.

93

4.4 Checklist of Fish: Superclass Agnatha – Jawless Fish. Class Cephalaspidomprphi Petromyzonidae - Lampreys Sea Lamprey Petromyzon marinus River Lamprey Lampetra fluviatilis Brook Lamprey Lampetra planeri Superclass Gnathostomata – Jawed Fish. Class Chondrichthyes Anguillidae European Eel Anguilla anguilla Clupeidae Twaite Shad Allis Shad

Alosa fallax.

Alosa alosa

Esocidae Pike

Esox lucius.

Salmonidae Atlantic Salmon Brown Trout Grayling

Salmo salar Salmo trutta Thymallus thymallus

Cyprinidae Barbel Gudgeon Roach Chub Dace Minnow

Barbus barbus Gobio gobio Rutilus rutilus, Squalius cephalus Leuciscus leuciscus Phoxinus phoxinus

Percidae Perch

Perca fluviatilis

Cottidae Bullhead

Cottus gobio

Cobitidae Stone Loach

Barbatula barbatula.

Scomberesocidae Skipper

Scomberesox saurus.

Pleuronectidae Flounder

Platichthys flesus

All records here come from surveys of the Wye by the Environment Agency via NBN. Edwards & Brooker (1982) in their book The Ecology of the Wye claim 29 fish species in the Wye catchment.

Taxonomy and nomenclature follow Miller, PJ et al (1997). Fish of Britian and Europe. Collins.

94

95