Northeast Area Monitoring and Assessment Program (NEAMAP)
Mid-Atlantic Nearshore Trawl Program
Progress Report: Fall 2007 Survey Data Summary
20 March 2008
Submitted to: Atlantic States Marine Fisheries Commission Washington, DC
By: Christopher F. Bonzek James Gartland J. David Lange Robert J. Latour, Ph.D.
Department of Fisheries Science Virginia Institute of Marine Science College of William and Mary Gloucester Point, VA
NEAMAP ASMFC Progress Report I.
Project Title: Data collection and analysis in support of multispecies stock assessments in the mid-Atlantic: Northeast Area Monitoring and Assessment Program Nearshore Trawl Program.
II.
Grantee State and Contact Name: Virginia/Virginia Institute of Marine Science – Christopher F. Bonzek
III.
Project Period: 1 August 2005 – 31 May 2009 Reporting Period: 1 August 2007 – 31 January 2008
IV.
Project Description: This is a new fisheries-independent bottom trawl survey operating in the near coastal ocean waters of the Mid-Atlantic region. The survey is an element of the ASMFC Northeast Area Monitoring and Assessment Program (NEAMAP) and is designed to sample fishes and invertebrates from coastal waters bounded by the 20ft.and 60ft. depth contours between Montauk, New York and Cape Hatteras, North Carolina and waters between the 60ft.and 120ft. depth contours in Rhode Island Sound and Block Island Sound using a bottom trawl. The main objective of the survey is the estimation of biomass, length and age structures, various other assessment related parameters and diet compositions of select finfishes inhabiting the area.
V.
Project Summary/Accomplishments: The fall 2007 survey was successfully completed during a research cruise which occurred between 27 September and 20 October 2007. A total of 150 stations, the target number, were sampled. About 1.1 million individual fishes weighing almost 50,000kg and representing about 130 species (including selected invertebrates which where processed similarly to fishes) were captured. Individual length measurements were recorded for over 73,000 specimens. Lab processing is proceeding on the 5,150 otoliths and 3,905 stomach samples which were collected (2,433 otoliths and 1,930 stomachs have been fully processed as of the date of this report). A full report is attached to this standard project summary.
VI.
Challenges/Changes: Beyond completion of laboratory samples, no significant challenges remain for this contract segment.
VII.
Participants: Primary program personnel remain unchanged.
VIII. Quality Assurance: Previous progress reports provided brief descriptions of quality assurance procedures in selecting fishing gear, conducting fishing operations, and processing the catch. These are interwoven into the attached report as well. Data collected during the survey have been processed through several data quality checks which were previously developed for other survey work and new checks developed specifically for NEAMAP. IX.
Funding Status: Expenditures have been generally in line with expectations. We anticipate sufficient funds will be available until Research Set Aside Program funds are dispersed later in 2008.
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X.
Future Activities: The future of this program is dependent upon continued funding. We anticipate sufficient funds to complete both the spring and fall 2008 cruises.
XI.
Presentations/Public Outreach: Since completion of field sampling, presentations of survey results have been (or will be) made as follows: • • • • • • •
February 2008: Mid-Atlantic Fishery Management Council February 2008: Cape May NJ Party and Charter Boat Association February 2008: NMFS NEFSC Trawl Advisory Panel February 2008: Bass Pro Shops Fishing Classic (Hampton, VA), Booth exhibit March 2008: NEAMAP Operations Committee March 2008: NEAMAP Board April 2008: New England Fishery Management Council
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Introduction Concerns regarding the status of fishery-independent data collection from the continental shelf waters between Cape Hatteras, North Carolina and the U.S. / Canadian border led the Atlantic States Marine Fisheries Commission’s (ASMFC) Management and Science Committee (MSC) to draft a resolution in 1997 calling for the formation the Northeast Area Monitoring and Assessment Program (NEAMAP) (ASMFC 2002). NEAMAP is a cooperative state-federal program modeled after the Southeast Area Monitoring and Assessment Program (SEAMAP), which had been coordinating fishery-independent data collection south of Cape Hatteras since the mid-1980s (Rester 2001). The four main goals of this new program directly address the deficiencies noted by the MSC for this region and include 1) developing fishery-independent surveys where current sampling is either inadequate or absent 2) coordinating data collection amongst existing surveys as well as any new surveys 3) providing for efficient management and dissemination of data and 4) establishing outreach programs (ASMFC 2002). The NEAMAP Memorandum of Understanding was signed by all partner agencies by July 2004. One of the first major efforts of the NEAMAP was to design a trawl survey intended to operate in the coastal zone of the Middle Atlantic Bight (MAB - i.e., Montauk, New York to Cape Hatteras, North Carolina). While the National Marine Fisheries Service (NMFS) Northeast Fisheries Science Center’s (NEFSC) bottom trawl survey has been sampling from Cape Hatteras to the U.S. / Canadian border in waters less than 91.4m since 1963, few stations are sampled in waters less than 27.4m due to the sizes of the sampling area and vessels (NEFSC 1988, R. Brown, NMFS, pers. comm). In addition, of the six coastal states in the MAB, only New Jersey conducts a fishery-independent trawl survey in its coastal zone (Byrne 2004). This new NEAMAP Inshore Trawl Survey is intended to fill the aforementioned gap in fisheryindependent survey coverage, which is consistent with the program goals. In early 2005, the ASMFC made $250,000 of “plus-up” funds that it had received through the Atlantic Coastal Fisheries Cooperative Management Act (ACFCMA) available for pilot work in an effort to assess the viability of the NEAMAP Inshore Trawl Survey. The Virginia Institute of Marine Science provided the sole response to the Commission’s request for proposals and was awarded the funding in August 2005. Two brief pre-pilot cruises and the full pilot cruise were conducted in 2006 (Bonzek et al., 2007). Early in 2007 ASMFC bundled funds from a combination of sources which were sufficient to begin full scale sampling operations in the fall of 2007. This report summarizes results from the fall 2007 cruise. Two significant changes to the area sampled by the NEAMAP Nearshore Trawl Program occurred prior to the fall 2007 cruise: • In 2007 NEFSC took delivery of the FSV Henry B. Bigelow, began preliminary sampling operations, and determined that the vessel could safely operate in waters as shallow as 18.3m. NEFSC then made a determination that future surveys would likely extend inshore to that depth contour (R. Brown, NMFS, pers. comm.). The NEAMAP Operations Committee subsequently decided that the offshore boundary of the NEAMAP survey coastal sampling (i.e., Montauk to Cape Hatteras) should be realigned to coincide
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•
with the inshore boundary of the NEFSC survey, and that NEAMAP should discontinue sampling between the 18.3m and 27.4m contours along the coast. NEFSC contributed significant funds toward NEAMAP full implementation with the provision that the additional under-sampled areas of Block Island Sound and Rhode Island Sound be added to the NEAMAP sampling area. These areas are deeper than other NEAMAP regions but from a ‘distance from shore’ standpoint are within the range covered by NEAMAP in other states.
Methods Station Selection Primary consideration in regards to survey stratification was consistency with the NMFS bottom trawl surveys. However, those surveys will be redesigned and re-stratified for 2009 (and beyond) and so re-stratification for the inshore NEAMAP areas was open for consideration as well. Examination of existing NMFS strata revealed that the major divisions among survey areas (latitudinal divisions from New Jersey to the south, longitudinal divisions off Long Island) generally corresponded well with major estuarine outflow areas. Therefore these boundary definitions, with minor modifications so that regional boundaries would more closely correspond to state borders, were used for the NEAMAP survey. However, examination of the current NMFS depth stratum definitions reveals that in some areas (primarily off the southern states) current stratum boundaries do not correspond well to actual depth contours. Depth stratum assignments were redrawn using depth sounding data from the National Ocean Service using depth strata 20ft.-40ft. and 40ft.-60ft. Finally, each stratum was subdivided into a grid pattern of potential sampling locations, with each cell measuring 1.5 x 1.5 minutes (2.25sq. nm). The number of stations (cells) selected for each stratum was assigned by proportional sampling according to surface area within the stratum, with a minimum of two stations per stratum. During the pilot cruise, an equal number of stations was selected for each depth stratum within a particular region. This assured that sufficient sampling occurred in the small, shallow strata closest to shore. As the large offshore (60ft. – 90ft.) strata were not sampled in the fall of 2007, this procedure was no longer required and straight proportional sampling was implemented. Species Priority Lists During the survey design phase, the NEAMAP Operations Committee developed a set of species priority lists. Priority ‘A’ species were to be subjected to the full processing procedure (see Procedures at Each Station below) at each station in which they were collected. Compared to the list used for the 2006 pilot survey, several Priority ‘A’ species were added due to the expanded survey area (should lead to collections of additional species of management importance) and the requests of the Mid-Atlantic Fisheries Management Council. Priority ‘B’ species were to be sampled for full processing as time allowed. Priority ‘C’ species would only be taken for full processing if sampling of A and B species would not be affected. These three categories might be summarized as ‘must have’ ‘great to have’ and ‘nice to have,’ respectively. All other species (here called Priority ‘D’) were to have aggregate weights recorded and all or an appreciable subsample to be measured. A fifth category (‘E’) was later defined, including
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species which required special handling. This category included sharks (other than dogfish) and sturgeon, which were measured, tagged, and released; and selected invertebrates which were processed similarly to Priority D fish species. Species included in categories A-C are presented below (Table 1). Table 1. Species priority lists (categories A-C only). A LIST
Atlantic Cod Black Sea Bass Bluefish Butterfish Haddock Pollock Scup Silver Hake Striped Bass Summer Flounder Weakfish Winter Founder American Shad Atlantic Menhaden Atlantic Croaker Monkfish Skate and Ray Species Smooth Dogfish Spiny Dogfish Spot Yellowtail Flounder
Alewife Atlantic Herring Atlantic Mackerel Black Drum Blueback Herring Red Drum Speckled Trout Tautog
Gadus morhua Centropristis striata Pomatomus saltatrix Peprilus triacanthus Melanogrammus aeglefinus Pollachius virens Stenotomus chrysops Merluccius bilinearis Morone saxatilis Paralichthys dentatus Cynoscion regalis Pleuronectes americanus B LIST Alosa sapidissima Brevoortia tyrannus Micropogonias undulatus Lophius americanus Mustelus canis Squalus acanthias Leiostomus xanthurus Limanda ferruginea C LIST Alosa pseudoharengus Clupea harengus Scomber scombrus Pogonias cromis Alosa aestivalis Sciaenops ocellatus Cynoscion nebulosus Tautoga onitis
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Gear Performance Wingspread, doorspread, and headrope height were measured on each tow during the fall 2007 cruise using a digital Netmind Trawl Monitoring System. Wingspread sensors were positioned on the middle net ‘jib’ in accordance with NFMS procedures. The headrope sensor was mounted at the midpoint of the headrope. A catch sensor was mounted in the cod-end, set to signal when the catch reached roughly 5,000lbs. GPS coordinates and vessel speed were recorded every 60 seconds using the ship’s chartplotting software. These data can be used to plot tow tracks for each station. The same computer used to record Netmind readings was also employed to plot station locations (and the corners of each sampling cell) and to run the countdown clock for each tow. Procedures at Each Station All fishing operations were conducted during daylight hours. Each tow was 20 minutes in duration with a target tow speed of between 2.9 and 3.3 knots. No tows were truncated due to known hangs in the tow path, surface traffic etc. One tow was terminated at 15 minutes due to an anticipated (and realized – 6,400kg) large catch, as indicated by triggering of the catch sensor midway through the tow. At each station several standard parameters were recorded. These included: • All necessary station identification parameters (date, station number, stratum, depth). • All necessary vessel operation parameters (beginning and ending GPS position, beginning and ending tow times, compass course, engine RPMs). • All necessary gear identification and operational parameters (net type code and net number, door type code and door numbers, amount of cable deployed). • Atmospheric and weather data (air temperature, wind speed, wind direction, general weather state, sea state, barometric pressure). • Hydrographic data at the surface and at the bottom (water temperature, salinity, and dissolved oxygen). Upon arrival near a sampling cell, the Captain and Chief Scientist jointly determined the desired starting point and tow path. Flexibility was allowed with regard to these parameters such that a clear tow could be accomplished while staying within the boundaries of the defined cell. Hydrographic data were taken prior to the beginning of each tow, except rarely when taking these readings would have delayed the tow until after sunset; in these cases hydrographic data were recorded at the completion of the tow. Vessel crew were responsible for all aspects of deployment and retrieval of the fishing gear. Due to the relatively shallow waters, 75fm. or less of warp was set out at all stations. One scientist was present in the wheelhouse during deployment and retrieval. The Captain signaled when the gear was fully set (winch brakes engaged), at which time the Netmind software and the countdown clock were both activated. At the conclusion of each tow, the scientist signaled the Captain when the clock reached zero, haulback commenced, and the Netmind recording software was stopped. Vessel crew dumped the catch into one of two enclosed locations on deck for sorting.
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The catch was sorted by species and modal size group. Aggregate biomass (kg) was measured for each species-size group combination. For priority A species, and often for priority B and C species, a subsample of five individuals from each group was selected for full processing (see next paragraph). For certain common priority B species including spot (Leiostomus xanthurus), Atlantic croaker (Micropogonias undulatus), skates, rays, and dogfish only three individuals per group were sampled for full laboratory processing. Data collected from each subsampled specimen included length (to the nearest millimeter), total and eviscerated weight (measured in grams, accuracy depended upon the balance on which individuals were measured), and macroscopic sex and maturity stage (immature, mature-resting, mature-ripe, mature-spent) determination. Stomachs were removed (except for spot and butterfish, for which previous sampling indicated that little useful data could be obtained from the stomach contents) and those containing prey items were preserved for subsequent examination. Otoliths or other appropriate ageing structures were removed from each subsampled specimen for later age determination. All specimens not selected for the complete processing were weighed (aggregate weight), and individual length measurements were recorded for either all or a large proportion, in accordance with approved subsampling procedures when necessary. Laboratory Methods Otoliths (or, depending upon the species, other appropriate ageing structures) were (and are being) prepared according to methodology established for other VIMS surveys. One otolith was selected and mounted on a piece of 100 weight paper with a thin layer of Crystal Bond. A thin transverse section was cut through the nucleus of the otolith using two Buehler diamond wafering blades and a low speed Isomet saw. The section was then mounted on a glass slide and covered with Crystal Bond. If necessary, the section was wet-sanded to an appropriate thickness before being covered with Crystal Bond. Some smaller, fragile otoliths were read whole. Both sectioned and whole otoliths were most commonly read using transmitted light under a dissecting microscope. Age was determined as the mode of three independent readings, one by each of three readers. Stomach samples were (and are being) analyzed according to standard procedures (Hyslop 1980). Prey were identified to the lowest possible taxon. Experienced laboratory personnel are able to process, on average, approximately 30 to 40 stomachs per person per day. Analytical Methods (Abundance) Estimates of abundance are expressed in terms of minimum trawlable number or biomass according to the general formula: cA , N= a
(1)
where N is the minimum number (or biomass) of fish present within the sampling area that are susceptible to the sampling gear, c is the mean number (or weight) of fish captured per tow, a is the area swept by one trawl tow, and A is the total survey area. Specifically, abundance was calculated in accordance with standard stratified random sampling:
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n
s Nˆ = ∑ As Nˆ s ,
(2)
s =1
where As is the area of stratum s, ns is the total number of strata in which striped bass were ˆ captured, and N s is an estimate of the mean area-swept catch in stratum s given by: nt ,s
Nˆ s =
∑ aˆ
ci
i =1
nt , s
i
,
(3)
In equation (3), ci and aˆi represent the catch (number or weight) and an estimate of the trawl area-swept at sampling location i, respectively, and nt,s is the number of tows in stratum s. Note that the ai estimates were calculated using vessel GPS data for distance towed and net mensuration gear for measurements of net opening (an average value was calculated from the measurements taken during each tow). As no correction is made for gear efficiency these estimates represent the minimum number (or biomass) of fish present within the sampling area that are susceptible to the sampling gear. This method produces estimates of abundance for each stratum, which are totaled to produce estimates for the entire survey area. As regional stratum boundaries were drawn to generally correspond with state borders, estimates of abundance (and certain other stock parameters) can be produced on a state-specific basis. While usually not biologically meaningful, for some parameters it was considered worthwhile to present results in this way due to the potential usefulness for fishery managers. Analytical Methods (Sex Ratios) Sex ratios were determined by summation of data from fully processed specimens. For this and several other stock parameters, data from fully processed specimens are expanded to the entire sample (i.e., catch level) for parameter estimation. Because workup procedures result in differential subsampling rates among size groups, failure to account for such factors would bias resulting stock parameter estimates. In the NEAMAP database each specimen has a calculated expansion factor associated with it which represents the number of fish that the specimen represents in the total sample for that station. Analytical Methods (Length Frequency) Length frequency histograms were designed using 10mm bins and the expansion factors as previously described. Length bins were identified using the bin midpoint (e.g. the 250mm bin represents individuals between 245mm and 254mm). Analytical Methods (Length-Weight Regressions) Power regressions for length-weight relationships were calculated and plotted by sex as well as both sexes combined. No tests for differences in growth by sex were performed. No expansion factors were used in these calculations as they do not depend on subsampling rates.
