THE DEPARTMENT OF PHYSIOLOGY AND PHARMACOLOGY
CONNECTIONS VOLUME 1 | ISSUE 1 | OCTOBER 2014
PHYSIOLOGY AND PHARMACOLOGY
IN THIS EDITION... 2.
CELEBRATING A 100 YEAR HISTORY
4.
ANNUAL RESEARCH DAY AND STEVENSON LECTURE
6.
MEET OUR NEW FACULTY MEMBERS
7.
THE MARILYN ROBINSON AWARD FOR EXCELLENCE IN TEACHING
8.
AWARD WINNING FACULTY, STAFF AND TRAINEES
9.
UPCOMING EVENTS AND RECAP
10. DEPARTMENT BY THE NUMBERS
MESSAGE FROM THE CHAIR R Welcome to the inaugural edition of the Department of Physiology and Pharmacology newsletter, Connections. This new electronic newsletter is designed to reach out to our current Department members, alumni, former colleagues and friends on an annual basis. The goal is to use this as a means to keep everyone abreast of changes in our Department and academic programs, new additions to our
faculty and staff, and to update people on our achievements and newsworthy events. 2014 is a special year for our Department as it the 100th anniversary of the creation of our original component Departments: the Department of Physiology and the Department of Pharmacology. Because of this, we have included a history of the Departments in this newsletter to highlight some of the important dates and events in our evolution. We are proud of our history and the role that we have played over the past century in the growth of Western and our School. We will include activities that celebrate our 100th anniversary during a number of our annual events this academic year. This newsletter includes a feature on our Annual Research Day and Stevenson Lecture, and new additions we have made to the speaker line up to recognize a former and a current PhD student. We are also excited to introduce you to our newest Department faculty members in this newsletter. We are
fortunate to have recruited some bright and energetic new faculty members over the past year that both build on our current areas of strength and bring new expertise. This newsletter also celebrates some of our student, staff and faculty member award winners. Because of space limitations, we are not able to include everyone who has been recognized over the past year for their achievements in this newsletter. We are indeed fortunate to have outstanding Department members that bring honor to both themselves and our programs. Finally, on a personal note, I would like to thank you all for your dedication and commitment to our academic environment, and especially to those that worked so hard to bring Connections to fruition.
R. JANE RYLETT, PhD, FCAHS Distinguished University Professor Chair, Physiology and Pharmacology Schulich Medicine & Dentistry
CELEBRATING A 100 YEAR HISTORY He was succeeded by Dr. R. Waud who directed the department until 1960. Dr. Charles W. Gowdey became Head of the Department in 1960 and retained that position until 1981. Under the guidance of Dr. Gowdey, the department entered an expansion phase with the focus being on neuropharmacology.
PHYSIOLOGY AND PHARMACOLOGY
This year the Department of Physiology and Pharmacology marks its 100th anniversary of the creation of the two legacy departments, the Department of Physiology and the Department of Pharmacology, which were originally separate. The department as a whole has a rich history dating back to the 1900s, when Western’s medical school, now known as the Schulich School of Medicine & Dentistry, entered a new era by establishing seven new departments. A BRIEF HISTORY: DEPARTMENT OF PHYSIOLOGY In 1914, Dr. F.R. Miller was appointed as the first full Professor of Physiology, and was involved in research, teaching, and initiating student laboratory instruction that included both human and animal experimentation. In 1947, Dr. Miller was appointed to the newly established position of Research Professor, and was briefly succeeded as Head by Professor O. E. Edholm. In 1951, Dr. J.A.F. Stevenson was appointed Professor and head of the Department of Physiology, and he served with great distinction for 19 years. During his tenure, the number of full-time faculty members increased, and teaching and research programs were reviewed and strengthened. The teaching commitment of Physiology was expanded to include students in the Faculties of Physical Education, Nursing, and Science. An Honours BSc in Physiology was also established with the first class graduating in 1969.
