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Ohio University

Botany NewS 1e tter 1 989

College ofArts and Sciences Athens, Ohio 45701



REPORT FROM THE CHAIR The department continues to prosper thanks to the efforts of faculty, students and office personnel, and the continued support of alumnae, alumni and friends. This year, Dr. Larry Larson announced his intention to seek early retirement. We will sorely miss Larry’s exceptional service to the department as a teacher and advisor of undergraduates and his whimsical good humor. He will continue to teach in the winter quarter, allowing us to be weaned gradually. Larry wifi be replaced—in terms of FTE, though not in style and discipline—by Jan Salick, a tropical botanist, who will enhance our traditionally strong ecology program and will add courses and research opportunities in tropical botany and conservation ecology. This year we were successful in competing for In-house monies, including a $61,000 award from House Bill 810 money to establish an equipment laboratory, and a $6,000 UPAC award to permanently increase our supplies budget. In last year’s newsletter we called upon you to support our botanical garden project. We were gratified by the response, which made possible physical landscaping of several sites, Including the establishment of a fern garden. As the physical appearance of the garden improves, the necessity to purchase plants of special botanical importance becomes a priority. Unfortunately, the additional monies mentioned above are earmarked for teaching and research, leading us again to seek your support for the botany garden. We hope that you will “stand up and be counted” by making at least a small gift, designated to the Department ofBotany, when Dean Eckelmann invites you to participate in this year’s College ofArts and Science Annual Roll Callfor Excellence.

.Wfai Ivan K. Smith Professor and Chair Torn Vierheller and Chenzhao Jian


Botany Newsletter 1989



The department has strengthened its program by the addition of a new faculty member in the area of tropical botany and conservation biology. We have selected a young, bright and established scientist, DR. JAN SALJCIC Jan is currently a Visiting Scientist In Ecology and Evolutionary Biology at Cornell University, on leave from her position as Assistant Scientist at the New York Botanical Garden. She wifijoin our faculty as an Assistant Professor this coming September.

This year, one of our most loved and respected faculty members has opted for early retirement. DR. LARRYLARSON has been a valued member of the department for the past 26 years, serving as both teacher, advisor and mentor, par excellence! Larry was honored June 2, at a retirement dinner attended by Dean Eckelmann and his wife, Jean, all current faculty and their spouses, and emeritus faculty Art Blickle and Warren Wlstendahl. He was presented with a wall plaque with the following Inscription:

Jan receIved her B.A. from Wisconsin, M.S. from Duke University and Ph.D. from Cornell University In 1983. Her current studies deal with the Interactions of plant populations and communities and human beings in both agricultural and tropical forest habitats. She states that “For conservation to succeed In a realm beyond the preservation of small reserved areas, human interactions with plant resources need to be understood. To this end, she has integrated her research on populations and genetics with conservation of habitats and germplasm. She has extensive experience In the tropics, Including Malaysia, Indonesia, Venezuela, Mexico, Central America, Colombia and Peru. She spent about four years in Central Selva, Peru, studying the ethnobotany of the Amuesha Indigenous people of the upper Amazon. These studies included population genetic studies of crop domestication, community analysis of agroforestry systems and regeneration and use of native tropical forest. She has studied the community ecology of rainforests, includ ing termite decomposition, In Malaysia; development of longterm conservation and resource management in Indonesia; ecology ofwild cassava species and plant? insect interactions on cassava in Mexico, Central America and Colombia. This past year at Cornell, she has worked on the genetics of peach tomato (SolanumsessilWorum) and has been involved with the development of Cornell’s conservation biology program, incorporating tropical field studies in ecology with conservation education. This summer, she wifi be teaching tropical conservation and applied ecology with the Organization for Tropical Studies in Costa Rica. “

Jan’s present work is supported by a $200,000 grant from the Andrew W. Mellon Foundation. She has received numerous other grants from The Agency for International Development, The Institute for Plant Resources Investigation, The International Board for Plant Genetic Resources and the National Science Foundation. She Is fluent in Spanish, French and Indonesian, as well as her native English.

“Presented to Laurence A. Larson in recognition of 26 years ofdedicated service, loyalty andfriendshp which will long be remembered. “

His guests at the dinner included his wife, Betty, daugh ters Amy and Betsy, Reid Huntley, Margaret Hill and Martha Allen. Larry has graciously submitted the following “life-history”, for his present and former students, wherever theymight be.