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Analytical Methods (Maturity Regressions) Logistic regressions were calculated and plotted separately for males and females. No expansion factors were used in these calculations as they do not depend on subsampling rates. Analytical Methods (Diets) Diets for each species were determined by estimating the mean proportional contribution of prey type (k) to predator (x) by weight or number (Wk,x) using the following equation: n
Wk , x =
∑M i =1
i,x
qi ,k , x ,
n
∑M i =1
(4)
i,x
where
qi ,k , x =
wi ,k , x wi , x
,
(5)
where Mi,x is the number of predator x captured at station i, wi,k,x is the total weight (or number) of prey type k encountered in the stomachs of predator x collected at station i, and wi,x is the total weight (or number) of all prey items encountered in these stomachs (Buckel et al. 1999). This cluster sampling estimator was used since trawl collections yield a cluster of each predator at each station. Results Gear Performance As was the case during the pilot survey and prior work, the 4-seam net performed consistently within the expected parameters (Figure 1). The addition of doorspread sensors provided valuable supplementary information. Mounting these sensors onto the relatively small trawl doors however (Figure 2), decreased the efficiency (i.e., spreading power) of the doors, and early in the cruise wingspread measurements were 1-2m less than previously achieved. Slight modifications to the warp and backstrap attachment points on the doors, combined with additional scope, brought the net back within specifications and provided the expected door spread measurements. Discounting the early stations at which the gear was slightly underspread, there did not appear to be an increasing trend in either the doorspread or wingspread measurements as was observed in the 2006 pilot survey. The same net was used for the fall 2007 survey as was used for the pilot survey. We will monitor whether this phenomenon appears in future surveys using new nets. No major gear damage occurred during the survey.
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Stations Sampled Based on a specified sampling rate of one station per 30sq.nm, the target number of stations to be sampled was 150 for the entire sampling area (1,938 cells x 2.25sq.mi. per cell / 30 stations per sq.nm. = 145 stations) and 150 stations were successfully occupied. The number of stations available and the number sampled in each stratum is given (Table 2). Of the 150 stations sampled, 129 were sampled within the specified primary sampling cell and 21 were chosen from the available randomly selected alternate sites, due to issues such as known hangs or other obstructions, fixed gear, or vessel traffic. One extra station was occupied in Region 12 (Virginia) to make up for one station not sampled in BI Sound (data from this station were assigned to Region 12, not to BI Sound). The highest number of alternate stations occupied was in BI Sound (2 out of 10) and RI Sound (7 out of 17) due to a high degree of caution, to unfamiliarity with the area, and to a relatively small number of towable locations in this area. For future surveys we anticipate obtaining a better sample of known towable locations through cooperation with local industry representatives. A region-by-region summary of these results is presented (Table 3). Maps comparing selected and actual stations sampled are shown (Figure 3).
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Table 2. Number of available sample cells and number sampled in each stratum. Region
State*
Stations Sampled 20ft.-40ft. Stations sampled
40ft. – 60ft.
Total cells
Stations sampled
Total cells
Stations sampled
Total cells
Stations sampled
Total cells
Stations per sq. nm.
60ft. – 90ft.
90ft. – 120ft.
RI Sound
RI
6
85
11
161
32.6
BI Sound
RI
4
42
5
88
32.5
1
NY
0
0
2
19
21.4
2
NY
2
8
3
19
12.2
3
NY
2
16
3
28
19.8
4
NY
2
16
3
29
20.3
5
NY
2
27
3
45
32.4
6
NJ
2
20
3
42
27.9
7
NJ
4
49
6
97
32.9
8
NJ
2
32
7
90
30.5
9
DE
4
53
8
113
31.1
10
MD
2
33
8
114
33.1
11
VA
5
62
9
122
29.6
12
VA
5
60
6
67
26.0
13
VA
7
94
11
142
29.5
14
NC
2
24
5
61
27.3
15
NC
2
25
4
55
43
519
81
1043
Total
30.0 10
127
16
249
29.1
* Note that region boundaries are not perfectly aligned with all state boundaries: • Some stations in RI Sound may occur in MA • Some stations in BI Sound may occur in NY • Region 5 spans the NY-NJ Harbor area • Some stations in Region 9 may occur in NJ
Table 3. Number of primary and alternate stations occupied in each region. Region
Primary Alternate Total Region Primary Alternate Stations Stations Stations Stations 10 7 17 8 1 RI Sound 8 7 2* 9 11 1 BI Sound 9 1 1 2 10 0 1 10 4 1 5 13 1 2 11 5 0 5 9 2* 3 12 5 0 5 15 3 4 13 5 0 5 7 0 5 14 4 1 5 5 1 6 15 10 0 10 7 Total 129 21 * One planned station in BI Sound was not sampled; an extra station was occupied in Region 12.
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Total 9 12 10 14 11 18 7 6 150
On the 17 full sampling days (i.e., no long steam times or port calls), an average of 7.9 stations per day were sampled. Counting all 24 days at sea, including transit days and partial sampling days, the number of stations averaged 6.25. Day-by-day vessel activities and work schedules are presented (Table 4). Table 4. Summary of activities conducted during each day at sea during the fall 2007 NEAMAP cruise. Hours Worked and Stations Sampled Each Day
Date 24-Sep 25-Sep 26-Sep 27-Sep 28-Sep 29-Sep 30-Sep 1-Oct 2-Oct 3-Oct 4-Oct 5-Oct 6-Oct 7-Oct 8-Oct 9-Oct 10-Oct 11-Oct 12-Oct 13-Oct 14-Oct 15-Oct 16-Oct 17-Oct 18-Oct 19-Oct 20-Oct 21-Oct
12: 6: 00 00 AM AM Final Survey Preparations Steaming Day - Hampton to Montauk Steaming Day - Hampton to Montauk
Time of Day 12: 00 PM
Crew Change / Survey Demo. Montauk
Survey Demo. Cape May
No Fishing / Crew Change and Resupply in Hampton No Fishing / Crew Change and Resupply in Hampton
Survey Finished
6: 00 PM
11: 00 No. PM Station 0 0 0 7 7 7 7 6 2 7 8 9 10 10 2 9 9 7 9 6 0 0 1 7 7 8 4 0
= Fishing hours = Personnel hours
Catch Summary A total of 1,101,152 specimens weighing 49,868kg were collected during the fall 2007 survey. A total of 73,473 individuals were measured (laying all individuals head-to-tail 11,465m, or 7.1miles, of fish were measured). Of those specimens taken for full workup, 5,150 otoliths (or other ageing structures) were taken and 3,905 full stomachs were preserved for later analysis. On average at each station, 7,132 (range 75 – 151,598) specimens were captured (Figure 4) weighing 332kg (range 7.4kg – 6,396kg) (Figure 5), 490 specimens were measured (range 57 – 1,394), and 34 specimens were processed for the full workup (range 4 – 77). At each station, an average of 20.4 species was captured (range 7 – 36) (Figure 6). The number of specimens processed for each species, separately for each priority category, is summarized in Table 5.
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Species Data Summaries Several graphical data summaries are shown for each species (Figures 7-173). Species are organized alphabetically. For priority A species, most or all of the following figures are presented (at the time of report preparation 2,433 otoliths and 1,930 stomachs for priority A species have been analyzed): • Distribution maps showing catch rates by number and biomass for all stations. • Bar graphs showing the minimum trawlable number (MTN) and biomass (MTB) by state, annotated with estimates of the overall MTN and MTB and percent standard error for each. • Length-frequency histograms annotated with the total number of specimens captured and measured and the number of otoliths and stomachs removed for processing. • Length-frequency histograms by age. • Histograms of sex ratio by state, and for species with adequate sample size, by size groups, annotated with the number of specimens examined. • Logistic regressions for maturity, by sex, annotated with sizes at 50% and 99% maturity. • von Bertalanffy growth regressions, by sex if appropriate. • Age distribution graphs (ageing has been completed only for black sea bass, scup, summer flounder, weakfish, and striped bass (not shown) ) shown with the total number of specimens by age if all specimens had been aged. • Length-weight power regressions for sexes combined and separately. • Diet compositions by weight and number, with prey types separated into broad taxonomic groups, annotated with the number of stomachs analyzed. These data summaries are numbered as follows: • Black seabass – Figures 33-42. • Bluefish – Figures 43-49. • Butterfish – Figures 59-64. • Scup – Figures 96-105. • Silver hake – Figures 106-111. • Summer flounder – Figures 139-148. • Weakfish – Figures 149-158. • Winter flounder – Figures 159-168.
For priority B species, some or all of the following data summaries are presented: • Distribution maps showing catch rates by number and biomass for all stations. • Bar graphs showing the MTN and MTB by state, annotated with estimates of the overall MTN and MTB and percent standard error for each. • Length-frequency histograms annotated with the total number of specimens captured and measured and the number of otoliths (or vertebrae for elasmobranchs) and stomachs removed for processing. • Histograms of sex ratio by state, and for species with adequate sample size, by size groups, annotated with the number of specimens examined. • Logistic regressions for maturity, by sex, annotated with sizes at 50% and 99% maturity.
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•
Length-weight power regressions for sexes combined and separately.
These data summaries are numbered as follows: • Atlantic croaker – Figures 12-17. • Atlantic menhaden – Figures 18-23. • Bluntnose stingray – Figures 50-52. • Bullnose ray – Figures 56-58. • Clearnose skate – Figures 65-70. • Cownose ray – Figures 71-73. • Little skate – Figures 82-86. • Smooth butterfly ray – Figures 115-117. • Smooth dogfish – Figures 118-123. • Spot – Figures 127-132. • Winter skate – Figures 169-173. No Priority C species were captured in sufficient numbers to allow meaningful data summaries to be presented. Several other species were captured in significant numbers as well (labeled as Priority D in Table 5). Data summaries for these species that are shown are: • Distribution maps showing catch rates by number and biomass for all stations. • Bar graphs showing the MTN and MTB by state, annotated with estimates of the overall MTN and MTB and percent standard error for each. • Length-frequency histograms annotated with the total number of specimens captured and measured. These data summaries are numbered as follows: • Atlantic spadefish – Figures 24-26. • Atlantic thread herring – Figures 27-29. • Bay anchovy – Figures 30-32. • Kingfish spp. – Figures 79-81. • Northern searobin – Figures 90-92. • Pinfish – Figures 93-95. • Silver perch – Figures 112-114. • Spanish mackerel – Figures 124-126. • Striped anchovy – Figures 133-135. • Striped searobin – Figures 136-138. Finally, several invertebrate species (or species groups) and selected sharks were captured in sufficient numbers to be of note (described as Priority D species in Table 5). Data summaries for these species that are shown are: • Distribution maps showing catch rates by number and biomass for all stations. • Bar graphs showing the MTN and MTB by state, annotated with estimates of the overall MTN and MTB and percent standard error for each.
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• • •
Length-frequency histograms annotated with the total number of specimens captured and measured. Histograms of sex ratio by state, annotated with the number of specimens examined (horseshoe crab only). Length-weight power regressions for sexes combined and separately (horseshoe crab only).
These data summaries are numbered as follows: • American lobster – Figures 7-11. • Brown shrimp – Figures 53-55. • Horseshoe crab – Figures 74-78. • Loligo squid – Figures 87-89.
Literature Cited Atlantic States Marine Fisheries Commission (ASMFC). 2002. Development of a Cooperative State/Federal Fisheries Independent Sampling Program. ASMFC Document, Washington, DC. Bonzek, C.F., J. Gartland, R.J. Latour. 2007. Northeast Area Monitoring and Assessment Program (NEAMAP) Mid-Atlantic Nearshore Trawl Program Pilot Survey Completion Report. ASMFC. 97pp. Byrne, Don. 2004. Counting the fish in the ocean. Online. Internet. <http://www.state.nj.us/dep/fgw/artoceancount.htm> Hyslop, E. J. 1980. Stomach contents analysis – a review of methods and their application. Journal of Fish Biology 17:411-429. NEFC. 1988. An evaluation of the bottom trawl survey program of the Northeast Fisheries Center. NOAA Tech. Memo. NMFS-F/NEC-52, p. 83. Rester, J.K. 2001. Annual report to the Technical Coordinating Committee Gulf States Marine Fisheries Commission. Report of the Southeast Area Monitoring and Assessment Program (SEAMAP) to the Gulf States Marine Fisheries Commission, Ocean Springs, Mississippi.
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Table 5. Number of specimens captured and measured and number of otoliths (or other hard parts) and stomachs sampled, by species priority level.