This focus was promoted in 1961 with recruitment of Dr. J.T. Hamilton as a lecturer and Richard B. Philp and Lewis W. Hersey as graduate students. The department began to diversify in 1980 and its name was changed to the Department of Pharmacology and Toxicology to reflect the growth of toxicological research and extension of its teaching mandate. In 2001, Dr. M.A. Cook took on the role of Acting Chair and guided the department through the steps leading to the merger with the Department of Physiology. A BRIEF HISTORY: DEPARTMENT OF PHYSIOLOGY AND PHARMACOLOGY The Department of Physiology and Pharmacology was formed on July 1, 2002 as a result of the merger of the Department of Physiology and the Department of Pharmacology and Toxicology. This coincided with the recruitment of Dr. J.L. Henry to Western University as Chair of the Department. With the departure of Dr. Henry in December 2004, Dr. R.J. Rylett was appointed and remains as the current Chair. In the past decade, the department has seen some major changes including updated state-of-the-art undergraduate student labs and the renaming of the graduate degree awarded by the department to read MSc or PhD in Physiology and Pharmacology. The department is currently comprised of more than 90 faculty members and cross-appointees, more than 115 graduate students and post- doctoral fellows, and more than 30 administrative and technical staff.
While several other Chairs served between the early 1970’s and the early 2000’s, Dr. S.M. Sims, who became Acting Chair in 2001 and 2002, played a key role in facilitating the merger with the Department of Pharmacology and Toxicology. A BRIEF HISTORY: DEPARTMENT OF PHARMACOLOGY The Department of Pharmacology was founded in 1914 with Dr. J.M. Crane serving as Head from 1915 to 1934.
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CONNECTIONS | THE DEPARTMENT OF PHYSIOLOGY AND PHARMACOLOGY
Departments established •
Prior to 1914
1914
• •
The Medical School entered a new era with the establishment of seven departments, including Physiology and Pharmacology
There were no individual departments Teachers were part-time and professors were elected to teach individual subjects
1921
The Medical School moves
•
Dr. F. R. Miller was appointed as the Head of Physiology (top photo)
•
A new building on Ottaway Avenue (South Street) opened
•
Dr. J. M. Crane was appointed as the Head of Pharmacology (bottom photo)
•
It served the Faculty of Medicine for 44 years
New Head of Pharmacology •
1934
Dr. R.A. Waud
Celebrating 100 Years
Physiology and Pharmacology
New Head of Physiology 1947 •
Dr. O. G. Edholm
1951
New Head of Physiology •
New Head of Pharmacology •
1960
Dr. C. W. Gowdey
Dental Sciences Building opens New Chair of Physiology •
1977
New Chair of Physiology •
1980
1981
1984
1986
1992
Dr. R. B. Philp
New Chair of Pharmacology •
Dr. J. Bend
Department of Physiology and Pharmacology forms
2004
2002
Dr. R. J. Rylett
New Chair of Physiology and Pharmacology •
New undergraduate labs open for use
Dr. G. J. Mogensen
New Chair of Pharmacology •
Dr. R. L. Kline
New Chair of Physiology and Pharmacology •
Transition from “Head” to “Chair” of Department
Dr. J. Kraicer
New Chair of Physiology •
1968
Medical Sciences Building on Western campus complete
1972
Pharmacology changes to Pharmacology and Toxicology
•
1967
1965
Dr. V. B. Brooks
New Chair of Physiology
Dr. J. A. F. Stevenson
Dr. J. Henry
2012
2014
100 year anniversary
PHYSIOLOGY AND PHARMACOLOGY
THE STEVENSON LECTURE AND RESEARCH DAY In 1972, a lecture series was created to honour the memory of Professor James A.F. Stevenson (1918-1971) who served as Chair of the Department of Physiology from 1951 to 1970. Under his dynamic leadership, the Department was expanded and teaching and research programs were strengthened. He directed a vigorous research group in Regulatory and Integrative Physiology concerned in particular with neural mechanisms in the control of water and energy balance.
of the Association of Scientific, Engineering and Technological Communities of Canada, and Editor of the Canadian Journal of Physiology and Pharmacology. He received national and international recognition for his contributions, including election as a Fellow of the Royal Society of Canada in 1969. Professor Stevenson believed that physiological processes are integrated components of adaptive responses of the whole animal to environmental challenges. This was a belief he firmly held as teacher and scientist. The Stevenson Lecture is dedicated to Professor Stevenson and his commitment and contributions to the Department.
Professor Stevenson was a pioneer in organizing scientific activities as founder and then as president of the Canadian Physiological Society and of the Biological Council of Canada. He also served as the Vice President
THE 2014 JAMES A.F. STEVENSON DISTINGUISHED LECTURER: DR. HARRY C. DIETZ III The 2014 Stevenson Lecture will take place as part of the Department’s annual Research Day. It will feature Dr. Harry C. Dietz III. Dr. Dietz is the Victor A. McKusick Professor of Pediatrics, Medicine, and Molecular Biology & Genetics in the Institute of Genetic Medicine at the Johns Hopkins University School of Medicine. He is also an Investigator in the Howard Hughes Medical Institute. His undergraduate training in biomedical engineering was performed at Duke University and his MD degree was received from the Health Sciences University of Syracuse. Clinical and research training in pediatrics, pediatric cardiology, and genetics occurred at Johns Hopkins University School of Medicine.
Dr. Dietz heads a multidisciplinary clinic for the diagnosis and management of individuals with heritable forms of cardiovascular disease, with a special emphasis on Marfan syndrome and related connective tissue disorders. He is Director of the William S. Smilow Center for Marfan Research, a group of dedicated molecular biologists focused on improvement of the lives of individuals with Marfan syndrome and related disorders through the development of novel diagnostic and treatment strategies. Dr. Dietz has received multiple prestigious awards including the Curt Stern Award from the American Society of Human Genetics, the Taubman Prize for excellence in translational medical science, and the Harrington Prize from the American Society for Clinical Investigation and the Harrington Discovery Institute. He is an inductee of the American Society for Clinical Investigation, American Association for the Advancement of Science, Academy of American Physicians, Institute of Medicine, Association of American Physicians, and the National Academy of Sciences.
JOIN US FOR THE JAMES A.F. STEVENSON DISTINGUISHED LECTURE AND RESEARCH DAY TUESDAY, NOVEMBER 4, 2014
SPEAKERS:
10:30 A.M. STUDENT POSTER PRESENTATIONS Great Hall, Somerville House
Michael Pest, PhD Candidate: “Cartilage specific deletion of Mig6 disrupts mouse joint homeostasis”
2:00 P.M. RESEARCH TALKS London Health Sciences Centre, University Hospital, Auditorium A
Cory Yamashita, MD, FRCPC: “Host-Defense Peptides for the Treatment of Cystic Fibrosis”
4:00 P.M. JAMES A.F. STEVENSON DISTINGUISHED LECTURER London Health Sciences Centre, University Hospital, Auditorium A
Jim Petrik, PhD: “The Paradox of Anti-Angiogenic Therapy in Cancer" Harry C. Dietz III, MD: “Found in Translation: New Insights into the Pathogenesis and Treatment of Marfan Syndrome and Related Disorders”
STEVENSON LECTURE AND RESEARCH DAY: SPEAKERS Learn more about our Annual Research Day Speakers:
MICHAEL PEST, PhD CANDIDATE Michael Pest is a senior PhD candidate in Dr. Frank Beier’s lab. His research focuses on the role of epidermal growth factor receptor (EGFR) signaling in cartilage health and joint pathology. He is investigating the role of EGFR regulatory proteins such as mitogen inducible gene 6 (Mig6) in musculoskeletal diseases like osteoarthritis. Pest has been funded by a variety of sources during his studies including the Canadian Arthritis Network, the collaborative CIHR STIHR Joint Motion Program, and most recently he was awarded the Doctoral Banting and Best Canada Graduate Scholarship.
CORY YAMASHITA, MD, FRCPC Cory Yamashita is a clinician-scientist in the Department of Medicine and Division of Respirology. Yamashita received his MD from the University of Manitoba in 2001 and pursued his training in Internal Medicine and fellowship training in Adult Respirology at Schulich Medicine & Dentistry. After completing his clinical training, he completed a research fellowship under the direction of Dr. Jim Lewis and Dr. Ruud Veldhuizen at Schulich Medicine & Dentistry in the area of surfactant biology and acute lung injury. Yamashita joined Schulich Medicine & Dentistry, Western University in 2007 and is an active member of the Department of Medicine and is cross appointed to the Department of Physiology and Pharmacology. His research program is centered around host-defense peptides for the treatment of bacterial lung infections and acute lung injury. His research is currently supported by the Ontario Thoracic Society and the Physicians’ Services Incorporated Foundation.
JIM PETRIK, PhD Jim Petrik graduated from the Kinesiology program at Western University with an MSc in 1995. He carried out the research for his PhD degree in Physiology at Schulich Medicine & Dentistry with Dr. David Hill on the in utero regulation of organogenesis and metabolism. A postdoctoral fellowship at the University of Guelph followed. He is currently a full professor in the Department of Biomedical Sciences, and has a cross-appointment in the Faculty of Medicine at McMaster University. His research interests include the evaluation of angiogenesis and anti-angiogenic therapies in epithelial ovarian cancer as well as the roles of metabolism and inflammation in the onset and progression of ovarian cancer. His research is supported by grant funding from NSERC, CIHR, the Ontario Institute for Cancer Research, and Ovarian Cancer Canada, as well as infrastructure grants from the Canadian Foundation for Innovation.
SCHULICH SCHOOL OF MEDICINE & DENTISTRY
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MEET OUR NEW FACULTY MEMBERS DR. WATARU INOUE Originally from Japan, Dr. Wataru Inoue moved to Canada to pursue a PhD at McGill University. Prior to moving to London, Ontario, he conducted postdoctoral training at the University of Calgary. Dr. Inoue’s research focuses on synaptic plasticity caused by stress as an entry point to understand how we adapt or fail to adapt to stress. By using patch clamp electrophysiology and optogenetics, he studies synapses in the hypothalamus. Disregulation of the hypothalamic-pituitary-adrenal axis is closely linked to various stress-related disorders, including major depression.
DR. JAMIE KRAMER Dr. Jamie Kramer carried out the research for his PhD degree at Memorial University of Newfoundland with Dr. Brian Staveley, where he became excited about the use of D. Melanogaster as a model for human disease. He was then recruited to Radboud University in the Netherlands where he established a new fly facility for disease modeling in the Department of Human Genetics. There he studied the neuronal function of chromatin regulators that were found to be disrupted in intellectual disability. Dr. Kramer’s research led him toward his current research interest in understanding epigenetic dysregulation to human cognitive disorders. He explains that this is his dream job — he knew he wanted to be a scientist from a young age, as he always loved trying to figure things out.
DR. PETER STATHOPULOS Dr. Peter Stathopulos is a Western University alumnus who completed his MSc under the supervision of Dr. Qingping Feng on studies of L-arginine uptake during heart failure. He pursued his PhD at the University of Waterloo, investigating how protein misfolding is involved in motor neuron disease, and received postdoctoral training on the structural biology of calcium handling proteins at the Ontario Cancer Institute. Dr. Stathopulos plans to focus his research program on the molecular mechanisms of organelle-specific calcium signaling protein function in health and disease. Understanding precisely how proteins mediate signaling cascades at the atomic level opens up exciting possibilities for developing novel diagnostic tools and pharmacological regulators of these pathways.
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CONNECTIONS | THE DEPARTMENT OF PHYSIOLOGY AND PHARMACOLOGY
THE MARILYN ROBINSON AWARD FOR EXCELLENCE IN TEACHING The Department of Physiology was fortunate to have Dr. Marilyn Robinson as a faculty member for many years during the 1980’s and 90’s. Dr. Robinson was an exceptionally talented educator and an enthusiastic lecturer who was valued and respected by both colleagues and students. As an Assistant Professor in Physiology and Coordinator of the Educational Development Office, she raised the profile of teaching at Western. Dr. Robinson was also a leader in developing new educational approaches to engage students in active learning and problem solving. Her expertise was recognized with many teaching awards, including the 3M Teaching Fellowship and the Pleva Excellence in
Teaching Award. Following her untimely passing, Western established The Marilyn Robinson Award for Excellence in Teaching, awarded for the first time in the 1996-97 academic year. This Award is given annually to a junior faculty member who demonstrates excellence in teaching based on outstanding contributions in the area of classroom, laboratory, or clinical instruction. The Department of Physiology and Pharmacology is fortunate to have several early-career faculty members following in Dr. Robinson’s footsteps as outstanding educators. Three of our faculty members have won this prestigious award: Tom Stavraky in 2001-02, Dr. Anita Woods in 2012-13, and Dr. Brad Urquhart in 2013-14.
AWARD WINNING TEACHERS
TOM STAVRAKY Tom Stavraky is an outstanding classroom teacher and manager of the third-year physiology laboratories, and has consistently received high ratings for his teaching. Stavraky is renowned for his in-class demonstrations of physiological principles. Following his success as a recipient of the Marilyn Robinson Award, he was awarded the University Students’ Council Award of Excellence, 2005 Schulich Undergraduate Education Leader Award, a Fellowship in Teaching from Western’s Teaching Support Centre, and the prestigious Leadership in Faculty Teaching Award given by the Ontario Ministry of Training, Colleges and Universities.
ANITA WOODS Anita Woods began her appointment in Physiology and Pharmacology in 2008, and has demonstrated a unique flare for teaching and education scholarship. She indicates that she is motivated to see light-bulb moments on students’ faces when they understand a new concept. Woods excels at using new technologies in her teaching including: real-time Twitter in lecture; both pre-lecture and review podcasts; experiential lectures; and augmented reality laboratory aids. Her contributions to Western go far beyond the classroom, with research into course evaluation and outcomes, and serving in mentorship and advisory roles.
BRAD URQUHART Brad Urquhart has been a professor in Physiology and Pharmacology since 2009, and has shown himself to be a superlative well-rounded teacher. Soon after he began his faculty appointment, Urquhart saw the need for a major course revision in the large online second-year Pharmacology course. This very popular course now incorporates many innovative teaching activities including animations and videos, as well as CSI scenarios in which students are walked through virtual crime scenes to study the pharmacological aspects of the crime. Students praise his teaching style, especially the methods he uses to engage students.
SCHULICH SCHOOL OF MEDICINE & DENTISTRY
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CONGRATULATIONS TO OUR TRAINEES RECIPIENTS OF NSERC OR CIHR MASTER’S SCHOLARSHIPS
RECIPIENTS OF CIHR DOCTORAL AWARDS
Ian Tobias (NSERC — supervisor Dean Betts) “Redox regulation of canine pluripotent stem cells”
Jake Bedore (supervisors Cheryle Seguin & Andrew Leask) “CCN proteins as emerging therapeutic targets for the treatment of degenerative disc disease”
Matthew Tsang (CIHR — supervisor Andrew Leask) “Role of skin progenitor cells in dermal fibrosis and wound healing” Chang-Hui Wang (CIHR — supervisor Peter Chidiac) “Role of regulator of G protein signaling 2 (RGS2) in cellular stress and apoptosis” Martin Woo (CIHR — supervisor Doug Fraser) “Alteration of human brain pericyte contractility induced by traumatic brain injury” RECIPIENT OF CIHR POSTDOCTORAL FELLOWSHIP Dr. Daniel Stolzberg (supervisor Stephen Lomber) “Cross-modal cortical plasticity following implantation of an artificial prosthesis”
Maha Hammad (supervisor Stephen Ferguson) “The role of PDZ proteins in the regulation of the signaling and trafficking of the corticotropin releasing factor 1 and serotonin 2A receptors” 2013 GRADUATE STUDENT RESEARCH AND TEACHING AWARDS IN PHYSIOLOGY AND PHARMACOLOGY Adrian Gunaratne - PhysPharm Graduate Student Council Award for Leadership (supervisor John Di Guglielmo)
Jason Moreau and Ashbeel Roy (co-winners) — The Gordon J. Mogenson Scholarship recognizes outstanding research achievements of a continuing PhD student in the Physiology Graduate Program (supervisors John Ciriello and Marco Prado, respectively) Dr. Gurjeev Sohi — The Esme Walker Award, given annually to an exceptional Physiology PhD graduate (supervisor Dan Hardy) Tom Velenosi — The George W. Stavraky Teaching Scholarship recognizes a continuing graduate student in Physiology and Pharmacology Graduate Programs for outstanding ability in undergraduate teaching (supervisor Brad Urquhart)
Dr. Michael Knauer — The Margaret Moffat Award, given annually to an exceptional Pharmacology PhD graduate (supervisor Rommel Tirona)
Tom Velenosi — The Hari and Gudrun Sharma Award recognizes excellence in research and academic performance of a continuing student in the Pharmacology & Toxicology Graduate Program (supervisor Brad Urquhart)
Noelle Ma — The Murray Fraser Award, given annually to an exceptional Pharmacology MSc graduate (supervisor Dan Hardy)
Peter Vo — The Bill Bottom Award, given annually to an exceptional Physiology MSc graduate (supervisor Dan Hardy)
AWARD WINNING FACULTY AND STAFF 2014 AWARDS OF EXCELLENCE AND SCHULICH EXCELLENCE IN EDUCATION AWARDS
Jamie Simek & Chris Webb — Dean’s Award for Excellence for Staff (Team)
Dr. Mandar Jog — Named a 2014 Faculty Scholar by Western University
Dr. Lina Dagnino — Dean’s Award for Excellence for Faculty (Individual)
Dr. Bradley Urquhart — 2014 Marilyn Robinson Award for Excellence in Teaching
Dr. Michael Rieder — Named a 2014 Distinguished University Professor by Western in recognition of his sustained excellence in scholarship
Dr. Cheryle Seguin — Dean’s Award for Excellence for Faculty (Junior) Dr. Rommel Tirona — Schulich Educator Award for Undergraduate Education
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Dr. Cheryle Seguin — CIHR New Investigator Award for her research program entitled “Mouse Models to Study Intervertebral Disc Development, Health and Disease”
Dr. Frank Beier — Awarded the Canada Research Chair Tier I in Musculoskeletal Research
CONNECTIONS | THE DEPARTMENT OF PHYSIOLOGY AND PHARMACOLOGY
EVENTS RECAP AND EVENTS BY THE NUMBERS
ANNUAL HOLIDAY PARTY The Department’s Annual Holiday Party was held at The Wave in early January to accommodate our undergraduate students who were studying for exams during the holiday season. Everyone enjoyed dinner and the annual “Battle-of-the-Wits” game. This year, faculty and staff competed with trainees in an Olympic-themed “Minute-to- Win-It” game. In a tight battle, the team of faculty and staff members beat the trainees for the second straight year!
ANNUAL DEPARTMENT FAMILY BARBEQUE AND GOLF TOURNAMENT On June 13, golfers of all abilities met at Llyndinshire Golf & Country Club. Everyone had a great time and the “RAB CITY” team were the winners. There was also a great turnout for the barbeque dinner at Weldon Park following the tournament. Peter Stathopulos and Michael Knauer cooked up burgers and hotdogs while attendees and their families played backyard games like bocce ball and frisbee!
ANNUAL DEPARTMENT CORN ROAST Our hallmark event, the Annual Department Corn Roast, was held at Fanshawe Conservation Area on September 4 to welcome our new Physiology and Pharmacology 4980 and graduate students. The event was a great success with more than 200 attendees! As is tradition, the students helped by husking corn and were required to read the “Ancient Fourth-Year Oath” in front of the Department. Great barbequed food and extracurricular games were enjoyed by all.
500
+
PANCAKES SERVED FOR UNITED WAY
-4
(UNDER PAR) the winning team’s score at this year’s spring BBQ & golf day
UPCOMING SEMINAR SERIES Lectures will be held at 4:00 p.m. in the Dental Sciences Building, Room 2016 November 10, 2014 Michael Pest, PhD Candidate “EGFR Signalling in Osteoarthritis and Cartilage Homeostasis” November 17, 2014 Amira Klip, PhD “Communication between Muscle and Immune Cells Leading to Insulin Resistance” November 24, 2014 Sigolene Meilhac, PhD “The Cell Orchestra of Heart Morphogenesis” December 1, 2014 Christian Casanova, PhD “What can the Extrageniculate Thalamus Tell us about the Visual Cortex?” December 8, 2014 Carmen Leung, PhD Candidate “Role of Rac1 in Cardiomyocyte Polarity and Embryonic Heart Development” For more information visit: www.schulich.uwo.ca/ physpharm/about_us
160 BURGERS FLIPPED at this year’s Annual Department Corn Roast
SCHULICH SCHOOL OF MEDICINE & DENTISTRY
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$7,000,000+
4,624
STUDENT REGISTRANTS in undergraduate courses for the 2014-15 academic year DEPARTMENT BY THE NUMBERS
101
graduate students registered in our MSc and PhD programs in September 2014
77
full-time and cross-appointed faculty
department faculty member research grant total
226
RESEARCH PAPERS published in 2013
750+
34
number of MSc and PhD graduates since 1950
undergraduate courses offered in the 2014-15 academic year
number of fourth-year honours students:
THE DEPARTMENT OF PHYSIOLOGY AND PHARMACOLOGY
CONNECTIONS Department of Physiology and Pharmacology Schulich School of Medicine & Dentistry Western University Medical Sciences Building, Room 216 London, Ontario, Canada, N6A 5C1 t. 519.661.3460 f. 519.661.3827 www.schulich.uwo.ca/physpharm
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