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Annual Report 2015


2 —  3


Director’s review


Looking towards a new phase in research and doctoral training


Focus on northern change and sustainability

12 – 13

Platform for co-operation

14 – 15

Main events in 2015

18 – 19

Scientific publications of the research groups

Annual Report 2015

Director’s review The Thule Institute promotes multidiscipli-

We organized the Thule Science Days

nary research and innovations in the fields of

under the topic ‘Research on the Change

environment, natural resources and north-

in the North’, where the highlights of our

ern issues at the University of Oulu through

research programme were presented. A

specific research and doctoral programmes.

great achievement for us was to arrange the

Since 2012, under the auspices of the Thule

16th International Congress on Circumpo-

Institute’s board, we have funded 13 inter-

lar Health in Oulu and in Rokua. The con-

nationally evaluated research groups and

gress and the associated summer school for

six associated research groups. These groups

PhD students focused on human health and

have formed a sound multidisciplinary core

well-being in the Arctic and northern areas.

of the Thule Institute’s research and doctoral

Besides scientific meetings, our personnel

training. The year 2015 was one of excellent

widely contributed to the Third International

results as researchers published 140 scien-

Conference on Arctic Research Planning

tific articles and 14 PhD students defended

(ICARPIII) during the Arctic Science Sum-

their theses. The Aurora doctoral programme

mit Week in Toyama, Japan. This conference

includes 80 registered PhD students, of

was very important, as its outcome will be a

whom 21 have been employed in our research

roadmap for integrated Arctic research and

groups using the strategic funds that the

its priorities for the next decade.

University has directed to our focus area. Thule Institute’s board members, chaired by

The year 2015 was one of planning organ-

Professor Eero Helle are warmly thanked for

izational changes at the University of Oulu,

their efforts and input during their four year

which concern the Thule Institute, too. The

(2012 – 2015) acting period connected to the

Institute will hold a position as a strategic

Institute’s research programme and decision

research unit of the University, and all its

making related to operations.

research will be carried out in the groups in the faculties or schools. We will also con-

One of our goals has been to promote the

tinue to promote our University’s expertise in

tenure track model for research personnel

Arctic research and polar science issues. The

according to the University’s strategy. Also,

new strategy and the organizational changes

cooperation between state research institutes

at the University will be fully implemented

and the University of Oulu research com-

in 2016. As the strategy states, our heritage

munity has deepened with the assistance

as one of the northernmost multidiscipli-

of the Thule Institute. Two state research

nary universities in the world is invaluable

institutes have currently settled in the cam-

when mankind’s new growth frontier is to

pus of the University of Oulu as neighbors to

the north. In accordance with the strategy,

the Thule Institute. The University of Oulu

we will be calling for new, top-level inter-

and the Thule Institute have a remarkable

national research projects during 2016 for a

role in the University of the Arctic’s activi-

four-year period starting at the beginning of

ties while hosting the Thematic Networks

the year 2017.

and Research Liaison Offices. The University of the Arctic Research Liaison Office was launched at the beginning of 2015.

4 —  5

Professor Kari Strand Director of the Thule Institute

Annual Report 2015

Dark-red Helleborine (Epipactis atrorubens) is one of the study species in a project, which combines molecular ecology with long-term life-history and demographic data to increase knowledge about survival and recoveries from population crashes. Photo by Hilde Hens.

Looking towards a new phase in research and doctoral training The Thule Institute has been focus-

6 —  7

The new University of Oulu strat-

ing for a long time on multidisciplinary

egy for 2016–2020 is based on five

research and doctoral training related to

thematic, internationally significant

the northern and Arctic environment,

research focus areas: ‘Creating sustain-

natural resources and sustainability. Due

ability through materials and systems’,

to the new organizational arrangements

‘Molecular and environmental basis

of the University of Oulu, 2015 was a

of lifelong health’, ‘Digital solutions in

year of reorganization for the Thule

sensing and interactions’, ‘Earth and

Institute, too.

near-space system and environmental change’, and ‘Understanding humans

Until the end of 2015, the Thule

in change’. Tackling those identified

Institute acted as a research centre with

global challenges requires multidiscipli-

an established research personnel. The

nary approaches and top-level research.

basis of its research consists of a multi-

From the beginning of 2016, the Thule

disciplinary research programme with

Institute acts as a strategic research

a total of 19 research groups that have

coordination unit of the University. The

all passed strict scientific evaluation

Institute represents two of those new

criteria. The groups are led by professors

thematic focus areas, ‘Creating sustain-

or associate professors at the University

ability through materials and systems’

of Oulu. The present groups will con-

and ‘Earth and near-space system and

tinue their work in the Thule Institute’s

environmental change’. A call and a

research programme until the end of

selection procedure for new, high-level

2016, and all the research personnel

research projects for the period 2017–

have their place in the faculties, accord-

2020 will be carried out in 2016.

ing to the University’s reorganization. The operational units of the Thule Insti-

Researcher and doctoral training

tute, the Centre for Arctic Medicine and

will continue in the Thule Institute’s

the Centre of Northern Environmental

research projects and in the Aurora DP,

Technology (NorTech Oulu), were inte-

a Doctoral Programme in Environment,

grated into the faculties of the Univer-

Society and Technology in the North.

sity. The Oulanka Research Station is

This solid cooperation will ensure high-

a part of the infrastructure unit of the

level and effective interaction at all lev-

University. The Innovation Centre for

els of high-grade academic research. The

Environment and Energy (CEE), which

Thule Institute also continues to pro-

was also located at the Thule Institute,

mote the University’s cooperation with

moved into the newly set up innovation

Finnish national research institutes and

arena at the University campus.

international science organizations. Annual Report 2015

Thule Institute in numbers in 2015


Research groups operated in the faculties of the University of Oulu and in the Thule Institute


Works published in scientific publications


Doctoral dissertations


International science events arranged


Coordination tasks carried out


Representatives in the staff in scientific organizations and other co-operative bodies


Students in the doctoral programme


Members of staff

Focus on northern change and sustainability In the programme period 2012–2016,

gathered wide interest with the topi-

the multidisciplinary research groups

cal contributions of the researchers and

and the doctoral programme in Environ-

doctoral students of the University and

ment, Society and Technology in

the state research institutes. The doc-

the North (Aurora DP) constitute a plat-

toral training events organized by the

form for three main research themes:

Aurora DP included training courses and

‘Climate Change Dynamics, Impacts

summer schools, in addition to informal

and Adaptation’, ‘Human Health and

joint student meetings. Fourteen PhD

Community Wellbeing’ and ‘Sustain-

students defended their thesis in 2015.

able Resources Management’. Aurora

The University of Oulu granted Aurora

DP, coordinated by the Thule Institute,

DP five new, four-year doctoral student

is one of the nine officially recognized

positions for 2016–2019.

8 —  9

doctoral programmes of the University of Oulu Graduate School. Doctoral

The research groups in the pro-

training in the Thule Institute’s research

gramme also carry out national and

programmes has been carried out since

international collaboration with other

2002. The period since 2012 has been

universities and state research institu-

fruitful in research and in doctoral

tions, especially utilizing the LYNET

training. By the end of 2015, a total of

coalition (the Finnish Partnership for

45 PhD students had defended their the-

Research on Natural Resources and the

ses. The 19 research groups financially

Environment). The new contract, which

supported by the Thule Institute have

includes the plan for research collabo-

been vitally important in understanding

ration and infrastructure development

the complexities of change in the North,

for coming years between the LYNET

and over 300 scientific articles have

organizations and the University of

been written during the programme

Oulu, was signed 1st October 2015. Pres-

period. The research contributions and

ently, there are over 40 ongoing projects,

doctoral training done within all these

which are coordinated and carried out

research groups are much appreciated.

by the professors and researchers at the Institute’s research programme. These

The main event in 2015 was ‘Thule

projects are mostly funded by the Acad-

Science Days – Research on the Change

emy of Finland, Tekes and the EU, and

in the North’ in October, where research

they form the framework for active col-

groups and PhD students presented their

laboration with the Thule Institute and

research. The Science Days included

its national and international partners.