Species black seabass bluefish butterfish scup silver hake (whiting) striped bass summer flounder weakfish winter flounder
Species American shad Atlantic croaker Atlantic menhaden Atlantic torpedo bluntnose stingray bullnose ray clearnose skate cownose ray Leucoraja spp. little skate monkfish roughtail stingray skate spp. smooth butterfly ray smooth dogfish southern stingray spiny butterfly ray spiny dogfish spot winter skate yellowtail flounder
Species alewife Atlantic herring Atlantic mackerel black drum blueback herring red drum tautog
Priority A Species Total Total Number Species Number Caught Weight (kg) Measured 401 85.3 401 4,635 394.5 2,613 148,182 1,904.9 6,015 276,234 3,928.7 13,718 346 24.8 346 17 66.3 17 960 625.5 926 60,990 4,168.1 5,747 391 98.7 391 Priority B Species Total Total Number Species Number Caught Weight (kg) Measured 9 0.8 9 58,763 7,616.5 2,843 740 30.2 288 8 92.3 8 349 1,178.9 307 731 1,155.0 631 1,499 1,847.7 1,355 451 3,976.6 150 20 5.7 20 5,288 3,026.2 2,659 6 31.2 6 92 510.1 92 60 37.0 60 292 557.1 292 1,690 1,555.6 765 18 139.0 18 133 1,366.7 133 17 51.3 17 44,437 3,942.0 2,507 951 925.3 735 1 0.1 1 Priority C Species Total Total Number Species Number Caught Weight (kg) Measured 56 3.1 56 198 5.4 198 3 0.3 3 35 5.8 35 50 1.6 50 2 14.9 2 4 3.7 4
Number of Otoliths 219 588 538 808 59 16 716 572 118
Number of Stomachs 210 485 11 797 59 16 448 471 114
Number of Otoliths 9 211 78 7
Number of Stomachs 9 193 78 3
340
324
194 6 1
187 6 1
202
200
13 160 171 1
12 9 160 1
Number of Otoliths 24 20 3 33 18 1 4
Number of Stomachs 24 20 1 25 18 1 4
continued
- 16 -
Table 5. cont..
Species African pompano Atlantic bumper Atlantic cutlassfish Atlantic moonfish Atlantic pomfret Atlantic spadefish Atlantic sturgeon Atlantic thread herring Atlantic threadfin banded drum bay anchovy Berycidae bigeye bigeye scad blackcheek tonguefish blue runner bluespotted cornetfish cobia conger eel crevalle jack dwarf goatfish Etropus sp. Florida pompano flying gurnard fourbeard rockling fourspot flounder fringed flounder gray triggerfish Gulf Stream flounder harvestfish hickory shad hogchoker inshore lizardfish jellyfish spp king mackerel kingfish spp lane snapper lined seahorse longspine snipefish lookdown
Priority D Species Total Total Number Species Number Caught Weight(kg) Measured 2 0.2 2 2 0.0 2 212 2.1 117 2,803 15.1 1,196 2 0.2 2 673 31.0 478 2 29.4 2 3,345 167.7 554 3 0.3 3 655 50.7 210 119,741 203.4 3,961 5 13.7 5 11 1.7 11 75 2.4 75 3 0.2 3 175 9.9 160 1 0.0 1 4 33.1 4 1 0.7 1 31 1.1 31 13 0.2 8 24 0.3 24 43 7.6 43 1 0.0 1 1 0.1 1 73 15.7 73 27 0.4 27 23 9.2 23 91 1.9 91 102 10.0 57 7 2.4 7 130 14.3 129 514 56.9 514 818.3 5 22.1 5 9,124 1,398.8 1,707 4 0.1 4 2 0.0 2 1 0.2 1 5 0.2 5
Number of Otoliths
Number of Stomachs
continued
- 17 -
Table 5. cont. Priority D Species (continued)
Species mackerel scad mantis shrimp northern pipefish northern puffer northern searobin northern sennet northern stargazer orange filefish pigfish pinfish planehead filefish red goatfish red hake rock crab rough scad round herring round scad sea raven sharksucker sheepshead short bigeye silver anchovy silver perch smallmouth flounder Spanish mackerel Spanish sardine species unidentified spotfin mojarra spotted hake striped anchovy striped burrfish striped searobin threadfin shad triggerfish spp. windowpane
Total Number Caught 38 14 1 93 881 222 12 1 287 2,744 1 1 74 117 140 452 258 4 3 6 3 42 1,398 32 161 276 40 87 207 224,369 25 760 2 1 744
Total Species Weight (kg) 0.4 0.4 0.0 5.8 104.2 19.7 21.1 0.1 18.7 107.3 0.0 0.0 8.4 12.6 2.9 9.1 1.9 3.8 0.2 21.5 0.1 0.5 39.0 0.5 42.5 5.2 22.8 1.3 25.7 2,519.3 11.2 171.5 0.2 0.1 114.0
Number Measured 38 14 1 93 782 76 12 1 269 331 1 1 74 117 70 447 243 4 3 6 3 42 141 32 161 114 40 82 207 4,990 23 546 2 1 694
Number of Otoliths
Number of Stomachs
continued
- 18 -
Table 5. cont. Priority E Species
Species American lobster Atlantic angel shark Atlantic sharpnose shark blue crab - juvenile female blue crab - male blue crab, adult female brown shrimp dusky shark horseshoe crab jonah crab Loligo squid sandbar shark sea scallop squid spp thresher shark white shrimp Total
Total Number Caught 262 3
Total Species Weight (kg) 59.0 12.8
Number Measured 262 3
68
257.4
68
1 4 12 898 4 795 2 119,501 15 32 11 5 48 1,101,152
0.0 0.1 1.6 21.6 18.1 1,447.9 0.2 2,277.5 100.1 4.3 1.1 73.6 1.8 49,868
1 4 12 459 4 342 2 9,614 15 32 11 5 20 73,473
Number of Otoliths
Number of Stomachs
10
8
1
1
9
9
5,150
3,905
Figure 1. Chronological summary of average net performance parameters for each tow. Accepted ranges for each parameter are given by the dotted lines. 40
7
D. Spread - 32.6m
35
6
5
25 4
Speed – 3.2kts 20
3 15 2
W. Spread – 13.1m
10
1
5
Height – 5.3m
0
0
01
10 10
19 20
2830 3740 46 50 55 60 64 7073
80 82
90 91
Tow Number Station Number - 19 -
100 100 109 110 118 120 127 130 136140145 150
Speed (kts)
Distance (m)
30
Figure 2. Thyboron Type IV 66â&#x20AC;? trawl doors frontside (A) and backside (B) with mounting bracket for Netmind sensor. A.
B.
- 20 -
Figure 3. Comparison of stations selected (red) to locations sampled (black) in Rhode Island to New York (A), New Jersey to Maryland (B), and Virginia to North Carolina (C).
A
2
1 BI
RI
3 5
4
B
C
11
6 12
7
13
14
8
9 10
15
- 21 -
0 50 1, 0 00 1, 0 50 2, 0 00 2, 0 50 3, 0 00 3, 0 50 4, 0 00 4, 0 50 5, 0 00 5, 0 50 6, 0 00 6, 0 50 7, 0 00 7, 0 50 8, 0 00 8, 0 50 9, 0 00 9, 0 50 10 0 ,0 10 00 ,5 11 00 ,0 11 00 ,5 12 00 ,0 12 00 ,5 13 00 ,0 13 00 ,5 14 00 ,0 14 00 ,5 15 00 ,0 15 00 ,5 16 00 ,0 16 00 ,5 17 00 ,0 17 00 ,5 18 00 ,0 18 00 ,5 19 00 ,0 19 00 ,5 20 00 ,0 25 00 ,0 26 00 ,0 27 00 ,0 28 00 ,0 29 00 ,0 30 00 ,0 31 00 ,0 32 00 ,0 33 00 ,0 34 00 ,0 35 00 ,0 50 00 ,0 68 00 15 , 00 0, 0 00 0
Number of Tows
- 22 -
Number of Fish per Tow 40
38
36
34
32
30
28
26
24
22
20
18
16
14
12
10
8
6
4
2
0
Number of Tows
Figure 4. Frequency histogram of number of species captured at each station. 20
10
0 Number of Species per Tow
Figure 5. Frequency histogram of number of specimens captured at each station (note irregularly incremented values at the high end of the x-axis).
18 17 16 15 14 13 12 11 10 9 8 7 6 5 4 3 2 1 0
Figure 6. Frequency histogram of biomass of all specimens captured at each station (note irregularly incremented values at the high end of the x-axis).
20
10
10 0 15 0 20 0 25 0 30 0 35 0 40 0 45 0 50 0 55 0 60 0 65 0 70 0 75 0 80 0 85 0 90 0 95 0 1, 00 0 1, 05 0 1, 10 0 1, 15 0 1, 20 0 1, 25 0 1, 30 0 3, 60 0 6, 50 0
0
0 50
Number of Tows
30
Total Weight per Tow (kg)
- 23 -
American Lobster (Priority E)
A
B
Figure 7. Number (A) and biomass (B) of specimens captured at each station for American lobster.
- 24 -
Figure 8. Minimum trawlable number and biomass by state for American lobster. 1,200,000
300,000
1,100,000
Minimum Trawlable Number = 1,164,274
1,000,000
Minimum Trawlable Biomass (kg)= 262,773 Pct. S.E. = 50% (number) 46% (biomass)
900,000 800,000
200,000
Biomass (kg)
Number
700,000 600,000 500,000
100,000
400,000 300,000 200,000 100,000
Avg Wt: 0.224kg
Avg Wt: 0.344kg
Avg Wt: 0.284kg
0
0
RI
NY
NJ
DE
MD
VA
NC
State Figure 9. Length frequency histogram for American lobster. 80 70
Number Captured = 262 Number Measured = 262
Expanded Number
60 50 40 30 20 10 0 0
1
2
3
4
5
6
7
Carapace Length (cm) - 25 -
8
9
10
11
12
Figure 10. Sex ratios for American lobster, by state.
n= 100
126
2
3
100
100
100
51
100
33
90 80 70 Percent
67 60 50
49 40 30 20
Sex U M F
10 0 1-RI
2-NY STATE
- 26 -
3-NJ
Figure 11. Length-weight regression for American lobster, sexes combined (A) and by sex (B). 700
600
Weight(g) = 1.8115 x Length(cm)2.4836 (n = 131)
Weight(g)
500
400
300
200
A
100
0 3
4
5
6
7
8
9
10
11
Carapace Length(cm)
700
Fe m ale s: We ight(g) = 1.923 x Le ngth(cm ) 2.4553 (n = 65)
600
Male s: (n = 65)
We ight(g) = 1.6928 x Le ngth(cm ) 2.5167
Weight(g)
500
400
300
200
100
B
0
3
4
5
6
7
8
Carapace Len gth(cm) - 27 -
9
10
11
Atlantic Croaker (Priority B)
A
B
Figure 12. Number (A) and biomass (B) of specimens captured at each station for Atlantic croaker.
- 28 -
Figure 13. Minimum trawlable number and biomass by state for Atlantic croaker. 150,000,000
18,000,000
140,000,000
17,000,000
130,000,000 120,000,000
16,000,000
Minimum Trawlable Biomass (kg)= 30,231,523
15,000,000 14,000,000
Pct. S.E. = 42% (number) 40% (biomass)
110,000,000
13,000,000
100,000,000
12,000,000
90,000,000
11,000,000 10,000,000
80,000,000
9,000,000 70,000,000
8,000,000
60,000,000
7,000,000
50,000,000
6,000,000 Avg Wt: 0.281kg
40,000,000
5,000,000 Avg Wt: 0.194kg
30,000,000
Avg Wt: 0.122kg
4,000,000
Avg Wt: 0.086kg
3,000,000 20,000,000 10,000,000
2,000,000
Avg Wt: 0.305kg
Avg Wt: 0.794kg
1,000,000 0
0
RI
NY
NJ
DE
MD
VA
NC
State Figure 14. Length frequency histogram for Atlantic croaker. 9,000
Number Captured = 58,763
8,000
Number Measured = 2,843
Expanded Number
7,000
Number of Otoliths = 211 6,000
Number of Stomachs = 193
5,000 4,000 3,000 2,000 1,000 0 0
5
10
15
20
25
Total Length (cm) - 29 -
30
35
40
45
Biomass (kg)
Number
Minimum Trawlable Number = 231,508,457
Figure 15. Sex ratios for Atlantic croaker by state (A) and length group (B).
A
n= 100
3
54
25
31
71
27
100.00
100.00
99.99
100.00
100.00
100.00
100
59
74
57
42
38
90 80
Percent
70 60
62 58
Sex
50 40
U M F
42
41
30
25
20 10 0 2-NY
3-NJ
4-DE
100
6-VA
7-NC
STATE
B Percent SUM n=
5-MD
7 100
14 100
32 100
43 100
40 100
30 100
18 100
15 100
8 100
3 100
1 100
74
14
43
52
39
74
46
18
15
100
100
90
86
80
82
85
70 60
60
57 50
Sex
53
48
U M F
40 30 20
26
26
10 5
6
7
8
9
10
11
12
13
14
15 Inch-class
Length (cm)
- 30 -
.5 40 .0 -4 2
.0 37 .5 -4 0
.5 35 .0 -3 7
.0 32 .5 -3 5
.5 30 .0 -3 2
.0 27 .5 -3 0
.5 25 .0 -2 7
.0 22 .5 -2 5
.5 20 .0 -2 2
.0 17 .5 -2 0
15 .0 -1 7
.5
0
Figure 16. Maturity logistic regression for Atlantic croaker, by sex. 1.0 99% maturity
0.9
Probability of Maturity
0.8 0.7 0.6 0.5
50% maturity
0.4 0.3 0.2 0.1
SEX
F M
0.0 10
20
30
Total Length (cm)
- 31 -
40
50
Figure 17. Length-weight regression for Atlantic croaker, sexes combined (A) and by sex (B). 1100
A
1000 900 800
Weight(g) = 0.0104 x Length(cm) 3.0628 (n = 209)
Weight(g)
700 600 500 400 300 200 100 0
10
20
30
40
50
Total Length(cm) 1100 1000
Females: W eight(g) = 0.0103 x Length(cm) 3.0676 (n = 132)
900
Males: W eight(g) = 0.0123 x Length(cm) 3.0124 (n = 75)
800
B
Weight(g)
700 600 500 400 300 200 100 0 10
20
30
T otal Len gth (cm)
- 32 -
40
50
Atlantic Menhaden (Priority B)
A
B
Figure 18. Number (A) and biomass (B) of specimens captured at each station for Atlantic menhaden.
- 33 -
Figure 19. Minimum trawlable number and biomass by state for Atlantic menhaden. 60,000
1,600,000 1,500,000 1,400,000
Minimum Trawlable Number = 2,255,876
1,300,000
Minimum Trawlable Biomass (kg)= 109,297
50,000
1,200,000
Pct. S.E. = 70% (number) 32% (biomass)
1,100,000
40,000
1,000,000
Number
30,000
800,000 700,000 600,000
20,000
500,000 400,000 300,000
10,000
Avg Wt: 0.012kg
200,000 Avg Wt: 0.018kg
100,000
Avg Wt: 0.202kg
Avg Wt: 0.269kg
Avg Wt: 0.330kg
Avg Wt: 0.187kg
0
0
RI
NY
NJ
DE
MD
VA
NC
State
Expanded Number
Figure 20. Length frequency histogram for Atlantic menhaden. 200 190 180 170 160 150 140 130 120 110 100 90 80 70 60 50 40 30 20 10 0
Number Captured = 740 Number Measured = 288 Number of Otoliths = 78 Number of Stomachs = 78
0
5
10
15
20
Fork Length (cm) - 34 -
25
30
35
Biomass (kg)
900,000
Figure 21. Sex ratios for Atlantic menhaden by state (A) and length group (B).
A
n= 100
100
7
12 100
100
33
100
8
100
7
100
6
20
94
26
13
100
67
90 38
80
60
53
Percent
70 60
Sex
50
50
U M F
40 33
30 20
21
20
10 6
0 1-RI
2-NY
3-NJ
4-DE
B Percent SUM n= 100
5-MD
6-VA
STATE
12 100
2 100
54
1 100
2 100
9 100
18 100
50
100
100
81
96
90
9
6
14
100
100
100
7
48
17
62
80
83
70 60 50 40
Sex
53
50
U M F
46
30
32
20
19 10 3
4
5
7
8
9
10
11
12
Length (cm)
- 35 -
.5 30 .0 -3 2
.0 27 .5 -3 0
.5 25 .0 -2 7
22 .5 -2 5
.0
.5 20 .0 -2 2
.0 17 .5 -2 0
.0 12 .5 -1 5
.5 10 .0 -1 2
07 .5 -1 0
.0
0
Inch-class
Figure 22. Maturity logistic regression for Atlantic menhaden, by sex. 1.0
99% maturity 0.9
Probability of Maturity
0.8 0.7 0.6 0.5
50% maturity 0.4 0.3 0.2 0.1
SEX
F M
0.0 0
10
20
Fork Length (cm)
- 36 -
30
40
Figure 23. Length-weight regression for Atlantic menhaden, sexes combined (A) and by sex (B). 600
500
Weight(g) = 0.0331 x Length(cm)2.7901 (n = 78)
Weight(g)
400
300
200
100
A
0 0
10
20
30
40
Fork Length(cm) 600
Fe m ale s: We ight(g) = 0.0358 x Le ngth(cm ) 2.7632 (n = 19) Male s: (n = 35)
500
We ight(g) = 0.0319 x Le ngth(cm ) 2.8059
Weight(g)
400
300
200
100
B
0
0
10
20
Fork Length(cm)
- 37 -
30
40
Atlantic Spadefish (Priority D)
A
B
Figure 24. Number (A) and biomass (B) of specimens captured at each station for Atlantic spadefish.
- 38 -
Figure 25. Minimum trawlable number and biomass by state for Atlantic spadefish. 90,000
1,800,000 1,700,000 1,600,000
Minimum Trawlable Number = 2,595,794
1,500,000
Minimum Trawlable Biomass (kg)= 102,415
1,400,000
80,000
70,000
Pct. S.E. = 35% (number) 32% (biomass)
1,300,000
60,000
1,200,000
50,000
Number
1,000,000 900,000
40,000
800,000 700,000
30,000
600,000 500,000
20,000
400,000 300,000 Avg Wt: 0.045kg
200,000
Avg Wt: 0..047kg
10,000
100,000
0
0
RI
NY
NJ
DE
MD
VA
NC
State Figure 26. Length frequency histogram for Atlantic spadefish. 300
Number Captured = 673
Expanded Number
Number Measured = 478 200
100
0 0
5
10
15
20
Fork Length (cm) - 39 -
25
30
35
Biomass (kg)
1,100,000
Atlantic Thread Herring (Priority D)
A
B
Figure 27. Number (A) and biomass (B) of specimens captured at each station for Atlantic thread herring.
- 40 -
Figure 28. Minimum trawlable number and biomass by state for Atlantic thread herring. 600,000
7,000,000
Minimum Trawlable Number = 13,526,564 6,000,000
Minimum Trawlable Biomass (kg)= 710,948
500,000
Pct. S.E. = 52% (number) 65% (biomass) 5,000,000
400,000
Number
300,000 3,000,000
200,000 2,000,000
100,000 1,000,000 Avg Wt: 0.005kg
Avg Wt: 0.141kg
Avg Wt: 0.080kg
Avg Wt: 0.027kg
0
0
RI
NY
NJ
DE
MD
VA
NC
State Figure 29. Length frequency histogram for Atlantic thread herring. 900
Number Captured = 3,345
800
Number Measured = 554
Expanded Number
700 600 500 400 300 200 100 0 0
5
10
15
Fork Length (cm) - 41 -
20
25
Biomass (kg)
4,000,000
Bay Anchovy (Priority D)
A
B
Figure 30. Number (A) and biomass (B) of specimens captured at each station for bay anchovy.
- 42 -
Figure 31. Minimum trawlable number and biomass by state for bay anchovy. 400,000
180,000,000 170,000,000 160,000,000
Minimum Trawlable Number = 452,812,195
150,000,000
Minimum Trawlable Biomass (kg)= 799,420
140,000,000
Pct. S.E. = 28% (number) 15% (biomass)
300,000
130,000,000 120,000,000
Number
100,000,000
200,000
90,000,000 80,000,000 70,000,000 60,000,000 50,000,000
100,000 40,000,000 30,000,000 20,000,000 Avg Wt: 0.005kg
10,000,000
Avg Wt: 0.001kg
Avg Wt: 0.002kg
Avg Wt: 0.002kg
Avg Wt: 0.002kg
Avg Wt: 0.003kg
MD
VA
0
0
RI
NY
NJ
DE
NC
State Figure 32. Length frequency histogram for bay anchovy. 40,000
Number Captured = 119,741 Number Measured = 3,961
Expanded Number
30,000
20,000
10,000
0 0
5
10
Fork Length (cm) - 43 -
15
Biomass (kg)
110,000,000
Black Sea Bass (Priority A)
A
B
Figure 33. Number (A) and biomass (B) of specimens captured at each station for black sea bass.
- 44 -
Figure 34. Minimum trawlable number and biomass by state for black sea bass. 110,000
600,000
100,000
Minimum Trawlable Number = 1,568,759 500,000
Minimum Trawlable Biomass (kg)= 326,991
90,000
Pct. S.E. = 22% (number) 23% (biomass)
80,000
400,000
60,000 300,000
50,000
40,000 200,000
30,000
20,000
100,000
Avg Wt: 0.741kg
Avg Wt: 0.253kg
Avg Wt: 0.198kg
Avg Wt: 0.102kg
Avg Wt: 0.074kg
Avg Wt: 0.111kg
Avg Wt: 0. 069kg
10,000
0
0
RI
NY
NJ
DE
MD
VA
NC
State Figure 35. Length frequency histogram for black sea bass. 40
Number Captured =401 Number Measured = 401
Expanded Number
30
Number of Otoliths = 219 Number of Stomachs = 210
20
10
0 0
5
10
15
20
25
30
35
Total Length (cm) - 45 -
40
45
50
55
60
Biomass (kg)
Number
70,000
Figure 36. Sex ratios for black sea bass, by state (A) and length group (B).
A
n= 100
24
45
100.00 7
90
69
22
18
34
7
100.00
100.00
100.00
100.00
100.00
100.01
31
5 8
100
22
16
43
14
87
80
82 79
74
70 Percent
67
60 14
50 40
Sex U M F
43
30 20 10 0 1-RI
2-NY
3-NJ
4-DE
100
6-VA
7-NC
STATE
B Percent SUM n=
5-MD
2 100.0
13 100.0
100
6
94
90
97 100.0
32 100.0
29 100.0
10
11
18
86
86
24 100.1
11 100.0
5 100.0
2 100.0
3 100.0
1 100.0
10
26
40
100
33
100
38
80
78 74
70
67
60
60
Sex
50
52
U M F
40 30 20 10 2-4
4-6
6-8
8-10
10-12
12-14
14-16
16-18
18-20
20-22
22-24 Inch-class
Length (cm)
- 46 -
55 -6 0
50 -5 5
45 -5 0
40 -4 5
35 -4 0
30 -3 5
25 -3 0
20 -2 5
15 -2 0
10 -1 5
05 -1 0
0
Figure 37. Maturity logistic regression for black sea bass, by sex. 1.0 99% maturity
0.9
0.7 0.6 0.5
50% maturity
0.4 0.3 0.2 0.1
SEX
F M
0.0 10
20
30
40
50
60
Total Length (cm) Figure 38. Expanded age frequency histogram for black sea bass. 169
170 160 150 140
Catch curve Z ages 1-9 = -0.68 Catch curve Z ages 2-9 = -0.63
130 120 110 100
93
90 80 70
61
60
54
50 40 30 20
6
10
3
2
1
2
0
0
Age (Year Class)
- 47 -
(1 99 7) 10
(1 99 8) 9
(1 99 9) 8
(2 00 0) 7
(2 00 1) 6
5
(2 00 2)
(2 00 3) 4
(2 00 4) 3
(2 00 5) 2
(2 00 6) 1
(2 00 7)
0
0
Expanded Number
Probability of Maturity
0.8
Figure 39. Age-specific length-frequency histograms for black sea bass for ages 0 through 3, and 5.
AG E = 0 11 10 9 8 7 6 5 4 3 2 1 0 0
5
10
15
20
25
30
35
40
45
50
55
60
40
45
50
55
60
40
45
50
55
60
40
45
50
55
60
40
45
50
55
60
To ta l Le n g th (c m ) AG E = 1 14 13 12 11 10 9 8 7 6 5 4 3 2 1 0 0
5
10
15
20
25
30
35
To ta l Le n g th (c m ) AG E = 2 5
4
3
2
1
0 0
5
10
15
20
25
30
35
T o t a l Le n g t h ( c m ) AG E = 3 5
4
3
2
1
0 0
5
10
15
20
25
30
35
T o t a l Le n g t h ( c m ) AG E = 5 2
1
0 0
5
10
15
20
25
30
35
T o t a l Le n g t h ( c m )
- 48 -
Figure 40. von Bertalanffy growth curves for black sea bass, by sex. 60
Fe m ale s : L(m ax) = 70.5 T(ze ro) = 0.263 K = 0.232
50
Length (cm)
M ale s :
L(m ax) = 46.2 T(ze ro) = 0.401 K = 0.453
40
30
20
10 0
1
2
3
4
5
Age
- 49 -
6
7
8
9
10
Figure 41. Length-weight regression for black sea bass, sexes combined (A) and by sex (B).. 3000
Weight(g) = 0.0286 x Length(cm) 2.7973 (n = 219)
Weight(g)
2000
1000
A 0 0
10
20
30
40
50
60
Total Length(cm)
3000 Females: W eight(g) = 0.0251 x Length(cm) 2.8433 (n = 177) Males: W eight(g) = 0.1191 x Length(cm) 2.3719 (n = 26)
Weight(g)
2000
1000
B 0 10
20
30
40
T otal Len gth (cm)
- 50 -
50
60
Figure 42. Diets of black sea bass by percent weight (A) and percent number (B).
A
B
- 51 -
Bluefish (Priority A)
A
B
Figure 43. Number (A) and biomass (B) of specimens captured at each station for bluefish.
- 52 -
Figure 44. Minimum trawlable number and biomass by state for bluefish. 500,000
8,000,000
Minimum Trawlable Number = 16,765,929 7,000,000
Minimum Trawlable Biomass (kg)= 1,519,012 400,000
Pct. S.E. = 17% (number) 13% (biomass)
6,000,000
5,000,000
Number
4,000,000
200,000 3,000,000
2,000,000 Avg Wt: 0.297kg Avg Wt: 0.453kg
1,000,000
Avg Wt: 0.028kg
Avg Wt: 0.103kg
NY
NJ
Avg Wt: 0.134kg
Avg Wt: 0.100kg
100,000
Avg Wt: 0. 149kg
0
0
RI
DE
MD
VA
NC
State Figure 45. Length frequency histogram for bluefish. 600
Number Captured = 4,635
500
Expanded Number
Number Measured = 2,613 Number of Otoliths = 588
400
Number of Stomachs = 485 300
200
100
0 0
5
10
15
20
25
30
35
40
45
Fork Length (cm) - 53 -
50
55
60
65
70
75
Biomass (kg)
300,000
Figure 46. Sex ratios for bluefish, by state (A) and length group (B).
A
n= 100 90
64
99
164
43
70
104
100
100
100
100
100
100
100
37
9.2
84.7
22.3
46.5
5.1 57.0
47.1
17.0
66.2
54.3
80 45.7
Percent
70 60 53.5
50
Sex
21.3
U M F
40 37.9
30
32.0
31.6
28.7
24.6
20
13.9
10 0 1-RI
2-NY
3-NJ
4-DE
5-MD
6-VA
7-NC
STATE
B Percent SUM n= 100
18 100
78 100
123 194 56 100 100 100
99
78
45
14
58
67 100
13 5 100 100
36
12
40
2 100
7 4 11 2 1 100 100 100 100 100
50
86
50
27
100
100
90 88
47
80 70
73 64
60
60 Sex
35
50
50
50
40 39
U M F
42
30 20
21
20
4-6
6-8
10
2-4
14 8-10 10-12 12-14 14-16 16-18 18-20 20-22 22-24 24-26 26-28 28-30 Inch-class
Length (cm)
- 54 -
70 -7 5
65 -7 0
60 -6 5
55 -6 0
50 -5 5
45 -5 0
40 -4 5
35 -4 0
30 -3 5
25 -3 0
20 -2 5
15 -2 0
10 -1 5
05 -1 0
0
Figure 47. Maturity logistic regression for bluefish, by sex. 1.0
99% maturity 0.9
Probability of Maturity
0.8 0.7 0.6 0.5
50% maturity 0.4 0.3 0.2 0.1
SEX
F M
0.0 0
10
20
30
40
50
Fork Length (cm)
- 55 -
60
70
80
Figure 48. Length-weight regression for bluefish, sexes combined (A) and by sex (B).. 5000
Weight(g)
4000
Weight(g) = 0.0211 x Length(cm)2.8645 (n = 588)
3000
2000
1000
A
0 0
10
20
30
40
50
60
70
80
Fork Length(cm)
5000
Fe m ale s: We ight(g) = 0.0218 x Le ngth(cm ) 2.8541 (n = 236) Male s: We ight(g) = 0.0179 x Le ngth(cm ) 2.9133 (n = 229)
Weight(g)
4000
3000
2000
1000
B 0
0
10
20
30
40
50
Fork Length(cm) - 56 -
60
70
80
Figure 49. Diets of bluefish by percent weight (A) and percent number (B).
A
B
- 57 -
Bluntnose Stingray (Priority B)
A
B
Figure 50. Number (A) and biomass (B) of specimens captured at each station for bluntnose stingray.
- 58 -
Figure 51. Minimum trawlable number and biomass by state for bluntnose stingray. 4,000,000
1,000,000
Minimum Trawlable Number = 1,342,462
900,000
Minimum Trawlable Biomass (kg)= 4,613,305 800,000
Pct. S.E. = 21% (number) 23% (biomass)
3,000,000
700,000
Number
2,000,000
500,000
400,000
300,000
1,000,000 200,000
100,000
Avg Wt: 0.492kg
Avg Wt: 0.772kg
Avg Wt: 2.493kg
Avg Wt: 3.874kg
Avg Wt: 2.542kg
0
0
RI
NY
NJ
DE
MD
VA
NC
State Figure 52. Width frequency histogram for bluntnose stingray. 30
Number Captured = 349
Expanded Number
Number Measured = 307
20
10
0 0
5
10
15
20
25
30
35
40
45
50
Disk Width (cm)
- 59 -
55
60
65
70
75
80
85
Biomass (kg)
600,000
Brown Shrimp (Priority E)
A
B
Figure 53. Number (A) and biomass (B) of specimens captured at each station for brown shrimp.
- 60 -
Figure 54. Minimum trawlable number and biomass by state for brown shrimp. 60,000
3,000,000
Minimum Trawlable Number = 3,364,019 Minimum Trawlable Biomass (kg)= 81,052
50,000
Pct. S.E. = 20% (number) 16% (biomass)
40,000
30,000
20,000
1,000,000
Avg Wt: 0.026kg Avg Wt: 0.021kg
Avg Wt: 0.024kg
10,000
Avg Wt: 0.023kg
0
0
RI
NY
NJ
DE
MD
VA
NC
State Figure 55. Length frequency histogram for brown shrimp. 300
Number Captured = 898
Expanded Number
Number Measured = 459 200
100
0 0
5
10
Total Length (cm) - 61 -
15
20
Biomass (kg)
Number
2,000,000
Bullnose Stingray (Priority B)
A
B
Figure 56. Number (A) and biomass (B) of specimens captured at each station for bullnose stingray.
- 62 -
Figure 57. Minimum trawlable number and biomass by state for bullnose stingray. 3,000,000
1,600,000 1,500,000
Minimum Trawlable Number = 2,989,245
1,400,000
Minimum Trawlable Biomass (kg)= 4,747,851
1,300,000
Pct. S.E. = 16% (number) 17% (biomass)
1,200,000 1,100,000
2,000,000
1,000,000
Biomass (kg)
Number
900,000 800,000 700,000 600,000
1,000,000
500,000 400,000
Avg Wt: 1.127kg
300,000 Avg Wt: 1.209kg
200,000
Avg Wt: 1.360kg
Avg Wt: 1.236kg
Avg Wt: 1.938kg
100,000
0
0
RI
NY
NJ
DE
MD
VA
NC
State Figure 58. Width frequency histogram for bullnose stingray. 80
Number Captured = 731 70
Number Measured = 631
Expanded Number
60
50
40
30
20
10
0 0
5
10
15
20
25
30
35
40
45
Disk Width (cm)
- 63 -
50
55
60
65
70
75
80
Butterfish (Priority A)
A
B
Figure 59. Number (A) and biomass (B) of specimens captured at each station for butterfish.
- 64 -
Figure 60. Minimum trawlable number and biomass by state for butterfish. 5,000,000
300,000,000
Minimum Trawlable Number = 556,339,149 Minimum Trawlable Biomass (kg)= 7,433,615 Pct. S.E. = 30% (number) 18% (biomass)
4,000,000
200,000,000
Number
2,000,000 100,000,000 Avg Wt: 0.018kg
Avg Wt: 0.004kg Avg Wt: 0.037kg Avg Wt: 0.012kg
Avg Wt: 0.018kg
Avg Wt: 0.023kg
Avg Wt: 0.043kg
1,000,000
0
0
RI
NY
NJ
DE
MD
VA
NC
State Figure 61. Length frequency histogram for butterfish. 40,000
Number Captured = 148,182 Number Measured = 6,015
Expanded Number
30,000
Number of Otoliths = 538 Number of Stomachs = 11
20,000
10,000
0 0
5
10
Fork Length (cm) - 65 -
15
20
Biomass (kg)
3,000,000
Figure 62. Sex ratios for butterfish, by state (A) and length group (B).
A n = 100.0 120
100
79.3
90
107
30
54
104
24
100.0
100.0
100.0
100.0
100.0
100.1
97.8
81.8
99.6
77.7
52.3
9.1
90
30.5
80
Percent
70 60
60.5 Sex
50
U M F
10.4
40 37.3
30 20
14.0
5.3 17.0
6.7
10
11.5 6.7
0 1-RI
2-NY
3-NJ
100
5-MD
6-VA
7-NC
66 100.1
20 100.0
STATE
B Percent SUM n=
4-DE
33 100.0
135 100.0
121 100.0
75 100.0
79 100.0
100
100
99
57
11
42
18
90
50 80
82
70 60
55
50 40
26
Sex U M F
39
30 20
17 10 1
2
3
4
5
6
7
Length (cm)
- 66 -
.0 17 .5 -2 0
.5 15 .0 -1 7
.0 12 .5 -1 5
.5 10 .0 -1 2
.0 07 .5 -1 0
.5 05 .0 -0 7
02 .5 -0 5
.0
0
Inch-class
Figure 63. Maturity logistic regression for butterfish, by sex. 1.0 99% maturity
0.9
Probability of Maturity
0.8 0.7 0.6 0.5
50% maturity
0.4 0.3 0.2 0.1
SEX
F M
0.0 7
8
9
10
11
12
13
14
Fork Length (cm)
- 67 -
15
16
17
18
19
Figure 64. Length-weight regression for butterfish, sexes combined (A) and by sex (B). 180 170 160 150 140 130
Weight(g) = 0.015 x Length(cm) 3.1259 (n = 535)
Weight(g)
120 110 100 90 80 70 60 50 40 30
A
20 10 0 3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
Fork Length(cm)
180 170 Females: W eight(g) = 0.0269 x Length(cm) 2.9149 (n = 106)
160 150
Males: W eight(g) = 0.0113 x Length(cm) 3.2363 (n = 96)
140 130
Weight(g)
120 110 100 90 80 70 60 50 40 30
B
20 10 0 7
8
9
10
11
12
13
14
Fork Len gth (cm)
- 68 -
15
16
17
18
19
Clearnose Skate (Priority B)
A
B
Figure 65. Number (A) and biomass (B) of specimens captured at each station for clearnose skate.
- 69 -
Figure 66. Minimum trawlable number and biomass by state for clearnose skate. 3,000,000
2,100,000 2,000,000
Minimum Trawlable Number = 5,728,724
1,900,000 1,800,000
Minimum Trawlable Biomass (kg)= 6,975,177
1,700,000
Pct. S.E. = 8% (number) 8% (biomass)
1,600,000 1,500,000
2,000,000
1,400,000
Number
Biomass (kg)
1,300,000 1,200,000 1,100,000 1,000,000 900,000 800,000
1,000,000
700,000 600,000 500,000 Avg Wt: 1.177kg
400,000 Avg Wt: 1.542kg
300,000
Avg Wt: 1.414kg
Avg Wt: 1.052kg
Avg Wt: 1.084kg
Avg Wt: 1.295kg
Avg Wt: 1.178kg
200,000
0
100,000
RI
NY
NJ
DE
MD
VA
NC
State Figure 67. Width frequency histogram for clearnose skate. 130
Expanded Number
120 110
Number Captured = 1,499
100
Number Measured = 1,355
90
Number of Vertebrae = 340
80
Number of Stomachs = 324 70 60 50 40 30 20 10 0 0
5
10
15
20
25
30
Disk Width (cm)
- 70 -
35
40
45
50
55
Figure 68. Sex ratios for clearnose skate, by state (A) and width group (B).
A
n= 100
9
58
39
22
121
100
100
100
63
100
100
100
100
28
31.3
88.2
74.6
61.0
75.0
27.0
33.3
90 80 73.0
70 Percent
68.7
66.7
60 Sex
50
U M F
40 39.0
30 25.4
20 10
25.0
11.8
0 1-RI
2-NY
3-NJ
4-DE
100
6-VA
7-NC
STATE
B Percent SUM n=
5-MD
4 100
2 100
18 100
46 100
45 100
72 100
69 100
48 100
28 100
7 100
1 100
28
100
55
52
75
61
59
27
38
40
100
62
60
90 80 70
73
72
60
Sex
50
45
40
48 39
U M F
41
30
25
20 10
10
11
12
13
14
15
16
17
18
19
21
DiskLength Width(cm) (cm) - 71 -
.0 52 .5 -5 5
.0 47 .5 -5 0
.5 45 .0 -4 7
.0 42 .5 -4 5
.5 40 .0 -4 2
.0 37 .5 -4 0
.5 35 .0 -3 7
.0 32 .5 -3 5
.5 30 .0 -3 2
.0 27 .5 -3 0
25 .0 -2 7
.5
0
Inch-class
Figure 69. Maturity logistic regression for clearnose skate, by sex. 1.0
99% maturity 0.9
Probability of Maturity
0.8 0.7 0.6 0.5
50% maturity
0.4 0.3 0.2 0.1
SEX
F M
0.0 20
30
40
Disk Width (cm)
- 72 -
50
60
Figure 70. Width-weight regression for clearnose skate, sexes combined (A) and by sex (B). 3000
Weight(g) = 0.0283 x Length(cm) 2.9061 (n = 339)
Weight(g)
2000
1000
A 0 20
30
40
50
60
Disk Width(cm)
3000 Females: W eight(g) = 0.0265 x Length(cm) 2.9201 (n = 166) Males: W eight(g) = 0.0231 x Length(cm) 2.9641 (n = 173)
Weight(g)
2000
1000
B 0 20
30
40
Disk Width (cm) - 73 -
50
60
Cownose Ray (Priority B)
A
B
Figure 71. Number (A) and biomass (B) of specimens captured at each station for cownose ray.
- 74 -
Figure 72. Minimum trawlable number and biomass by state for cownose ray. 1,900,000
19,000,000
1,800,000
18,000,000
1,700,000
17,000,000
Minimum Trawlable Number = 2,015,335
1,600,000
16,000,000
Minimum Trawlable Biomass (kg)= 18,366,308
15,000,000
1,500,000
Pct. S.E. = 63% (number) 88% (biomass)
14,000,000
1,300,000
13,000,000
1,200,000
12,000,000
1,100,000
11,000,000
1,000,000
10,000,000
900,000
9,000,000
800,000
8,000,000
700,000
7,000,000
600,000
6,000,000
500,000
5,000,000
Biomass (kg)
Number
1,400,000
4,000,000
400,000 300,000
Avg Wt: 1.681kg
Avg Wt: 1.705kg
Avg Wt: 10.089kg
3,000,000
Avg Wt: 1.232kg
200,000
2,000,000
100,000
1,000,000 0
0
RI
NY
NJ
DE
MD
VA
NC
State Figure 73. Width frequency histogram for cownose ray. 30
Number Captured = 451
Expanded Number
Number Measured = 150
20
10
0 0
5
1 0
1 5
2 0
2 5
3 0
3 5
4 0
4 5
5 0
5 5
6 0
6 5
7 0
Disk Width (cm)
- 75 -
7 5
8 0
8 5
9 0
9 5
1 0 0
1 0 5
1 1 0
1 1 5
Horseshoe Crab (Priority E)
A
B
Figure 74. Number (A) and biomass (B) of specimens captured at each station for horseshoe crab.
- 76 -
Figure 75. Minimum trawlable number and biomass by state for horseshoe crab. 3,000,000
1,100,000
Minimum Trawlable Number = 3,394,754
1,000,000
Minimum Trawlable Biomass (kg)= 6,168,907
900,000
Pct. S.E. = 31% (number) 32% (biomass)
800,000
2,000,000
Biomass (kg)
Number
700,000
600,000
500,000
400,000
1,000,000 300,000
200,000
Avg Wt: 0.136kg
100,000
Avg Wt: 0.246kg
Avg Wt: 0.158kg
Avg Wt: 0.118kg
Avg Wt: 0.122kg
Avg Wt: 0.120kg
Avg Wt: 0.092kg
0
0
RI
NY
NJ
DE
MD
VA
NC
State Figure 76. Width frequency histogram for horseshoe crab. 120
Expanded Number
110 100
Number Captured = 795
90
Number Measured = 342
80 70 60 50 40 30 20 10 0 0
5
10
15
20
Carapace Width (cm) - 77 -
25
30
35
Figure 77. Sex ratios for horseshoe crab, by state (A) and width group (B).
A n= 100
100
5
100
52
100
36
18 100
32 100
21 100
40.0
36.9
21.2
38.2
41.8
35.0
90 80
78.8
Percent
70 60
63.1
60.0
65.0
61.8 58.2
Sex
50
U M F
40 30 20 10 0 1-RI
2-NY
3-NJ
4-DE
B n= Percent SUM
100
5-MD
6-VA
STATE
1 100
1 100
12 100
44 100
40 100
100
100
89
84
15
42 100
97
22 100
2 100
100
100
90
85
80 70 60
Sex
50
U M F
40 30 20
17
10 5
6
11 7
8
9
10
11
12
Carapace Width (cm) Length (cm) - 78 -
.5 30 .0 -3 2
.0 27 .5 -3 0
.5 25 .0 -2 7
.0 22 .5 -2 5
.5 20 .0 -2 2
.0 17 .5 -2 0
.5 15 .0 -1 7
12 .5 -1 5
.0
0
Inch-class
Figure 78. Width-weight regression for horseshoe crab, sexes combined (A), and by sex (B). 5000
4000
Weight(g)
Weight(g) = 0.0297 x Length(cm) 3.4455 (n = 166) 3000
2000
1000
A
0 13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
Carapace Width (cm)
4000 Females: W eight(g) = 0.1044 x Length(cm) 3.068 (n = 108) Males: W eight(g) = 0.2982 x Length(cm) 2.6636 (n = 56)
Weight(g)
3000
2000
1000
B 0 13
14
15
16
17
18
19
20
21
22
23
24
25
Carapace Width (cm) - 79 -
26
27
28
29
30
31
32
Kingfish spp. (Priority D)
A
B
Figure 79. Number (A) and biomass (B) of specimens captured at each station for kingfish (spp).
- 80 -
Figure 80. Minimum trawlable number and biomass by state for kingfish (spp). 5,000,000
40,000,000
Minimum Trawlable Number = 36,116,481 Minimum Trawlable Biomass (kg)= 5,611,106
4,000,000
Pct. S.E. = 33% (number) 35% (biomass)
30,000,000
Number
20,000,000
2,000,000
10,000,000
1,000,000 Avg Wt: 0.119kg
Avg Wt: 0.142kg
Avg Wt: 0.159kg
RI
NY
NJ
Avg Wt: 0.117kg
Avg Wt: 0. 205kg
Avg Wt: 0.158kg
Avg Wt: 0.120kg
VA
NC
0
0
DE
MD
State Figure 81. Length frequency histogram for kingfish (spp). 1,400
Expanded Number
1,300 1,200
Number Captured = 9,124
1,100
Number Measured = 1,707
1,000 900 800 700 600 500 400 300 200 100 0 0
5
10
15
20
25
Total Length (cm) - 81 -
30
35
40
Biomass (kg)
3,000,000
Little Skate (Priority B)
A
B
Figure 82. Number (A) and biomass (B) of specimens captured at each station for little skate.
- 82 -
Figure 83. Minimum trawlable number and biomass by state for little skate. 7,000,000
11,000,000
10,000,000
Minimum Trawlable Number = 18,483,896
6,000,000
Minimum Trawlable Biomass (kg)= 10,732,786
9,000,000
Pct. S.E. = 14% (number) 13% (biomass)
8,000,000
5,000,000
7,000,000
Number
5,000,000
3,000,000 4,000,000
2,000,000
3,000,000
2,000,000 Avg Wt: 0.592kg
Avg Wt: 0.549kg
Avg Wt: 0.566kg
RI
NY
NJ
Avg Wt: 0.530kg
1,000,000
Avg Wt: 1.254kg
1,000,000
0
0
DE
MD
VA
NC
State Figure 84. Width frequency histogram for little skate. 1,500 1,400 1,300
Number Captured = 5,288
1,200
Number Measured = 2,659
Expanded Number
1,100
Number of Vertebrae = 194
1,000
Number of Stomachs = 187
900 800 700 600 500 400 300 200 100 0 0
5
10
15
20
25
30
35
Disk Width (cm) - 83 -
40
45
50
55
60
Biomass (kg)
4,000,000 6,000,000
Figure 85. Sex ratios for little skate, by state (A) and width group (B).
A
n= 100
81
74
38
1
100
100
100
100
35
16
100 96
90
84
80 70
65
Percent
60
Sex
50
U M F
40 30 20 10 0 1-RI
2-NY
B
3-NJ
4-DE
STATE
n= Percent SUM
100
3 100
17 100
117 100
44 100
4 100
1 100
1 100
1 100
2 100
2 100
1 100
1 100
100
25
25
23
13
100
100
100
100
100
100
100
90
87
80
75
70
75
77
60
Sex
50
U M F
40 30 20 10 8
9
10
11
12
13
16
18
19
20
21
22
Disk Width Length (cm)(cm) - 84 -
.5 55 .0 -5 7
.0 52 .5 -5 5
.5 50 .0 -5 2
.0 47 .5 -5 0
.5 45 .0 -4 7
.5 40 .0 -4 2
.0 32 .5 -3 5
.5 30 .0 -3 2
.0 27 .5 -3 0
.5 25 .0 -2 7
.0 22 .5 -2 5
20 .0 -2 2
.5
0
Inch-class
Figure 86. Width-weight regression for little skate, sexes combined (A) and by sex (B). 5000
4000
Weight(g)
Weight(g) = 0.0554 x Length(cm) 2.8085 (n = 194) 3000
2000
1000
A
0 20
30
40
50
60
Disk Width(cm)
5000 Females: W eight(g) = 0.2536 x Length(cm) 2.3443 (n = 149) Males: W eight(g) = 0.0695 x Length(cm) 2.7514 (n = 45)
Weight(g)
4000
3000
2000
1000
B
0 20
30
40
Disk Width (cm) - 85 -
50
60
Loligo Squid (Priority E)
A
B
Figure 87. Number (A) and biomass (B) of specimens captured at each station for Loligo squid.
- 86 -
Figure 88. Minimum trawlable number and biomass by state for Loligo squid. 5,000,000
300,000,000
Minimum Trawlable Number = 435,880,830 Minimum Trawlable Biomass (kg)= 8,546,121 Pct. S.E. = 16% (number) 14% (biomass)
4,000,000
200,000,000
Number
2,000,000 100,000,000
Avg Wt: 0.025kg
Avg Wt: 0.021kg
Avg Wt: 0.015kg
Avg Wt: 0.028kg
Avg Wt: 0.018kg
Avg Wt: 0.032kg
1,000,000
Avg Wt: 0.037kg
0
0
RI
NY
NJ
DE
MD
VA
NC
State Figure 89. Length frequency histogram for Loligo squid. 30,000
Number Captured = 119,501
Expanded Number
Number Measured = 9,614 20,000
10,000
0 0
5
10
15
Mantle Length (cm) - 87 -
20
25
Biomass (kg)
3,000,000
Northern Searobin (Priority D)
A
B
Figure 90. Number (A) and biomass (B) of specimens captured at each station for northern searobin.
- 88 -
Figure 91. Minimum trawlable number and biomass by state for northern searobin. 500,000
4,000,000
Minimum Trawlable Number = 3,978,887 Minimum Trawlable Biomass (kg)= 456,313
400,000
Pct. S.E. = 21% (number) 28% (biomass)
3,000,000
Number
2,000,000
200,000
1,000,000
100,000
Avg Wt: 0.017kg
Avg Wt: 0.104kg
Avg Wt: 0.124kg
NY
NJ
Avg Wt: 0.086kg
Avg Wt: 0.059kg
Avg Wt: 0.034kg
Avg Wt: 0.034kg
0
0
RI
DE
MD
VA
NC
State Figure 92. Length frequency histogram for northern searobin. 130 120
Expanded Number
110 100
Number Captured = 881
90
Number Measured = 782
80 70 60 50 40 30 20 10 0 0
5
10
15
20
Total Length (cm) - 89 -
25
30
35
Biomass (kg)
300,000
Pinfish (Priority D)
A
B
Figure 93. Number (A) and biomass (B) of specimens captured at each station for pinfish.
- 90 -
Figure 94. Minimum trawlable number and biomass by state for pinfish. 9,000,000
400,000
Minimum Trawlable Number = 8,245,445 8,000,000
Minimum Trawlable Biomass (kg)= 323,903 Pct. S.E. = 85% (number) 83% (biomass)
7,000,000
300,000
6,000,000
Number
200,000 4,000,000
3,000,000
100,000 2,000,000
1,000,000 Avg Wt: 0.097kg
Avg Wt: 0.039kg
0
0
RI
NY
NJ
DE
MD
VA
NC
State Figure 95. Length frequency histogram for pinfish. 1,400 1,300 1,200
Number Captured = 2,744
Expanded Number
1,100
Number Measured = 331
1,000 900 800 700 600 500 400 300 200 100 0 0
5
10
Fork Length (cm) - 91 -
15
20
Biomass (kg)
5,000,000
Scup (Priority A)
A
B
Figure 96. Number (A) and biomass (B) of specimens captured at each station for scup.
- 92 -
Figure 97. Minimum trawlable number and biomass by state for scup. 5,000,000
600,000,000
Minimum Trawlable Number = 943,933,018 Minimum Trawlable Biomass (kg)= 15,083,844 500,000,000
Pct. S.E. = 23% (number) 13% (biomass)
4,000,000
400,000,000
Number
300,000,000
2,000,000 200,000,000 Avg Wt: 0.014kg
Avg Wt: 0.005kg
Avg Wt: 0.036kg
Avg Wt: 0.062kg
Avg Wt: 0.048kg
Avg Wt: 0.050kg
Avg Wt: 0.047kg
1,000,000 100,000,000
0
0
RI
NY
NJ
DE
MD
VA
NC
State Figure 98. Length frequency histogram for scup.
Expanded Number
90,000 80,000
Number Captured = 276,234
70,000
Number Measured = 13,718 Number of Otoliths = 808
60,000
Number of Stomachs = 797 50,000 40,000 30,000 20,000 10,000 0 0
5
10
15
20
25
Fork Length (cm) - 93 -
30
35
40
Biomass (kg)
3,000,000
Figure 99. Sex ratios for scup by state (A) and length group (B).
A n = 219
142
149
77
40
128
40
100.0
100.0
100.0
100.1
100.1
100.0
99.9
92.7
99.9
60.9
37.8
10.3
14.0
29.9
100 90
49.0 36.1
80
Percent
70
21.1
60
26.9
Sex
50 40
49.9
48.9
6-VA
7-NC
U M F
40.8
18.3 35.4
30 20
20.8
10
5.7
0
1-RI
2-NY
3-NJ
4-DE
B
5-MD
STATE
n= Percent SUM
100
1 100.0
100
95 159 100.0 100.0
100
99
107 99.9
46
205 170 100.1 100.0
24 100.0
13
66
18 9 100.0 100.0
11
33
3 100.0
3 100.0
1 100.0
97
33
100
41
90
89
36 80 70
67
67
60
23
50
52
55
Sex U M F
40
35
30
30
20 10 1
2
3
4
5
6
7
8
9
10
12
13
Length (cm)
- 94 -
.0 32 .5 -3 5
.5 30 .0 -3 2
.5 25 .0 -2 7
.0 22 .5 -2 5
.5 20 .0 -2 2
.0 17 .5 -2 0
.5 15 .0 -1 7
.0 12 .5 -1 5
.5 10 .0 -1 2
.0 07 .5 -1 0
.5 05 .0 -0 7
02 .5 -0 5
.0
0
Inch-class
Figure 100. Maturity logistic regression for scup, by sex. 1.0 99% maturity
0.9
0.7 0.6 0.5
50% maturity
0.4 0.3 0.2 0.1
SEX
F M
0.0 0
10
20
30
40
Fork Length (cm) Figure 101. Expanded age frequency histogram for scup.
2007
300000
Catch curve Z ages 0-4 = -2.82
241790 200000
100000
28476 1027
108
3
0
Age (Year Class) - 95 -
(2 00 2) 5
(2 00 3) 4
(2 00 4) 3
(2 00 5) 2
(2 00 6) 1
(2 00 7)
0
0
Probability of Maturity
0.8
Figure 102. Age-specific length-frequency histograms for scup for ages 0 through 3. AG E = 0 4 ,0 0 0
3 ,0 0 0
2 ,0 0 0
1 ,0 0 0
0 0
5
10
15
20
25
30
35
40
T o t a l Le n g t h ( c m ) AG E = 1 800 700 600 500 400 300 200 100 0 0
5
10
15
20
25
30
35
40
25
30
35
40
25
30
35
40
To t a l L e n g t h ( c m ) AG E = 2 30
20
10
0 0
5
10
15
20
To ta l Le n g th (c m ) AG E = 3 30
20
10
0 0
5
10
15
20
F o r k Le n g th (c m )
- 96 -
Figure 103. von Bertalanffy growth curves for scup, by sex. 40
Fe m ale s : L(m ax) = 70.5 T(ze ro) = 0.263 K = 0.232 M ale s :
Length (cm)
30
L(m ax) = 46.2 T(ze ro) = 0.401 K = 0.453
20
10
0 0
1
2
3
Age
- 97 -
4
5
Figure 104. Length-weight regression for little skate, sexes combined (A) and by sex (B). 900 800 700 Weight(g) = 0.0144 x Length(cm) 3.1465 (n = 806)
Weight(g)
600 500 400 300 200
A
100 0 0
10
20
30
40
Fork Length(cm)
900 800
Females: W eight(g) = 0.015 x Length(cm) 3.1337 (n = 305)
700
Males: W eight(g) = 0.013 x Length(cm) 3.1791 (n = 174)
Weight(g)
600 500 400 300 200
B
100 0
0
10
20
Fork Length(cm)
- 98 -
30
40
Figure 105. Diets of scup by percent weight (A) and percent number (B).
A
B
- 99 -
Silver Hake (Whiting) (Priority A)
A
B
Figure 106. Number (A) and biomass (B) of specimens captured at each station for silver hake.
- 100 -
Figure 107. Minimum trawlable number and biomass by state for silver hake. 110,000
1,300,000 1,200,000
100,000
1,100,000
90,000
Minimum Trawlable Number = 1,480,829 1,000,000
Minimum Trawlable Biomass (kg)= 107,251 900,000
80,000
Pct. S.E. = 60% (number) 66% (biomass) 70,000
700,000
60,000
600,000
50,000
500,000
40,000
400,000
30,000 300,000
20,000 Avg Wt: 0.078kg
200,000
Avg Wt: 0.006kg
Avg Wt: 0.005kg
Avg Wt: 0.083kg
10,000
100,000
0
0
RI
NY
NJ
DE
MD
VA
NC
State Figure 108. Length frequency histogram for silver hake. 70
Expanded Number
60
Number Captured = 346 Number Measured = 346
50
Number of Otoliths = 59 Number of Stomachs = 59
40
30
20
10
0 0
5
10
15
Fork Length (cm) - 101 -
20
25
30
Biomass (kg)
Number
800,000
Figure 109. Sex ratios for silver hake by state (A) and length group (B).
A
n= 100
46
100
5100
100
7
80
100
33
90 80
Percent
70
63
60
Sex
50
U M F
40 30 20
20
10 0 1-RI
2-NY
3-NJ
STATE
B n= Percent SUM
100
3 100.0
10 100.0
4 100.0
7 100.0
5 99.9
6 100.0
14 100.0
8 100.0
1 100.0
100
100
100
20
43
36
39
100
100
90 80
13
70
68
64
60
61
14
50 40
Sex U M F
43
30 20 10 2
3
4
5
6
8
9
10
11
Length (cm)
- 102 -
.0 27 .5 -3 0
.5 25 .0 -2 7
.0 22 .5 -2 5
.5 20 .0 -2 2
.5 15 .0 -1 7
.0 12 .5 -1 5
.5 10 .0 -1 2
.0 07 .5 -1 0
05 .0 -0 7
.5
0
Inch-class
Figure 110. Maturity logistic regression for silver hake, by sex. 1.0 99% maturity
0.9
Probability of Maturity
0.8 0.7 0.6 0.5
50% maturity
0.4 0.3 0.2 0.1
SEX
F M
0.0 12
13
14
15
16
17
18
19
20
21
Fork Length (cm)
- 103 -
22
23
24
25
26
27
28
Figure 111. Length-weight regression for silver hake, sexes combined (A) and by sex (B). 140 130 120 110 Weight(g) = 0.0074 x Length(cm) 2.9604 (n = 59)
100
Weight(g)
90 80 70 60 50 40 30
A
20 10 0 0
10
20
30
Fork Length(cm)
140 130
Females: W eight(g) = 0.0079 x Length(cm) 2.9399 (n = 30)
120
Males: (n = 8)
110
W eight(g) = 0.0195 x Length(cm) 2.6403
100
Weight(g)
90 80 70 60 50 40
B
30 20 10
12
13
14
15
16
17
18
19
20
21
Fork Length(cm)
- 104 -
22
23
24
25
26
27
28
Silver Perch (Priority D)
A
B
Figure 112. Number (A) and biomass (B) of specimens captured at each station for silver perch.
- 105 -
Figure 113. Minimum trawlable number and biomass by state for silver perch. 140,000
6,000,000
130,000
Minimum Trawlable Number = 6,971,143 5,000,000
120,000
Minimum Trawlable Biomass (kg)= 189,861 110,000
Pct. S.E. = 38% (number) 30% (biomass) 100,000 4,000,000
90,000
Number
70,000
3,000,000
60,000 50,000 2,000,000
40,000 30,000 1,000,000
Avg Wt: 0.052kg
Avg Wt: 0.025kg
Avg Wt: 0.032kg
Avg Wt: 0.039kg
20,000 10,000 0
0
RI
NY
NJ
DE
MD
VA
NC
State Figure 114. Length frequency histogram for silver perch. 500
Number Captured = 1,398
400
Expanded Number
Number Measured = 141
300
200
100
0 0
5
10
Total Length (cm) - 106 -
15
20
Biomass (kg)
80,000
Smooth Butterfly Ray (Priority B)
A
B
Figure 115. Number (A) and biomass (B) of specimens captured at each station for smooth butterfly ray.
- 107 -
Figure 116. Minimum trawlable number and biomass by state for smooth butterfly ray. 1,800,000
600,000
1,700,000
500,000
Minimum Trawlable Number = 1,090,128
1,600,000
Minimum Trawlable Biomass (kg)= 2,182,600
1,500,000
Pct. S.E. = 21% (number) 19% (biomass)
1,400,000 1,300,000 1,200,000
400,000
Number
1,000,000 900,000
300,000
800,000 700,000 600,000
200,000
500,000 400,000 100,000
Avg Wt: 3.032kg
300,000
Avg Wt: 0.854kg
200,000 100,000 0
0
RI
NY
NJ
DE
MD
VA
NC
State Figure 117. Width frequency histogram for smooth butterfly ray. 30
Number Captured = 292
Expanded Number
Number Measured = 292 20
10
0 0
5
1 0
1 5
2 0
2 5
3 0
3 5
4 0
4 5
5 0
5 5
6 0
6 5
7 0
Disk Width (cm) - 108 -
7 5
8 0
8 5
9 0
9 5
1 0 0
1 0 5
1 1 0
1 1 5
Biomass (kg)
1,100,000
Smooth Dogfish (Priority B)
A
B
Figure 118. Number (A) and biomass (B) of specimens captured at each station for smooth dogfish.
- 109 -
Figure 119. Minimum trawlable number and biomass by state for smooth dogfish. 4,000,000
2,000,000
Minimum Trawlable Number = 7,381,432
1,900,000 1,800,000
Minimum Trawlable Biomass (kg)= 6,667,209
1,700,000
Pct. S.E. = 32% (number) 53% (biomass)
1,600,000
3,000,000
1,500,000 1,400,000 1,300,000
Number
1,100,000
2,000,000
1,000,000 900,000 800,000 700,000 600,000
1,000,000
500,000 400,000
Avg Wt: 2.821kg
300,000
Avg Wt: 1.995kg
Avg Wt: 0.592kg
Avg Wt: 0.240kg
Avg Wt: 0.551kg
Avg Wt: 0.516kg
200,000 100,000
0
0
RI
NY
NJ
DE
MD
VA
NC
State Figure 120. Length frequency histogram for smooth dogfish. 300
Number Captured = 1,690
Expanded Number
Number Measured = 765 Number of Vertebrae = 202 200
Number of Stomachs = 200
100
0 30
35
40
45
50
55
60
65
70
75
Precaudal Length (cm) - 110 -
80
85
90
95
100
105
Biomass (kg)
1,200,000
Figure 121. Sex ratios for smooth dogfish by state (A) and length group (B).
A
n= 100
11 100
36 100
100
52
100
28
100
37
100
38
88.5
37.0
87.6
39.4
70.6
76.7
90 80
Percent
70 63.0
60
60.6 Sex
50
U M F
40 30 29.4
23.3
20 10
12.4
11.5
0 1-RI
2-NY
3-NJ
4-DE
100
6-VA
STATE
B n= Percent SUM
5-MD
2 100
33 100
91 100
8 100
5 100
7 100
4 100
8 100
14 100
12 100
9 100
5 100
1 100
3 100
77
67
61
94
29
79
100
68
35
37
28
24
100
33
90 80
76
70
72
71 65
60
67
63
Sex
50
U M F
40
39 30 20
33
32
23
21
10
Length (cm)
- 111 -
34-36
36-38
38+ Inch-class 95 +
32-34
90 -9 5
30-32
85 -9 0
28-30
80 -8 5
26-28
75 -8 0
24-26
70 -7 5
22-24
65 -7 0
20-22
60 -6 5
6
55 -6 0
40 -4 5
35 -4 0
30 -3 5
18-20
50 -5 5
14-16 16-18
45 -5 0
12-14
0
Figure 122. Maturity logistic regression for smooth dogfish, by sex. 1.0
99% maturity 0.9
Probability of Maturity
0.8 0.7 0.6 0.5
50% maturity
0.4 0.3 0.2 0.1
SEX
F M
0.0 30
40
50
60
70
80
90
Precaudal Length (cm)
- 112 -
100
110
120
130
Figure 123. Length-weight regression for smooth dogfish, sexes combined (A) and by sex (B). 7000
6000
Weight(g) = 0.0871 x Length(cm) 2.3552 (n = 202)
Weight(g)
5000
4000
3000
2000
A
1000
0 30
40
50
60
70
80
90
100
Precaudal Length(cm)
7000 Females: W eight(g) = 0.0312 x Length(cm) 2.6113 (n = 90)
6000
Males: W eight(g) = 0.3867 x Length(cm) 1.9733 (n = 112)
Weight(g)
5000
4000
3000
2000
B
1000
0 30
40
50
60
70
Precaudal Len gth (cm) - 113 -
80
90
100
Spanish Mackerel (Priority D)
A
B
Figure 124. Number (A) and biomass (B) of specimens captured at each station for Spanish mackerel.
- 114 -
Figure 125. Minimum trawlable number and biomass by state for Spanish mackerel. 80,000
400,000
Minimum Trawlable Number = 618,880 Minimum Trawlable Biomass (kg)= 167,037
70,000
Pct. S.E. = 25% (number) 21% (biomass) 60,000
300,000
40,000
200,000
30,000
100,000
Avg Wt: 0.010kg
Avg Wt: 0.660kg
Avg Wt: 0.409kg
Avg Wt: 0.296kg
Avg Wt: 0.231kg
20,000
10,000
0
0
RI
NY
NJ
DE
MD
VA
NC
State Figure 126. Length frequency histogram for Spanish mackerel. 30
Number Captured = 161
Expanded Number
Number Measured = 161 20
10
0 0
5
10
15
20
25
Fork Length (cm) - 115 -
30
35
40
45
Biomass (kg)
Number
50,000
Spot (Priority B)
A
B
Figure 127. Number (A) and biomass (B) of specimens captured at each station for spot.
- 116 -
Figure 128. Minimum trawlable number and biomass by state for spot. 10,000,000
110,000,000
Minimum Trawlable Number = 174,432,892
100,000,000
9,000,000
Minimum Trawlable Biomass (kg)= 15,589,723
90,000,000
8,000,000
Pct. S.E. = 31% (number) 31% (biomass) 80,000,000
7,000,000 70,000,000
Number
60,000,000
5,000,000 50,000,000
4,000,000 40,000,000
3,000,000 30,000,000 Avg Wt: 0.111kg
Avg Wt: 0.109kg
Avg Wt: 0.105kg
Avg Wt: 0.090kg
Avg Wt: 0.080kg
20,000,000
2,000,000
1,000,000
10,000,000
0
0
RI
NY
NJ
DE
MD
VA
NC
State Figure 129. Length frequency histogram for spot. 12,000
Expanded Number
11,000 10,000
Number Captured = 44,437
9,000
Number Measured = 2,507
8,000
Number of Otoliths = 160
7,000
Number of Stomachs = 9
6,000 5,000 4,000 3,000 2,000 1,000 0 0
5
10
15
Fork Length (cm) - 117 -
20
25
Biomass (kg)
6,000,000
Figure 130. Sex ratios for spot by state (A) and length group (B).
A n= 100
2
38
100
100
100
33
3
29
100
100
33
37
57
100
30
100 75
56
90 80
64
60
67 63
Sex
50 40
U M F
40
30 25
20 10 0 1-RI
3-NJ
4-DE
5-MD
6-VA
7-NC
STATE
100
4 100.0
70 100.1
82 100.0
3 100.0
43.8
8.6
65.9
95.7
90
40.8
80 70 60 50
56.2
Sex 50.7
U M F
40
34.1
30 20 10 5
6
7
8
Length (cm)
- 118 -
.5 20 .0 -2 2
17 .5 -2 0
15 .0 -1 7
.0
.5
0 .0
B
Percent SUMn =
12 .5 -1 5
Percent
70
Inch-class
Figure 131. Maturity logistic regression for spot, by sex. 1.0
99% maturity 0.9
Probability of Maturity
0.8 0.7 0.6 0.5
50% maturity 0.4 0.3 0.2 0.1
SEX
F M
0.0 13
14
15
16
17
18
19
Fork Length (cm)
- 119 -
20
21
22
23
Weight(g)
Figure 132. Length-weight regression for spot, sexes combined (A) and by sex (B). 220 210 200 190 180 170 160 150 140 130 120 110 100 90 80 70 60 50 40 30
Weight(g) = 0.0126 x Length(cm)3.1211 (n = 159)
A
13
14
15
16
17
18
19
20
21
22
Fork Length(cm)
Weight(g)
220 210
Fe m ale s: We ight(g) = 0.007 x Le ngth(cm ) 3.3293 (n = 94)
200 190 180 170 160 150 140 130 120 110 100 90 80 70 60 50 40 30
Male s: (n = 62)
We ight(g) = 0.0295 x Le ngth(cm ) 2.819
B 13
14
15
16
17
18
Fork Length(cm) - 120 -
19
20
21
22
Striped Anchovy (Priority D)
A
B
Figure 133. Number (A) and biomass (B) of specimens captured at each station for striped anchovy.
- 121 -
Figure 134. Minimum trawlable number and biomass by state for striped anchovy. 5,000,000
500,000,000
Minimum Trawlable Number = 878,956,591 Minimum Trawlable Biomass (kg)= 9,935,140 Pct. S.E. = 41% (number) 40% (biomass)
400,000,000
4,000,000
200,000,000
2,000,000
Number
3,000,000
Avg Wt: 0.015kg
Avg Wt: 0.011kg
Avg Wt: 0.011kg
1,000,000
100,000,000
Avg Wt: 0.020kg
Avg Wt: 0.008kg
Avg Wt: 0.015kg
Avg Wt: 0.013kg
0
0
RI
NY
NJ
DE
MD
VA
NC
State Figure 135. Length frequency histogram for striped anchovy. 80,000
70,000
Number Captured = 224,369 Number Measured = 4,990
Expanded Number
60,000
50,000
40,000
30,000
20,000
10,000
0 0
5
10
Fork Length (cm) - 122 -
15
Biomass (kg)
300,000,000
Striped Searobin (Priority D)
A
B
Figure 136. Number (A) and biomass (B) of specimens captured at each station for striped searobin.
- 123 -
Figure 137. Minimum trawlable number and biomass by state for striped searobin. 600,000
1,900,000 1,800,000 1,700,000
Minimum Trawlable Number = 2,709,314
1,600,000
Minimum Trawlable Biomass (kg)= 659,547
1,500,000
Pct. S.E. = 38% (number) 57% (biomass)
500,000
1,400,000 1,300,000
400,000
1,200,000
Number
1,000,000
300,000 900,000 800,000 700,000
200,000
600,000 500,000 400,000
100,000
300,000
Avg Wt: 0.200kg
200,000
Avg Wt: 0.278kg
Avg Wt: 0.151kg
NY
NJ
Avg Wt: 0. 190kg
Avg Wt: 0. 032kg
Avg Wt: 0. 054kg
Avg Wt: 0. 016kg
100,000
0
0
RI
DE
MD
VA
NC
State Figure 138. Length frequency histogram for striped searobin. 70
Number Captured = 760 Number Measured = 546
Expanded Number
60
50
40
30
20
10
0 0
5
10
15
20
25
Total Length (cm) - 124 -
30
35
40
45
Biomass (kg)
1,100,000
Summer Flounder (Priority A)
A
B
Figure 139. Number (A) and biomass (B) of specimens captured at each station for summer flounder.
- 125 -
Figure 140. Minimum trawlable number and biomass by state for summer flounder. 700,000
1,300,000 1,200,000
Minimum Trawlable Number = 3,600,065
1,100,000
600,000
Minimum Trawlable Biomass (kg)= 2,283,875
1,000,000
500,000
900,000
Pct. S.E. = 9% (number) 10% (biomass)
800,000
Number
600,000
300,000 500,000 400,000
200,000
300,000 200,000
Avg Wt: 1.113kg
Avg Wt: 0.865kg
Avg Wt: 0.607kg
NY
NJ
Avg Wt: 0.686kg
Avg Wt: 0.579kg
Avg Wt: 0.333kg
100,000
Avg Wt: 0.325kg
100,000
0
0
RI
DE
MD
VA
NC
State Figure 141. Length frequency histogram for summer flounder. 60
Number Captured = 960 Number Measured = 926
50
Expanded Number
Number of Otoliths = 716 Number of Stomachs = 448
40
30
20
10
0 0
5
10
15
20
25
30
35
40
45
50
Total Length (cm) - 126 -
55
60
65
70
75
80
Biomass (kg)
400,000 700,000
Figure 142. Sex ratios for summer flounder by state (A) and length group (B).
A
n = 100.0 110
100
180
78
100.0
99.9
50
100.0
25.1
44.3
41.8
29.2
47
100.0
231
33.7
19
100.0
100.1
45.5
5.3 21.1
90 80
74.9
Percent
70
73.7
70.0
60
62.7
58.2
55.7
50
Sex
52.2
U M F
40 30 20 10 0 1-RI
2-NY
3-NJ
4-DE
B Percent SUM n= 100
5-MD
6-VA
7-NC
STATE
3 2 70 106 132 118 97 90 56 24 11 4 1 1 99.9 100.0 100.0 100.1 100.0 100.1 100.0 100.0 100.0 100.0 100.0 100.0 100.0 100.0
33
39
7
60
42
53
32
13
100
100
100
100
100
100
65
90
87 80 70 60
68
33 62 58
Sex
50
U M F
46
40
39 30
33 28
20 10 4-6
6-8
8-10 10-12
12-14 14-16 16-18 18-20 20-22 22-24 24-26
26-28 28-30 30-32 Inch-class
Length (cm)
- 127 -
75 -8 0
70 -7 5
65 -7 0
60 -6 5
55 -6 0
50 -5 5
45 -5 0
40 -4 5
35 -4 0
30 -3 5
25 -3 0
20 -2 5
15 -2 0
10 -1 5
0
Figure 143. Maturity logistic regression for summer flounder, by sex. 1.0 99% maturity
0.9
0.7 0.6 0.5
50% maturity
0.4 0.3 0.2 0.1
SEX
F M
0.0 10
20
30
40
50
60
70
80
Total Length (cm) Figure 144. von Bertalanffy growth curves for summer flounder, by sex. 80 Females: L(max) = 70.5 T(zero) = 0.263 K = 0.232 Males:
L(max) = 46.2 T(zero) = 0.401 K = 0.453
70
60
Length (cm)
Probability of Maturity
0.8
50
40
30
20
10 0
1
2
3
4
5
6
7
Age
- 128 -
8
9
10
11
12
13
14
- 129 -
Age (Year Class)
1)
0
(1 99
0)
1)
0
(1 99
16
(1 99
2)
Expanded Number
245
16
2)
1
(1 99
15
(1 99
3)
0
15
3)
0
(1 99
14
(1 99
4)
0
14
4)
2
(1 99
13
(1 99
5)
0
13
5)
3
(1 99
12
(1 99
6)
)
)
)
)
0
12
6)
2
(1 99
11
99 7
(1 99
(1
99 8
99 9
00 0
)
1
11
7)
3
(1 99
10
9
(1
(1
(2
00 1
)
)
)
)
0
10
)
3
99 8
215
(1
)
8
7
6
(2
00 2
00 3
00 4
00 5
)
3
9
99 9
)
)
7
(1
00 0
00 1
)
5
(2
(2
(2
(2
00 6
4
8
(2
(2
00 2
)
4
3
2
1
(2
5
7
6
(2
00 3
)
)
)
)
0
18
5
(2
00 4
00 5
00 6
00 7
209
4
(2
(2
(2
(2
100
3
2
1
0
Expanded Number
Figure 145. Age frequency histogram for summer flounder for 2006 (A) and 2007 (B). 300
A
200
94
70
17
0
0 0
Age (Year Class)
300
B
243
Catch curve Z ages 1-14 = -0.51
200
149
100
52
36
0
0
0
Figure 146. Age-specific length-frequency histograms for summer flounder for ages 0 through 6. AG E = 0 4 0
3 0
2 0
1 0
0 0
5
1 0
1 5
2 0
2 5
3 0
3 5
4 0
A G E =T o1t a
4 5
5 0
5 5
6 0
6 5
7 0
7 5
8 0
4 5
5 0
5 5
6 0
6 5
7 0
7 5
8 0
4 5
5 0
5 5
6 0
6 5
7 0
7 5
8 0
4 5
5 0
5 5
6 0
6 5
7 0
7 5
8 0
l L e n g th (c m )
4 0
3 0
2 0
1 0
0 0
5
1 0
1 5
2 0
2 5
3 0
3 5
4 0
A G E =T o2t a
l L e n g th (c m )
1 4 1 3 1 2 1 1 1 0 9 8 7 6 5 4 3 2 1 0 0
5
1 0
1 5
2 0
2 5
3 0
3 5
4 0
A G E =T o3t a
l L e n g th (c m )
1 8 1 7 1 6 1 5 1 4 1 3 1 2 1 1 1 0 9 8 7 6 5 4 3 2 1 0 0
5
1 0
1 5
2 0
2 5
3 0
3 5
4 0
T o ta l L e n g th (c m )
AG E = 4 4
3
2
1
0
0
5
1 0
1 5
2 0
2 5
3 0
3 5
4 0
ta A G E T=o 5
4 5
5 0
5 5
6 0
6 5
7 0
7 5
8 0
4 5
5 0
5 5
6 0
6 5
7 0
7 5
8 0
4 5
5 0
5 5
6 0
6 5
7 0
7 5
8 0
l L e n g th (c m )
3
2
1
0 0
5
1 0
1 5
2 0
2 5
3 0
3 5
4 0
T o ta l L e n g th (c m )
AG E = 6 2
1
0 0
5
1 0
1 5
2 0
2 5
3 0
3 5
4 0
T o ta l L e n g th (c m )
- 130 -
Figure 147. Length-weight regression for summer flounder, sexes combined (A) and by sex (B). 6000
5000 Weight(g) = 0.0033 x Length(cm) 3.3013 (n = 716)
Weight(g)
4000
3000
2000
A
1000
0 10
20
30
40
50
60
70
80
Total Length(cm)
6000 Females: W eight(g) = 0.003 x Length(cm) 3.3221 (n = 448)
5000
Males: W eight(g) = 0.006 x Length(cm) 3.1409 (n = 256)
Weight(g)
4000
3000
2000
1000
B
0
10
20
30
40
50
Total Len gth(cm)
- 131 -
60
70
80
Figure 148. Diets of summer flounder by percent weight (A) and percent number (B).
A
B
- 132 -
Weakfish (Priority A)
A
B
Figure 149. Number (A) and biomass (B) of specimens captured at each station for weakfish.
- 133 -
Figure 150. Minimum trawlable number and biomass by state for summer weakfish. 8,000,000
100,000,000
Minimum Trawlable Number = 230,584,843
90,000,000
7,000,000
Minimum Trawlable Biomass (kg)= 16,269,075 80,000,000
Pct. S.E. = 20% (number) 20% (biomass) 6,000,000
70,000,000
5,000,000
Number
4,000,000
50,000,000
40,000,000
3,000,000 30,000,000
Avg Wt: 0.031kg
Avg Wt: 0.061kg
Avg Wt: 0.112kg
Avg Wt: 0.109kg
Avg Wt: 0.074kg
Avg Wt: 0.078kg
Avg Wt: 0.081kg
2,000,000
20,000,000
1,000,000 10,000,000
0
0
RI
NY
NJ
DE
MD
VA
NC
State Figure 151. Length frequency histogram for weakfish. 9,000
Number Captured = 60,990
8,000
Number Measured = 5,747
Expanded Number
7,000
Number of Otoliths = 572 6,000
Number of Stomachs = 471
5,000 4,000 3,000 2,000 1,000 0 0
5
10
15
20
25
30
35
Total Length (cm) - 134 -
40
45
50
55
60
Biomass (kg)
60,000,000
Figure 152. Sex ratios for weakfish by state (A) and length group (B).
A
n=
77
100
50
161
16
72
129
100.1
100.0
100.1
100.0
100.0
100.0
100.0
26.5
12.6
52.6
74.6
49.3
11.1
53.1
67
90
40.9
87.4 80
31.6
Percent
70 60
Sex
50
50.7
47.3 40
48.0
46.8
6-VA
7-NC
U M F
42.0
30
25.4
20 10 0 1-RI
2-NY
3-NJ
4-DE STATE
B n= Percent SUM
100
5-MD
2 100
45 100
171 100
192 100
116 100
29 100
9 100
5 100
1 100
1 100
1 100
100
30
13
45
31
21
16
24
100
100
100
90
38
84
80
80 76
70
40
69
60
Sex
56
50
50
U M F
40 30
30
20 10 2-4
4-6
6-8
8-10
10-12
12-14
14-16
16-18
18-20
20-22
22-24 Inch-class
Length (cm)
- 135 -
55 -6 0
50 -5 5
45 -5 0
40 -4 5
35 -4 0
30 -3 5
25 -3 0
20 -2 5
15 -2 0
10 -1 5
05 -1 0
0
Figure 153. Maturity logistic regression for weakfish, by sex. 1.0 99% maturity 0.9
Probability of Maturity
0.8 0.7 0.6 0.5 50% maturity 0.4 0.3 0.2 0.1
SEX
F M
0.0 10
20
30
40
50
60
70
Total Length (cm) Figure 154. Age frequency histogram for weakfish.
40000
34312 Catch curve Z ages 0-3 = -1.49
30000
17622
20000
10000
6017 342
Age (Year Class) - 136 -
5
(2 00 3) 4
(2 00 4) 3
(2 00 5) 2
(2 00 6) 1
0
(2 00 7)
0
0 (2 00 2)
0
Figure 155. Age-specific length-frequency histograms for weakfish for ages 0 through 3. AG E = 0 500
400
300
200
100
0 0
5
10
15
20
25
30
35
40
45
50
55
60
40
45
50
55
60
40
45
50
55
60
To t a l L e n g t h ( c m ) AG E = 1 600 500 400 300 200 100 0 0
5
10
15
20
25
30
35
To t a l L e n g t h ( c m ) AG E = 2 160 140 120 100 80 60 40 20 0 0
5
10
15
20
25
30
35
To t a l L e n g t h ( c m ) AG E = 3 16 14 12 10 8 6 4 2 0 0
5
10
15
20
25
30
35
To ta l Le n g th (c m )
- 137 -
40
45
50
55
60
Figure 156. von Bertalanffy growth curves for weakfish, by sex. 60
Fe m ale s : L(m ax) = 70.5 T(ze ro) = 0.263 K = 0.232
50
Length (cm)
M ale s :
L(m ax) = 46.2 T(ze ro) = 0.401 K = 0.453
40
30
20
10 0
1
2
Age
- 138 -
3
4
Weight(g)
Figure 157. Length-weight regression for weakfish, sexes combined (A) and by sex (B).. 1800 1700 1600 1500 1400 1300 1200 1100 1000 900 800 700 600 500 400 300 200 100 0
Weight(g) = 0.0099 x Length(cm)2.9942 (n = 571)
A 0
10
20
30
40
50
60
Total Length(cm)
1800
Fe m ale s: We ight(g) = 0.0098 x Le ngth(cm ) 2.9949 (n = 280)
1700 1600
Male s: We ight(g) = 0.0103 x Le ngth(cm ) 2.9844 (n = 257)
1500
Weight(g)
1400 1300 1200 1100 1000 900 800 700 600 500 400 300 200
B
100 0
10
20
30
40
T otal Len gth(cm) - 139 -
50
60
Figure 158. Diets of weakfish by percent weight (A) and percent number (B).
A
B
- 140 -
Winter Flounder (Priority A)
A
B
Figure 159. Number (A) and biomass (B) of specimens captured at each station for winter flounder.
- 141 -
Figure 160. Minimum trawlable number and biomass by state for winter flounder. 500,000
1,700,000 1,600,000 1,500,000
Minimum Trawlable Number = 1,639,213
1,400,000
Minimum Trawlable Biomass (kg)= 413,370
400,000
1,300,000
Pct. S.E. = 32% (number) 27% (biomass)
1,200,000 1,100,000
Number
900,000 800,000 700,000
200,000
600,000 500,000
Avg Wt: 0.252kg
400,000
100,000
300,000 200,000 100,000
0
0
RI
NY
NJ
DE
MD
VA
NC
State Figure 161. Length frequency histogram for winter flounder. 60
Number Captured = 391 50
Number Measured = 391
Expanded Number
Number of Otoliths = 118 40
Number of Stomachs = 114
30
20
10
0 0
5
10
15
20
25
Total Length (cm) - 142 -
30
35
40
45
Biomass (kg)
300,000
1,000,000
Figure 162. Sex ratios for winter flounder by state (A) and length group (B).
A
n=
118
100
100
27
90 80 70
69 Percent
60
Sex
50
U M F
40 30 20 10 0 1-RI STATE
B n= Percent SUM
100
1 100
8 100
100
15 100
14 96
63
90
15 100
17 100
22
11 100
16 100
19 100
22
95
8 100
7 100
1 100
100
100
100
92
46
80
78
76
70 60
Sex
50 40
U M F
40
30
33
20 10 6
7
8
9
10
11
12
13
14
15
17
Length (cm)
- 143 -
.0 42 .5 -4 5
.0 37 .5 -4 0
.5 35 .0 -3 7
.0 32 .5 -3 5
.5 30 .0 -3 2
.0 27 .5 -3 0
.5 25 .0 -2 7
.0 22 .5 -2 5
.5 20 .0 -2 2
.0 17 .5 -2 0
15 .0 -1 7
.5
0
Inch-class
Figure 163. Maturity logistic regression for winter flounder, by sex. 1.0
99% maturity 0.9
Probability of Maturity
0.8 0.7 0.6 0.5
50% maturity
0.4 0.3 0.2 0.1
SEX
F M
0.0 10
20
30
40
50
Total Length (cm) Figure 164. Age frequency histogram for winter flounder. 300
235
200
100
87 62
4
0
2
1
0
0
0
0
Age (Year Class) - 144 -
(1 99 7) 10
(1 99 8) 9
(1 99 9) 8
(2 00 0) 7
(2 00 1) 6
(2 00 2) 5
(2 00 3) 4
(2 00 4) 3
(2 00 5) 2
(2 00 6) 1
(2 00 7)
0 0
Expanded Number
Catch curve Z ages 1-7 = -1.01
Figure 165. Age-specific length-frequency histograms for winter flounder for ages 1 through 3. AG E = 1 8 7 6 5 4 3 2 1 0 0
5
10
15
20
25
30
35
40
45
30
35
40
45
30
35
40
45
T o t a l Le n g t h ( c m ) AG E = 2 4
3
2
1
0 0
5
10
15
20
25
T o t a l Le n g t h ( c m ) AG E = 3 7 6 5 4 3 2 1 0 0
5
10
15
20
25
T o t a l Le n g t h ( c m )
- 145 -
Figure 166. von Bertalanffy growth curves for winter flounder, by sex. 50
Fe m ale s : L(m ax) = 70.5 T(ze ro) = 0.263 K = 0.232 M ale s :
Length (cm)
40
L(m ax) = 46.2 T(ze ro) = 0.401 K = 0.453
30
20
10 1
2
3
4
5
Age
- 146 -
6
7
8
Figure 167. Length-weight regression for winter flounder, sexes combined (A) and by sex (B). 1100 1000 900 800
Weight(g) = 0.0115 x Length(cm) 3.0331 (n = 118)
Weight(g)
700 600 500 400 300 200 100 0 10
20
30
40
50
Total Length(cm)
1100 1000
Females: W eight(g) = 0.0126 x Length(cm) 3.0084 (n = 95)
900
Males: W eight(g) = 0.0079 x Length(cm) 3.1393 (n = 18)
800
Weight(g)
700 600 500 400 300 200 100 0 10
20
30
T otal Length(cm) - 147 -
40
50
Figure 168. Diets of winter flounder by percent weight (A) and percent number (B).
A
B
- 148 -
Winter Skate (Priority B)
A
B
Figure 169. Number (A) and biomass (B) of specimens captured at each station for winter skate.
- 149 -
Figure 170. Minimum trawlable number and biomass by state for winter skate. 1,400,000
1,900,000 1,800,000
1,300,000
1,700,000
Minimum Trawlable Number = 3,048,125
1,600,000
1,200,000
Minimum Trawlable Biomass (kg)= 2,756,000
1,500,000
1,100,000
Pct. S.E. = 29% (number) 18% (biomass)
1,400,000
1,000,000 1,200,000
900,000
1,100,000
800,000
1,000,000
700,000 900,000 800,000
600,000
700,000
500,000
600,000
400,000
500,000
300,000
400,000 300,000
Avg Wt: 0.752kg
200,000
Avg Wt: 1.127kg
200,000
Avg Wt: 1.270kg
100,000
100,000
0
0
RI
NY
NJ
DE
MD
VA
NC
State Figure 171. Width frequency histogram for winter skate. 70
Number Captured = 951
Expanded Number
60
Number Measured = 735 Number of Vertebrae = 171
50
Number of Stomachs = 160 40
30
20
10
0 0
5
10
15
20
25
30
35
Disk Width (cm) - 150 -
40
45
50
55
60
Biomass (kg)
Number
1,300,000
Figure 172. Sex ratios for winter skate by state (A) and width group (B).
A
n= 100
81
84
100
100
100
55.7
22.5
75.0
4
90 80
76.7
Percent
70 60
Sex
50
U M F
44.3
40 30
25.0
20 10 0 1-RI
2-NY
3-NJ
STATE
B n= Percent SUM
100
1 100
6 100
21 100
28 100
34 100
100
19
60
23
28
37 100
12
90
30 100
11 100
1 100
51
62
100
85
80
81
77
70
72
60
Sex
50
49 40
40
U M F
38
30 20 10 4-6
6-8
8-10
10-12
12-14
14-16
16-18
18-20
22-24
Length (cm) (cm) Disk Width
- 151 -
55 -6 0
45 -5 0
40 -4 5
35 -4 0
30 -3 5
25 -3 0
20 -2 5
15 -2 0
10 -1 5
0
Inch-class
Figure 173. Width-weight regression for winter skate, sexes combined (A) and by sex (B). 6000
5000
Weight(g) = 0.0269 x Length(cm) 3.0104 (n = 171)
Weight(g)
4000
3000
2000
1000
0 10
20
30
40
50
60
Disk Width (cm)
5000 Females: W eight(g) = 0.0161 x Length(cm) 3.1552 (n = 109) Males: W eight(g) = 0.0314 x Length(cm) 2.9603 (n = 59)
Weight(g)
4000
3000
2000
1000
0 10
20
30
40
Disk Width (cm) - 152 -
50
60