THE RETIREMENT OF LAURENCE A. LARSON The Beginning ofa Journey I came to Athens in the fall of 1953, havingjust been I discharged from the U.S. Army, to visit a boyhood friend who was then a student at the university. I liked what I saw, applied for admission to O.U. and was accepted as a physics major.

While working on the physics degree I enrolled in several botany courses taught by Professors Vermfflion, Buckle and Gambill and as a consequence changed my professional goal from physics to botany. I graduated in 1956, and took ajob with the U.S. Hydrographic Office and did field work in northern Alaska and the Arctic Ocean. While In Alaska I made a small

collection of mosses which was sent to the Bartley Herbarium at O.U.

An added bonus to the department and university is that Jan’s husband, DR. CHARLES STAVER, has extensive expertise and experience in tropical agroecology and farming systems. He Is a 1971 graduate of Columbia University (AB, Geography) with graduate degrees (MPS, Agronomy, and Ph.D., Vegetable Crops) from Cornell University.

At the Arctic Research lab, at Pt. Barrow, Dr.

Royal Shanks, of the University of Tennessee convinced me to return to school and to consider the botany gradu ate program at Tennessee. In March of 1957 I enrolled at the University ofTennessee, earned the M.S. degree in I959 and stayed on as instructor ofbotany for one term.


Botany Newsletter 1989 The teaching experience at Tennessee convinced me to continue my graduate training in preparation for a university teachingjob. I enrolled In the Ph.D. program at Purdue University and was awarded the degree in 1963. I returned to Ohio University in the summer of 1963 as the 7th member of the Botany Department. I developed courses In plant physiology and radiation biology techniques with the help of grants from the Atomic Energy Commission and National Science Foun dation. During this time I developed and co-authored with C. E. Mifier a laboratory manual for General Botany. Over the years I wrote several more laboratory manuals and Study Guides for Botany and Biology. A sabbatical leave in 1970-71 was spent at Oxford University in England with a friend whom I met while at Purdue. That year was yet another turning point in my professional development. On my return to Athens I concentrated on undergraduate teaching and course. development. During the 70’s I attended workshops which contributed to experimental course development. Some of the more Interesting workshops were: Ultimate Reality and Meaning; Freedom and Creativity In Educa tion and the Arts; Evolution/Creation; Ethics in Science; Human Sexuality; Small Group Facilitation; and Bioeth ics. These workshops led to several experimental courses such as: Science and Religion; Human Sexuality; Technology and Human Values; Reproductive Biology. A real breakthrough in course development for me came in the early 1980’s when the University devel oped a General Education requirement for all students. One part of the requirement was that all students must complete a Tier-Ill course, a senior synthesis course. I developed two courses; Unity and Variety in Biology and Literature and The Human Life Cycle. The development of the Human Life Cycle course required time and energy. It integrates biology, art and poetry with the stages of creation, transformation sexuality and death of the human cycle. Art and poetry as metaphor of the biological life cycle is a major component of this course. Some of the experiences I had in developing and maintaining this course led me Into new areas of activity and interest. It is risky business In academia to transfer one’s interest and activity from one area to another. It requires a kind of courage on the part of the individual and an open-minded philosophy on the part of the department. I’m pleased to say that we both succeeded and I believe are more aware of the world around us as a consequence. I attended art history seminars and courses in pottery and became aware of what It was like to be a student again. After years of being “the instruc to?, the experience of going through the daily routine of a student was helpful to me to understand the problems

students have In their day to day life. Also I began to understand the significance of the sense of creativity in my work as; botany teacher, in biology course development and in the creation of a functional piece of pottery such as a bowl or plate. Just as the poet may use a walk through the woods as metaphor for his/her inspiration for a poem, I used the selection of clay, shaping of the bowl, selection of the glaze and firing temperature as metaphor for the human life cycle. And so you might well ask what wifi I do retiring at the age of 59. Ohio University has an excellent early retirement plant ofwhich I am able to take advantage. I will teach my two favorite courses; the Tier-Ill course The Human Life Cycle and Botany 101, the non majors biology course one quarter each year. I enjoy teaching 101. It is an important course and must be done well. For many students this may be the only experience they will have to taste the sweetness of livingness. Where else might students be exposed to their molecular origin and begin to be aware of the impact of those unsaid eternal questions they know they have but do not elucidate. Questions such as; who am I?; what am I doing here?; is there significance in my exis tence? Basic concepts in 101 such as, evolution, genet ics, energy transfer and structure do not directly answer those questions but can put them in a Knowing context. There are many ways of Knowing and knowledge of living systems and their relationship to non-living things contribute to that Knowing. Biology has the potential to transcend the concreteness of living things to become the poetry of the soul. To recognize that in yourself and to be able to expand and develop answers to those ancient eternal questions can be achieved once the facts of biology are assimilated and we can move on to questioning the significance of our being. This can be a magnifi cent breakthrough of those walls of intellectual wisdom in which many of us have become mired. ,

In addition to teaching, my wife and I will make return visits to England and Italy and plan to visit Ireland, France and Norway. I will do more ceramics and courses in drawing and poetry. And then there are the everyday things that one always does such as having tea parties for neighborhood children, working in the garden, reading, writing letters to relatives, friends, editors, congressmen and heads of state. After a few years of this I hope to settle down and begin to think about what to do for my life’s work!!! L.A.L. AsJlrmly cemented clamshells .

Fall apart in autumn

so i must take to the road again Farewell myfrienci.s


Botany Newsletter 1989

ACTIVITIES “IN THE NEWS”: “Science comes through—Crops with less STEVE RIOCH, OU honors student and water” East Coast director of Ecology Action, and WAN K. SMITH have been featured in both local and na tional press for research on biointensive gardening. iW wire services report that Steve is growing more food with less water for a world which is becoming increasingly desert-like each year. Botany’s oneeighth acre biointensive garden, located on West State Street, has raised a weekly toll of more than 150 pounds oftomatoes, red chard and cucumbers as well as lettuce, cabbage, radishes, Swiss chard, squash and other vegetables. Steve and Ivan have received letters and certificates of appreciation from the Tn-County Community Action Agency and the Southeastern Ohio Foodbank for contributing over 6,000 pounds ofvegetables to the needy of Southeastern Ohio. The garden uses only one-fourth to one-eighth as much water per pound of food, com pared to commercial farms. It has three essential components: compost-rich, fertile soil, deep-digging which introduces air to the soil, and close spacing of plants to reduce water loss from evaporation. Steve states that there is nothing new to this proc ess, it has been used for thousands ofyears. He goes on to say that blointensive gardening (which he refers to as “sophisticated low technology”) is the answer for individuals who are interested in growing high-quality food at home, in a small space, with less water. .


towns, small but bountiful biointensive gardens could lessen the misery by providing essential nourishment for the grower’s families and cash from the sale of surplus crops in the city centers.”


The potential of biointensive gardening is re ceiving more recognition: our one week summer workshop on this method attracted 28 particIpants from 1 1 states, as distant as California, Oregon, Florida and Massachusetts, and two participants from Mexico. Steve believes that many of the principles can be adapted to commercial farms, that people will be able to grow two to six times more food using less water, half the amount of fertilizer and one hundredth the energy. Ivan says that developing countries can utilize the practice as it holds promise of “true land reform, allowing people to make a living off their land.” Jim Cavender, malnour Rioch’s advisor, emphasizes this point ishment often drives people to the city, which they surround with “squalid shantytowns” packed with refugees. In Bangul, Central African Republic, food was scarce and much of what was available was imported at great expense from Europe. Cavender states that “If people could grow food where they are and could make a little money out of It, this flow of people to the cities would stop. Even in the shanty.



“Investigating the Life Cycle of the Lowly The research of DR. JAMES Slime Mold CAVENDER was featured in the April 12, 1989, issue of The Chronicle ofH4jher Education. He says that slime molds do not deserve their repugnant name. “They probably are no slimier than any other soil microbe.” Jim has over 1 ,000 freeze-dried samples of cellular slime molds stored in his walkin refrigerator in boxes of glass tubes filled with a milk suspension. This collection contains not only Cavender’s collections but also those of his former mentor, the late Kenneth Raper of the University of Wisconsin who donated his collection in 1987. Jim has described 29 of the 60 known species, collected on his numerous travels to much of the world. They occur in humus, leaf mold and the top four inches of the forest soils and feed on the bacteria that feed on the decaying litter. He says that they are important because of their activities at the bottom of the food chain and are vital for increasing soil fertility. Thus, without slime molds, forests, especially in the tropics, cannot regenerate. Following deforestation, the increase in sunlight heats the soil and destroys cellular slime molds and most other microbes. The frequent use of slash and burn agriculture diminishes their number and Western style agricultural practices devastates them. Jim believes his samples can be revived and introduced back into tropical forest soils and may be a key in forest regeneration. Although many scientists are studying slime molds in the laboratory, Cavender is one of only two participating in field studies, including taxonomy and ecology, and has been responsible for “landmark studies” on their ecology and biogeography. He says that “When you invert the soil it creates a desert almost immediately. But it’s hard for us to see what we are doing to microbes and easier for us to see what we are doing to trees.” Thus, he does not expect the popularity of slime molds to become widespread among conservation groups. They “are not apt to rally around the slime mold.” However, his work has pointed out the importance of the soil and its organisms and their significant role in conservation. ...

An “Dormancy: How Seeds Evolved” article by GENE MAPES, GAR ROTHWELL and M. T. HAWORTH, on the “Evolution of seed dor mancy” (Nature, 337: 645-646, 16 February, 1989) .




Botany Newsletter 1989 has been featured in the Science Notebook of the Februaiy 27, Washington Post, and the April 3, The Scientist. Seeds deposited 286 million years ago in Kansas were unearthed and appear to contain the earliest evidence of the ability of seeds to remain dormant. Although this may appear obscure, seed dormancy was a major step facilitating the occu pancy of thy habitats by land plants. Prior to dormancy, seeds would germinate immediately and those disseminated to dry areas would die quickly. Thus, plants were probably limited to humid, wet regions and drier upland areas were uninhabited. The Kansas seeds contained embryos and were within cones of early conifers, a group which Gene has been studying intensively at Ohio University. Gene reports that she has really enjoyed receiving reprint requests from all over the world and “even seed companies, who will be so surprised when they see the fossils.”

AWARDS AND HONORS: Alumni Award: DR. LESUE G. HICKOK (MS 1 971) was unanimously selected by the College of Arts and Sciences Awards Committee to receive the College’s 1989 SIgnificant Achievement Award in recognition of your distinguished record In academic research and teaching Les has distinguished himself in academic teaching and research since receiving his doctorate from the University of Massachusetts in 1975. He is this country’s leading plant geneticist specializing in ferns, with a worldwide reputation for excellence. His work has had continuous grant support of over $720,000, has received awards from his colleagues, and has led to a patent for obtaining agriculturally important genes. This work continues today in a search for salt tolerant mutants. Les writes that “I have always considered my training at O.U. to be first rate and influential in regards to my career.” Les and his wife Donna returned to Athens on May 19 and 20, where they attended the Society of Alumni and Friends Program of Recognition. On Friday afternoon, he presented a seminar to the department on his recent work on the selection and characterization of single and double gene NaCl tolerant mutants in the fern Ceratopteris. Les Is currently Professor of Botany at the University of Tennessee, Knoxville. “







DR. ROBERTB. GOLDBERG (BS, 1966) has been elected an Alumni Member of Phi Beta Kappa. Bob is a plant molecular biologist and is a professor in the Department of Biology at the University of California, Los Angeles. He is on the Board of Reviewing Editors of Science and is editor of Plant Cell Biology. DR. RICHARD B. RYPMA, greenhouse cura tor, was recognized by SEPAN (Special Education Parent Advocacy Network) for contributions to the lives of handicapped people. Senator Jan Michael Long was the master of ceremonies.

Awards to undergraduate students were passed out during our annual picnic at Strouds Run State Park in late May. C. Paul and Beth K. Stocker Scholarships ($1950.00) for the 1989-90 academic year were awarded to TODD Cl-lADWELL and HEIDI SCHOCH. Todd is a sophomore and a botany major in the Honors Tutorial College. Heidi is currently ajunior In the College ofArts and Sciences. Wolfe Awards are given each year to juniors and seniors with interests and expertise in conservation. The winners of the junior and senior Wolfe Awards this year are, respectively, JACKIE and JUDITH JARDINE. Judith will be entering our graduate program this coming fall and will be spending most of this summer in Costa Rica with Dr. Jan Salick at the Organization for Tropical Studies. Jackie was inducted into the Phi Beta Kappa honors society this summer.

GRANTS: Our faculty and students continue to be supported by both national and “in-house” grants. In addition to those continuing grants, new grants were awarded from the Baker Fund of Ohio Univer sity to DRS. JAMES BRASELTON and OAR ROTI-IWELL. Jim’s grant will support his work on the karyology and systematics of the Plasmodio phoromycetes, while Gar will continue his work on biostratigraphy of early gymnosperms. DR. IRWIN UNGAR has received a grant from the Ohio Univer sity Research Committee to study the seed ecology of Spergularia marina and a Ohio University Research Challenge Award to investigate the seed physiological ecology of this species.


Botany Newsletter 1989 DR. ALLAN M. SHOWALTER and DR. MARTIN TUCK (Chemistry) have been awarded a $25,000 grant from Ohio House Bill funds to equip a mo lecular and cell biology instructional laboratory. The laboratory will be located In Room 101 of the Botanical Research Facifity at the greenhouse cornplex.

Is designed to provide expertise in each of the three levels of biological organization: cellular (CELL AND MOLECULAR BIOLOGY), organismal (SYS TEMATICS AND MORPHOLOGY), and populational (ECOLOGY AND EVOLUTIONARY BIOLOGY) To that end, our faculty have been chosen to maintath a balance in these areas, and our students are exposed to the breadth of the spectrum of plant biology. .

CELL AND MOLECULAR BIOLOGY During the past ten to 1 5 years there has been an explosion of interest, information and excitement In cell and molecular biology. Research aided by the development of a wide variety of ex panding techniques, including those for the charac terization and manipulation of genetic material, cytophotometric analysis, and molecular cloning, has led to a revolution in our concepts of genes and their expression. This arena of research is impor tant not only for the value of basic knowledge gained but also for the potentially useful crops which may be produced by genetic engineering.

Dr. Allan Showalter and JUt Zhou JACKIE HARDYADAMS (PhD. candidate) received an award of $500 from the Roger Tory Peterson Institute of Natural History to support her research on “Phlox maculata and Habemaria pera moena: convergent evolution or a case of mimicry?’. MARYLOUISE (COOKIE) TRIVFflT (Ph.D. candidate) was awarded a Wray and Todd grant of $sOo from the Paleontological Society to support her research.

PROGRAMS AND FACULTY NEWS: The success that the department has achieved during the past two decades has been due in large part to our undergraduate and graduate students as well as to the excellence of the faculty In both teaching and research. The Botany program

DR. ALLANM. SHOWALTERjoined our de partment to enhance our program in molecular biology. Allan’s research involves determining the molecular structure of genes that encode proteins found in the plant cell wall and discovering the manner in which these genes are regulated by wounding or pathogen activities. More specifically, he Is focusing on three different cell wall proteins (extensins, glycine-rich proteins, and solanaceous lectins). Currently, his lab has succeeded in Isolat ing and characterizing genes from tomato plants which encode extensins and glycine-rich proteins. During the past year he has reported on this work at the Gordon Research Conference on Plant Molecular Biology at Andover, New Hampshire, the UCLA symposium on Plant Gene Transfer at Park City,Utah, the Biotechnology Center and Department of Plant Pathology, Ohio State University, and the Department of Biological Sciences at Bowling Green State University. Contributing to his re search were DOMINIQUE RUMEAU, a postdoctoral associate who has recently returned to France, DORIS POWELL, research technician, graduate student JINZHOU, and undergraduate student PETER TANDLER. Allan has also developed a new undergraduate/graduate course entitled “Biotech nology and Genetic Engineering” which covers modem recombinant DNA techniques and their application to fields of agriculture and medicine. He


Botany Newsletter 1989 is active In the interdepartmental Molecular and Cell Biology Program and organized the annual MCB retreat in September at Burr Oak State Park. In the Botanical Research Facility, the work of DR. NORMAN S. COHN and DR. JOHN P. MiTCHEIL focuses on peas, cytophotometiy, and what Is going on Inside a developing pea stem. Specifically, they are developing cytophotometric measurements, particularly of nuclei, to characterize developmental events. Their work has resulted In two publications on the localization of nuclear protein In pea seedlings during development and the ultrastructural immunolocalization of developmental proteins in phloem cell wall regions of shoot tissues. This work was presented at the meeting of the International Society for Histochemistry and Cytochemistry in August, 1988, held in Washington, D.C. New results of a morphological! anatomical study of the effects of gibberellic acid on pea internode growth will be presented this August at the Botanical Society ofAmerica meetings in Toronto. Graduate student research involves in vitro translation analy sis of gibberellic acid-induced proteins in dwarf pea mutants with cDNA clonal analysis. Undergradu ates are participating in preparing purified monoclonal antibodies for affinity column purification of GA-induced antigenic proteins for peptide sequenc Ing. John comments that the presence of two Chinese graduate students, two Korean Honors Tutorial students, one Kenyan student visiting from Dr. Ungar’s lab and two American undergraduates aided and abetted by an American and a Scots professor has led to some very interesting, and oc casionally non-botanical, discussions. This coming fall quarter, Norm will be visiting labs in Cardiff, Aberystwyth, Bristol and Edinburgh to consult on research In shoot development and GA-induced protein changes during development. The research of DR. WAN K. SMITH focuses on the role of glutathione (GSH), and glutathione reductase (GR) in cold tolerance. Several environmental stresses, including low or high temperature, drought, and atmospheric oxidants, are thought to inhibit plant growth and development by causing oxidation of cellular components. GSH, a sulfur containing tripeptide, is an Important reductant and it has been suggested that elevation of cellular GSH and/or GR could prevent oxidative damage. Ivan has two graduate students working in his lab. TOM VIERHELLER, a doctoral student, and CAROL THORNE, in the second year of a masters program. Tom has shown that a cold-sensitive cultivar of soybean accumulates oxidized GSH when exposed to moderately cold temperatures, despite the pres “



ence of large amounts of GR. He is currently com paring the behavior of this cultivar with a coldtolerant cultivar from Switzerland. Ivan’s duties as Chair of the department have encouraged him to travel. He attended the national meeting of botany chairs at Purdue Univer sity, presented an Invited symposium paper at the southern section meeting of the American Society of Plant Physlologists in Athens, Georgia, and accom panied Tom Vierheller to an international sulfur workshop in Gronigen, The Netherlands. This summer, Tom and Ivan will be attending the ASPP meetings in Toronto, Canada. DR. JAMES P. BRASELTON is completing his studies of the Plasmodiophoromycetes. This is a group of parasitic protists which sometimes are classified in the fungi. They are significant because they are serious plant diseases (e.g. clubroot of cabbage, powdery scab of potatoes) and may serve as vectors for viral infections. Jim has received a Baker Fund Award and will use most of the funds for a collecting trip to Scotland during August. He has recently published two papers on karyotype analysis of two species in the group. Jim’s Botany 100 course (The World of Plants) has grown beyond all expectations and he now reports that he taught more students this past year than any other instructor or professor at the University.


Current patterns of human activities throughout the world in pursuit of space, food and energy indicate that within the next century most of the tropical habitats will be destroyed and along with them most of the world’s plants and animals. The expertise of biological scientists will be called upon to facilitate both the preservation and rees tablishment of these habitats. To do that ade quately, we must have an understanding of the evolutionary history of plants, their current rela tionships, and the roles that they and other organisms play or have played. The role of microbial organisms Is being investigated byDR. JAMES C. CAVENDER. During this past year he investigated cellular slime molds In Japan In collaboration with Kunihiko Kawaba of Shizuoka University, with support from a grant from the Ohio University Research Committee. He also visited the laboratory of Hiromitsu Hagiwara,


Botany Newsletter 1989 Tsukuba University, his counterpart in Japan studying the ecology and systematics of this group. After returning from Japan, Jim has been studying the effect of slash and burn agriculture on cellular slime molds in the Central African Republic In col laboration with Jaques-Paulin Regner of the Univer sity of Bangui, work supported by a grant from the National Science Foundation. Slime molds are eliminated by slash and burn agriculture but return to original levels within 1 5 years after abandonment of the fields to fallow, if the soil Is not excessively manipulated. A paper on the cellular slime molds of Japan will appear this August in the journal Mycologia.

Jim has traveled to Belize twice with stu dents to study integrative tropical botany, an cxperimental course which combines tropical rain forest botany and sustainable agriculture. There are two graduate students currently working with him, MICHAEL HOLP{ES and EDUARDO VADELL. Both of these students are studying the cellular slime molds at the Mayan ruins of Tikal, Guate mala, where there is exceptional species diversity. Michael will return to Tikal this August to do addi tional field work and collecting. Dr. Cavender has developed a Tier III course, Alternative Agriculture. The 35 students enrolled are building a sustainable 30-bed garden at the Botany Experimental Garden on the Hocking River flood plain. DR. GAR ROTHWELL is currently focusing on the origin of major groups of vascular plants, with ongoing studies on the evolution of seed plants, the early evolution of conifers, and evolu tionary radiations among ferns. His other activities include coauthorlng a second edition of Wilson Stewart’s popular text “Paleobotany and the Evolu tion of Plants.” During the past year, he presented seminars on his research at the Smithsonian Insti tution, the University of Chicago, the annual meeting of the South-Central Section of the Geological Society ofAmerica In Lawrence, Kansas, and the North-Central Section in Akron, Ohio, the Botanical Society ofAmerica meetings In Davis, California, and the Third International Organization of Paleobo tanists Conference in Melbourne, Australia. This summer, he will visit Japan and will present his work at Chiba University and Tokyo University in June, and will speak on a symposium on lycopods, conifers and ferns at the Botanical Society of Amer ica meetings In Toronto. The remainder of this summer will be spent on book writing and In field explorations for Tertiary fossil ferns in western Canada. This coming year, facets of structural botany will be brought more solidly into the curricu

lum and he will be teaching plant anatomy each fall and plant morphology each winter. He has also been running an experimental course in Botanical Photography, which will soon be a permanent course. Oar’s plant paleontology research laboratory is active with three doctoral students now concen trating on completing their doctoral dissertation research. MARYLOUISE TRIVE1Ts work on growth architecture of fossil plants includes two recent papers on primitive seed plants and two presenta tions at the upcoming meetings of the Botanical Society ofAmerica in Toronto. JAWELLE PRYOR is continuing her community ecology studies of Paleo zoic coal-forming swamps, and is awaiting publica tion of her recent study of medullosan seed ferns. LI BAT, from Nanking, China, is continuing his characterization of one of the most ancient seed plants. This coming fall, RUDOLPH SERBET from the University ofAlberta, Canada, wilijoin his laboratory as a masters student. DR. GENE K. MAPES is investigating the biology and evolutionary ecology of primitive coni fers. This past February she published some of her work on fossil conifer embryos and the origin of seed dormancy with Dr. Rothwell and undergradu ate student M. T. Haworth (Zoology) in the British journal Nature. In April, Gene and Roy Mapes (Department of Geological Sciences) presented a study on fossil plant remains from shales of oceanic sediments to the South-Central Geological Society in Arlington, Texas. Although deposited many kilometers from the paleo-shorelme and their place of growth, these plants are from a species-rich conifer community and enable geologists to recog nize time-equivalent terrestrial and marine rocks. In May, they edited a guidebook to the Regional Geology and Paleontology of the Upper Paleozoic area in southeastern Kansas, published by the Kansas Geological Survey. They have also had productive excavations at Hamilton, Kansas, includ ing three weeks of digging with 18 paleontologists, graduate and undergraduate students from several countries, experiencing the “joy of one of the hottest summers on record in July in Kansas.” DR. PHILIP D. CANTINO has been on a faculty fellowship leave this past year at the Roytal Botanic Garden, Kew, where he has been continu Ing his studies on the mint family, Labiatae. He will return to campus late in August.


Botany Newàletter 1989 ECOLOGY AND EVOLUTIONARY BIOLOGY Organisms do not occur alone in nature but interact with enumerable other organisms and with an ablotic environment. Basic to understanding life on earth Is an understanding of how organisms cope and survive, and the mechanisms which facffitate their evolution. To the end, faculty and graduate students in these areas are Investigating three aspects of ecology and evolution: responses of plants, and their evolution, to stressful environments; the physiological ecology of fern gameto phytes and the population biology of sporophytes; and the evolution of flowering plant breeding sys tems. DR. GAYLE MUENCHOWhas been studying the underlying mechanisms which support the evolution of dioecy. Last summer, she found that flower-eating weevils attack staminate and monoecious (with male and female flowers) inflores cences of the local water “arrow-leaf,” Sagittaria 1atfo1ia, at a higher frequency than they attack solely pistillate Inflorescences. This differential herbivory and the potential causes of it are currently being Investigated further by graduate stu dent BRYAN BISHOP. The overall hypothesis suggests that this differential herbivoiy may have been a driving force for the evolution of dloecy in some populations of this species. Gayle and postdoctoral associate VERONIQUE DELESALIE will travel extensively this summer throughout much of the range of the species to start the process of comparing populations that are made up entirely of monoeclous plants to populations that are predomi nantly dioecious. Genetic differentiation within and between populations of Sagtttaria are being studied by the use of starch gel electrophoresis by research associate DR. THEODORA LEE GREGG. This past year, these studies have focused on the feasibility of using isozymes as genetic markers and the perfec tion of over 10 enzyme systems. Graduate student JACKIE M. ADAMS has recently completed her doctoral comprehensives and Is working on mimicry between orchids and a local species of Phlox. FiNIEYBRYANwil1 be working on a chasmogamous/ cleistogamous breeding system. Both Adams and Bryan will be working for the Ohio Department of Natural Resources this summer, doing ecological surveys ofJackson and Vinton Counties. Last fall, Gayle presented some of her work In a symposium on dioecy and herbivory, sponsored by the Entomo logical Society ofAmerica, at Syracuse University.

Gayle has also been active in the Women in

Science and Engineering Workshop, designed to provide women students with field and career infor matlon on scientific and engineering disciplines and COLLENCHYMA, a women’s support group with particular interest in gender issues. This group meets twice monthly and, although initially founded by women from botany, now includes women from various scientific disciplines throughout the univer sity. Issues under discussion have included sexual harassment, subtle and overt bias and how to deal with it, practical issues women face in graduate training and flndlngjobs, women’s participation in nontraditional fields, and the role of women In science, the professions and the household.

Dr. Irwin Ungar with (left to right) Kern Badger, Tortya Selby, Jirn Nellesen, and Williarn Katembe

In investigating the effects of biotic and abiotic factors on plant growth and distribution along environmental gradients, DR. IRWINA. UNGAR hopes to be able to determine which factors are limiting to species at both the most stressful and the least stressful ends of physical gradients. He is studying responses of species both in the field and in the laboratory. These studies include determin ing the seed bank dynamics of the annual halophyte Spergularia marina in a salt marsh habitat near Rittman, Ohio. Spergularia is a prolific seed producer and is estimated to produce more than one million seeds per meter square in the seed bank during the spring. He has several students working in his lab. KEM BADGER has just com pleted his doctoral research on the life history


Botany Newsletter 1989 dynamics and physiological responses of seed germination in the grass Hordeumjubatum. Kern has accepted a position as Assistant Professor of Biology at Missouri Valley College. JIM NELLESEN is currently completing his dissertation on the differences in physiological and phenological re sponses of the grass Andropogon virginicus In abandoned spoils and old field habitats. JIRA KATEMBE Is studying the effects of salt stress on the seedling ernergence and DNA content of Atriplex triartgularis. TONYA SELBYhas recentlyjolned his lab and will be cornpleting her M.S. research cornparing plant population size with that of the seed bank during the growing season in a saline rnarsh area at Constitution, Ohio. This past year, DR. MARLIS RAHMAN cornpleted his doctoral work on the dernography and physiological ecology of Echino chloa crusgalli and has returned to Indonesia to his faculty position at the University of Andalas. Irwin has also been active In a university and cornrnunlty wide committee to determine the use of the Athens Mental Health grounds, which have recently corne under university ownership, and has been elected to the Executive Cornrnittee of the Ohio Biological Survey. He presented serninars on his studies on the demography and physiology of halophytes at the University of Manitoba, Winnipeg, