(1928–1981), founded the School of Biology at the Universidad de San Carlos of Guatemala as well as the Biotopo Universitario para la Conservación del Quetzal, which is inhabited by this new species of salamander. He later became Rector of the Universidad de San Carlos of Guatemala. Juan Mario Dary Fuentes, his son, has followed strongly in his father’s path, rising to become the Guatemalan Minister of Environment and Natural Resources. Most of these slamanders were encountered on the ground, but 2 individuals were found in rotting vertical trunks about 2 m above the ground. Bolitoglossa tzultacaj is known only from the southern slopes of the Sierra de las Minas near the junction of the departments of Zacapa, Izabal, and Alta Verapaz. The forest at 1800 m is an admixture of pine-oak forest and cloud forest vegetation, which prevails at higher elevations. Bolitoglossa la (Fig. 8) is known from near Chichicastenango, the northern edge of the Guatemalan Plateau, and the extreme western portion of the Sierra de Chuacús. Localities are drained by the upper tributaries of the Río Chixoy and are separated from each other by less than 20 km airline. The elevational range is 2100–2390 m in pine-oak forest. In the vicinity of Chichicastenango, the species has been found most frequently in bromeliads up to heights of 6–7 m above the ground. A small series of specimens taken near Santa Rosa Chucuyub were encountered in or beneath rotting logs following rainy periods. Bolitoglossa nussbaumi is known only from the type-locality near Todos Santos in the Sierra de los Cuchumatanes. The specific epithet is a patronym for Ronald A. Nussbaum, the leading authority on caecilians, who joined us on several trips to Guatemala and was tireless in his pursuit of salamanders. Individuals of the type-series were collected from within and beneath rotted wood in a wooded ravine of a relatively dry forest containing pines, scattered oaks, laurels, and abundant brushy vegetation along open edges. Bolitoglossa centenorum is known only from the type-locality near San Mateo Ixtatán in the Sierra de los Cuchumatanes. The specific epithet is a patronym in honor of the Centeno family of Guatemala. Members of this family have done much to further education and appreciation of nature. Ing. Héctor A. Centeno, Rector emeritus (1992–2001) of the Universidad del Valle de Guatemala, has served in many positions, including Vicepresidential Advisor for Science and Technology and founder of the environmental and conservation organization Fundación Defensores de la Naturaleza. His son, Lic. Marco Vinicio Centeno, is founder of the Sociedad Guatemalteca de Ornitología, is a member of the Mesa Nacional de Aviturismo, and teaches biology at the Universidad del Valle de Guatemala. Individuals of the type-series were collected from beneath rotting logs. Bolitoglossa ninadormida (Fig. 9) is known from several highland localities in the Sierra de los Cuchumatanes, including near Todos Santos and near San Juan Ixcoy. This species has been collected in rotting trunks in a fir (Abies) and cypress (Cupressus) forest at 1600–1800 hr. Bolitoglossa huehuetenanguensis (Fig. 10) occurs at near San Mateo Ixtatán in the Sierra de los Cuchumatanes at elevations of 2450 to 2800 m. The type-series was collected mostly from
within and beneath rotted wood and a few individuals were taken under rocks. Most of the trees had been felled in the area and salamanders were also encountered under the loose bark at the base of stumps. Bolitoglossa psephena is known only from the highlands often referred to as the María Tecúm or the Tecún Umán Ridge. It inhabits montane wet forest that has abundant pines, cypress, firs, oaks, alders, and laurels. The type was taken at about 2500 m but the highest elevation attained in the area is the crest of the Tecún Umán Ridge at 3400 m. Relatively good patches of forest are still present. Bolitoglossa xibalba (Fig. 11) is relatively widespread, occurring at various localities in the Sierra de los Cuchumatanes and the Montañas del Cuilco. The elevational range is 1980–2760 m in wet montane forest. Most specimens were found under bark or moss at the base of rotting tree trunks. Individuals from the Montañas de Cuilco were taken between 19:30–21:00 hrs 50–120 cm above the surface of the ground on leaves along the side of a trail. Bolitoglossa nympha (Fig. 12) is a species belonging to a group of diminuitive, short-tailed salamanders including Bolitoglossa rufescens and Bolitoglossa occidentalis. It occurs in the Merendón region along the Guatemala-Honduras border at moderate elevations of 200 to 1200 m. Most specimens were found after dark sitting on leaves of low vegetation. The majority of individuals were taken less than a meter from the ground, but a few were encountered 1.6–2.0 m high. Nights following afternoon showers were especially productive in finding these salamanders, although they were also active during light night rains or mists. Although they might be found practically anywhere in the forest, they appeared to be more abundant in the humid environments presented by streamside vegetation. A new species of Dendrotriton is described from the mixed conifer-broadleaf forest of the northern Sierra de los Cuchumatanes and another from the cloud forest of the eastern extension of the Cuchumatanes. Dendrotriton chujorum (Fig. 13) was found along the road from Nentón to San Mateo Ixtatán in the northern portion of the Sierra de los Cuchumatanes. The slopes from where it was collected are drained by the upper tributaries of the Río Grijalva and are covered by small remnants of a mostly hardwood forest that is conspicuously drier than the forest at higher elevations or on opposing slopes of this range. All specimens were taken from bromeliads 3–10 m above the ground. Dendrotriton kekchiorum (Fig. 14) was taken in the mountains to the north of Uspantán, in the eastern portion of the Sierra de los Cuchumatanes. The holotype was collected on a rainy evening at about 1900 h at an air temperature of 12˚C. The specimen was sitting about 1 m above the ground on a fern along a small forest path. In addition to the new forms described in Campbell et al. (2010), we are aware of many undescribed species of Guatemalan salamanders that will be the subject of future studies. Given this knowledge, we suspect that salamander diversity in Nuclear Central America is still greatly underestimated. FrogLog Vol. 96 | May 2011 | 23