three relevant sessions: People in the North, Climate Change and the Environment, and Sustainable Resources Management. Besides the scientific sessions, Career Planning’ was arranged for the graduating doctoral students. The event

Annual Report 2015

a half-day workshop entitled ‘Research

Research groups in the 2012–2016

• Evolutionary approach to trophic interactions under climate change (Markku Orell)

programme by multidisciplinary theme (group leader’s name in parentheses):

• Outer space influence on the low and middle polar atmosphere (Ilya Usoskin)

Climate Change Dynamics, Impacts and Adaptation • Unravelling population history, demography and structure for management of northern populations: Molecular ecology meets population biology (Laura Kvist) • Rapid environmental changes in the Eurasian Arctic – lessons from the past to the future (Juha-Pekka Lunkka, Kari Strand) • Consortium: Streams in the landscape: effects of land use on biodiversity and eco-

• Consortium: The effect of climate change on Scots pine seed and seedling development (SEED) (Hely Häggman),

Biodiversity and people in the land-use (Anne Tolvanen)

• Transdisciplinary environmental research (Timo P. Karjalainen) • Climate change and boreal rivers (Heikki Mykrä) • Sedimentary and geochemical research on climate and environmental change (Kari Strand)

system processes in boreal streams (Timo Muotka),

Human impact on Northern groundwater resources: Impact of land-use and climate change on future management of aquifers and connected ecosystems (Björn Klöve)

Human Health and Community Wellbeing • Circles of violence – Challenges on building a culture on non-violence in family and school (CIRCLE) (Vappu Sunnari) • Structural racism and its impact on Indigenous Health – a comparative study of Canada, Finland and Norway (Arja Rautio) • Prevention of marginalization in Northern Finland (Juha Veijola, Anja Taanila, Irma Moilanen) • Health and well-being (Arja Rautio)

Sustainable Resources Management • Potentiality of geodiversity information in biodiversity assessments (GEOBIO) (Jan Hjort) • Research Group on Environmental Catalysis: Sustainable abatement and use of gaseous emissions in process industry (Riitta Keiski) • Waste and energy synergy networks in the context of eco-municipalities – Towards resource efficiency in the North (WE2North) (Eva Pongrácz) • Utilization of chemical non-wood resources in local bioeconomies (Jouko Niinimäki) • Environmental technology (Jaakko Rämö) • Environmental and resource economics (Ilmo Mäenpää) Field activities in Kilpisjärvi related to the GEOBIO research project. Photo by Jan Hjort.

10 —  11

The course on Northern hydrology – research methods and foreseen changes took the participants in the field to monitor environmental conditions and how the collected data brought it all up. Photo by Anssi Rauhala.

Annual Report 2015

Visiting researcher Weng-Si Chao applies the electron microprobe at the Centre of Microscopy and Nanotechology, University of Oulu for a specific mineralogical study of the central Arctic Ocean sediments from the Lomonosov Ridge.

� Participation in international and national networks and coordination of several networking projects and secretariats.�

Field technician Markku Rontti from the Oulanka research station measures snow depth and snow weight at Oulanka in February for the Finnish Environment Institute.

Platform for co-operation The research area of the University of

stories of cutting-edge Arctic research

the Arctic (UArctic) is led by the Thule

from landscape-forming processes to the

Institute. This includes hosting the The-

peoples of the Arctic, as told by scientists

matic Networks and Research Liaison

supported by INTERACT Transnational

Office. In addition, the Thule Institute


12 —  13

led two thematic networks: Health and Well-being in the Arctic (Centre for Arc-

The Thule Institute has been hosting

tic Medicine) and Communicating Arc-

a national portal for IODP (the Interna-

tic Research (Oulanka Research Station).

tional Ocean Discovery Program) and

These networks conducted doctoral

ICDP (the International Continental

training and educational activities, such

Scientific Drilling Program), provid-

as PhD summer schools, in collaboration

ing general information about these

with the Aurora DP (connected to the

programmes and activities of Finland in

ICCH16), and the second workshop on

2015. The portal offers information and

Visualization of Environmental Change

contact details for Finnish scientists and

at Oulanka.

students applying to participate in the research expeditions and meetings. The

EU-PolarNet was launched at the

Academy of Finland supports Finland’s

beginning of 2015. The Thule Institute

participation in the IODP, managed in

is a Finnish partner in this EU H2020-

Europe by the European Consortium for

funded European consortium of exper-

Ocean Research Drilling (ECORD).

tise and infrastructure of polar research. The Thule Institute is leading a task on

Research co-operation between the

stakeholder consultation, the work for

University of Oulu and state research

which began by convening scientific

institutes within the Lynet (Finnish

sessions in Paris at ‘Our Common Future

Partnership for Research on Natu-

under Climate Change’, a preparatory

ral Resources and the Environment)

congress for the COP21 Climate Change

Northern Cooperation Programme has

Conference and at the Arctic Circle

been promoted by the Thule Institute.

Assembly in Reykjavik.

Lynet is a network of the seven largest research institutes in the sectors of natural resources and the environment

INTERACT (the International Network

in Finland. The Northern Cooperation

for Terrestrial Research and Monitoring

Programme 2016–2018 was planned

in the Arctic), where it has been man-

and agreed during 2015 on the basis

aging the Transnational Access work

of the actions and good practices from

package for the past five years. A call for

the former NorNet cooperation. The

grants to conduct research at two Cana-

Programme focuses on both research

dian and two U.S. sites was arranged in

project and infrastructure cooperation

2015, resulting in the selection of four

as well as capacity-building and doc-

European projects to visit these sta-

toral training. The main themes of the

tions. One of the main events of the year

research are ICT solutions for environ-

was the launch of INTERACT: Stories

ment and nature resources, the north-

of Arctic Science at the Arctic Circle

ern bioeconomy, ecosystem change and

conference in Reykjavik. This highly

interactions, and human wellbeing in

illustrated popular science book contains

the north.

Annual Report 2015

The Thule Institute is a partner in

� Fourteen doctoral theses with Arctic attitude from the Aurora DP in 2015.�

Main events in 2015 • Course on spatial multicriteria analy-

• Presentations by the Thule Institute

sis for environmental decision-mak-

researchers and the game I’m the

ing. 21–23 January. Oulu.

King of the World by NorTech Oulu in

• Seminar on biological and environmental applications of Bayesian inference. 26 February. Oulu. • Summer School of the 16th International Congress on Circumpolar Health: Focus on Future Health and Wellbeing. 6–8 June. Oulu. • 16th International Congress on Circumpolar Health. 6–12 June. Oulu and Rokua.

14 —  15

the University of Oulu’s Science Days. 1-2 September. Oulu • Thule Science Days 2015 – Research on the Change in the North Oulu. 27–28 October. Oulu. • Course on Northern Hydrology – research methods and foreseen changes. 1–3 December. Oulu. • Polarcode project’s stakeholder workshop. 9 December. Helsinki.

• Barents Summer School 2015 on cli-

• Enterprise Forum: Academic Entre-

mate change effects on Arctic land-

preneurship – Doctors Creating New

scapes: integrating responses across

Business. 10 December. Oulu

terrestrial and aquatic ecosystems. 21–26 June. Abisko, Sweden.

Annual Report 2015

The doctoral defence ceremony of Lic.Sc.(Tech.) Jenni Ylä-Mella was held on November 27th, 2015. On the photo: custos and supervisor Eva Pongrácz, defendant Jenni Ylä-Mella and opponent Professor Mika Horttanainen, Lappeenranta University of Technology.

The doctoral defence ceremony of Master of Health Sciences Anastasia Emelyanova was held on November 26 th, 2015. On the picture (from left): opponent, Professor Reijo Tilvis, University of Helsinki, custos and supervisor, Professor Arja Rautio and defendant Anastasia Emelyanova.

ICCH16 participants discussing lively of population health in the Arctic. Photo by Lauri Autere.

16 —  17

Gáktennieiddat performed in the opening ceremony of the ICCH16. Photo by Seija Leskelä.

Annual Report 2015

Vice Rector Olli Silvén (right) and Dean of the Faculty of Medicine Kyösti Oikarinen represented University of Oulu in the opening ceremony of the ICCH16.

Scientific publications of the research groups Doctoral theses Emelyanova Anastasia. Cross-regional

Ylä-Mella Jenni. Strengths and challenges in the Finnish waste electrical

analysis of population aging in the

and electronic equipment recovery


system : consumers’ perceptions and

Grönroos Mira. Metacommunity structuring in stream systems. Disentan-

participation. Zoratti Laura. Effect of environmental,

gling the roles of regional and local

developmental and genetic factors on


flavonoid and carotenoid profile of

Hekkala Anne-Maarit. Restoration of

Vaccinium berries.

the naturalness of boreal forests. Irannezhad Masoud. Spatio-temporal climate variability and snow resource changes in Finland.

Refereed articles in scientif ic journals

Kangas Veli-Matti. Genetic and Pheno-

Abass K, Carlsen A, Rautio A (2015).

typic Variation of the Moose (Alces

Approaches to describe risks and


future needs. Chapter 4 in AMAP

Karhu Mirjam. Treatment and characterisation of oily wastewaters.

Assessment 2015: Human Health in the Arctic, pp. 99-110.

Muilu-Mäkelä Riina. Polyamine metabo-

Abbaspour KCE, Rouholahnejad N,

lism of Scots pine under abiotic stress.

Vaghefi S, Srinivasan R, Kløve B

Nordström Tanja. Predisposing fac-

(2015). Modelling hydrology and

tors and consequences of adolescent

water quality of the European Con-

ADHD and DBD : a longitudinal study

tinent at a subbasin scale: calibra-

in the Northern Finland Birth Cohort

tion of a high-resolution large-scale


SWAT model. Journal of Hydrology

Prokkola Hanna. Biodegradation studies of recycled vegetable oils, sur-

524:733–752. Ala-aho P, Isokangas E, Rossi PM, Kløve

face-active agents, and condensing

B (2015). Fully integrated sur-


face-subsurface flow modelling of

Tirkkonen Tiina. Early attachment,

groundwater-lake interaction in an

mental well-being and development

esker aquifer: Model verification with

of Finnish children at preschool age.

stable isotopes and airborne ther-

Twinship - risk or opportunity?

mal imaging. Journal of Hydrology

Tolkkinen Mikko. Biodiversity and ecosystem functioning in boreal

552:301-406. Alahuhta J (2015). Geographic pat-

streams: The effects of anthropogenic

terns of lake macrophyte communi-

disturbances and naturally stressful

ties and species richness at regional


scale. Journal of Vegetation Science

Visanko Miikka. Functionalized nanocelluloses and their use in barrier and membrane thin films.

26:564-575. Alahuhta J, Rääpysjärvi J, Hellsten S, Kuoppala M, Aroviita J (2015). Species sorting drives variation of boreal lake and river macrophyte communities. Community Ecology 16:76-85.

Alakortes J, Fyrstén J, Carter AS, Moilanen

Ceresa F, Monros J, Belda E, Kvist L (2015).

IK, Ebeling HE (2015). Finnish mothers’

Does fragmentation of wetlands affect

and fathers’ reports of their boys and girls

gene flow in sympatric Acrocephalus

by using the Brief Infant-Toddler Social and

warblers with different migration strat-

Emotional Assessment (BITSEA). Infant

egies? Journal of Avian Biology. Doi:

Behav Dev. 2015 May; 39:136-47. Doi: 10.1016/j.inf beh.2015.02.016. Annala M, Mykrä H, Tolkkinen M, Virtanen

10.1111/jav.00589. de Bildt A, Sytema S, Zander E, Bölte S, Sturm H, Yirmiya N, Yaari M, Charman T,

R, Kauppila T, Muotka T (2015). Land use

Salomone E, LeCouteur A, Green J, Bedia

impacts on stream community composition

RC, Primo PG, van Daalen E, de Jonge

and concordance along a natural stress gra-

MV, Guðmundsdóttir E, Jóhannsdóttir S,

dient. Ecological Indicators 62:14-21.

Raleva M, Boskovska M, Rogé B, Baduel

Apperl B, Pulido-Velazquez M, Andreu J,

S, Moilanen I, Yliherva A, Buitelaar J,

Karjalainen TP (2015). Contribution of

Oosterling IJ (2015). Autism Diagnostic

the Multi Attribute Value Theory to con-

Interview-Revised (ADI-R) Algorithms

flict resolution in groundwater manage-

for Toddlers and Young Preschool-

ment. Application to the Mancha Oriental

ers: Application in a Non-US Sample of

groundwater system, Spain. Hydrology and

1,104 Children. J Autism Dev Disord.

Earth System Sciences. Sci., 19:1325-1337.

2015 Jul;45(7):2076-91. Doi: 10.1007/

Doi:10.5194/hess-19-1325-2015. Ardanov P, Lyastchenko S, Karppinen K, Häg-

s10803-015-2372-2. Emelyanova A, Rautio A (2015). Policies of

gman H, Kozyrovska N, Pirttilä AM (2015).

Arctic countries to promote the volun-

Variable effect of Methylobacterium sp. on

teering in old age. In book: Ageing, Well-

potato (Solanum tuberosum L) germina-

being and Climate Change in the Arctic:

tion, yield and disease development is associated with the shift in endophytic micro-

An Interdisciplinary Analysis, pp. 49-68. Eskelinen R, Ronkanen A-K, Marttila H,

bial community. Plant and Soil, 01/2015.

Kløve B (2015). Purification efficiency of a


peatland-based treatment wetland during

Beier P, Sutcliffe P, Hjort J, Faith DP, Pressey RL and Albuquerque F (2015). A review of selection-based tests of abiotic surrogates

snow-melt and runoff events. Ecological Engineering 84:169-179. Guo JY, Huhtaniska S, Miettunen J, Jääskel-

for species representation. Conservation

äinen E, Kiviniemi V, Nikkinen J, Moil-

Biology 29:668–679.

anen J, Haapea M, Mäki P, Jones PB,

Bode MK, Lindholm P, Kiviniemi V, Moil-

Veijola J, Isohanni M, Murray GK (2015).

anen I, Ebeling H, Veijola J, Miettunen J,

Longitudinal regional brain volume

Hurtig T, Nordström T, Starck T, Remes

loss in schizophrenia: Relationship to

J, Tervonen O, Nikkinen J (2015). DTI

antipsychotic medication and change in

abnormalities in adults with past history

social function. Schizophrenia Research

of attention deficit hyperactivity disorder: a tract-based spatial statistics study. Acta Radiologica 56:990-996. Bulik-Sullivan BK, Loh PR, Finucane HK,

168;1-2:297-304. Häggman H, Sutela S, Fladung M (2015). Genetic engineering – contribution to forest tree breeding efforts. In Vettori C, Fla-

Ripke S, Yang J, Schizophrenia Working

dung M, Häggman H, Pilate G, Gallardo

Group of the Psychiatric Genomics Con-

F, Kazana V, Migliacci F (Eds.) Biosafety

sortium, Patterson N, Daly MJ, Price AL,

of forest Transgenic Trees: Improving the

Neale BM (2015). LD Score regression dis-

Scientific Basis for Safe Tree Development

tinguishes confounding from polygenicity

and Implementation of EU Policy Direc-

in genome-wide association studies. Nat

tives. Springer, Forestry Sciences. Annual Report 2015

Genet. 47:291-295.

18 —  19

Hawkins CP, Mykrä H, Van der Laan J,

Huuki T, Renold E (2015). Crush: mapping

Oksanen J (2015). Environmental distur-

historical, material and affective force

bance can increase beta diversity of stream

relations in young children’s hetero-sexual

macroinvertebrate assemblages. Global

playground play. Discourse: Studies in the

Ecology and Biogeography 24:483-494.

Cultural Politics of Education, [Special

Heiderscheidt E, Leiviskä T, Kløve B (2015). Chemical treatment response to vari-

Issue: Fabulous monsters: alternative discourses of childhood in education].

ations in non-point pollution water


quality: Results of a factorial design

Huuki T, Sunnari V (2015). Standing by…

experiment. Journal of Environmental

standing off — troublesome compassion

Management 150:164-172. Doi:10.1016/j.

in the relationships of school boys. Gender

jenvman.2014.10.021. Heiderscheidt E, Leiviskä T, Ronkanen AK,

and Education 27;5:555-574. Irannezhad M, Kløve B (2015). Do atmos-

Kløve B (2015). Evaluating the suitability of

pheric teleconnection patterns explain

synthetic organic polymers to replace iron

variations and trends in thermal grow-

salts in the purification of humic and sed-

ing season parameters in Finland? Inter-

iment-rich runoff. Desalination and water

national Journal of Climatology. Doi:

treatment 57:10948-10957. Heikkinen HI, Sarkki S (2015). Ympäristön-

10.1002/joc.4332. Irannezhad M, Chen D, Kløve B (2015). Inter-

muutos, poronhoidon sopeutumiskapa-

annual variations and trends in surface

siteetti ja suurpedot. In: ”Suden kanssa”,

air temperature in Finland in relation to

Eds. Hiedanpää J & Ratamäki O. pp. 97 –

atmospheric circulation patterns, 1961-

119. Lapland University Press. Heino J, Alahuhta J (2015). Elements of regional beetle faunas: faunal variation and

2011. International Journal of Climatology. Doi:10.1002/joc.4193. Irannezhad M, Ronkanen A-K, Kløve B

compositional breakpoints along climate,

(2015). Effects of Climate Variability and

land cover and geographical gradients.

Change on Snowpack Hydrological Pro-

Journal of Animal Ecology 84:427-441.

cesses in Finland. Cold Regions Science

Heino J, Alahuhta J and Fattorini S (2015). Phylogenetic diversity of regional beetle

and Technology 118:14-29. Irannezhad M, Ronkanen A-K, Kløve B

faunas at high latitudes: patterns, drivers

(2015). Wintertime Climate Factors Con-

and chance along ecological gradients. Bio-

trolling Snow Resource Decline in Fin-

diversity and Conservation 2:2751-2767.

land? International Journal of Climatol-

Heino J, Soininen J, Alahuhta J, Lappalainen J, Virtanen R (2015). A comparative

ogy. Doi:10.1002/joc.4332. Irannezhad M, Torabi Haghighi A, Chen D,

analysis of metacommunity types in the

Kløve B (2015). Variability in dryness and

freshwater realm. Ecology and Evolution

wetness in central Finland and the role


of teleconnection patterns. Theoretical

Hjort J, Gordon JE, Gray M, Hunter JR ML (2015). Why geodiversity matters in valuing nature’s stage. Conservation Biology 29:630–639. Hjort J, Hugg TT, Antikainen H, Rusanen J,

and Applied Climatology. Doi: 10.1007/ s00704-014-1305-x. Jukuri T, Kiviniemi V, Nikkinen J, Miettunen J, Mäki P, Mukkala S, Koivukangas J, Nordström T, Parkkisenniemi J, Moilanen I,

Sofiev M, Kukkonen J, Jaakkola MS, Jaak-

Barnett JH, Jones PB, Murray GK, Veijola

kola JJK (2015). Fine-scale Exposure to

J (2015). Cerebellar activation in young

Allergenic Pollen in the Urban Environ-

people with familial risk for psychosis -

ment: Evaluation of Land Use Regression

the Oulu Brain and Mind Study. Schizo-

Approach. Environmental Health Perspec-

phrenia Research 2015.

tives. Doi: 10.1289/ehp.1509761.


Jukuri T, Kiviniemi V, Nikkinen J, Miettunen

Kaparulina E, Strand K and Lunkka JP

J, Mäki P, Mukkala S, Koivukangas J, Nor-

(2015). Provenance analysis of central

dström T, Parkkisenniemi J, Moilanen I,

Arctic Ocean sediments: Implications for

Barnett JH, Jones PB, Murray GK, Veijola J

circum-Arctic ice sheet dynamics and

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Editor: Pirjo Taskinen Layout: Hannele Heikkilä-Tuomaala / Communications and Marketing / University of Oulu Photos: University of Oulu / Thule Institute Printing house: Erweko Oy – Oulu 2016 ISBN: 978-952-62-1247-0

Thule annual report 2015  
Thule annual report 2015  

Annual report 2015 of the Thule Institute, University of Oulu, Finland.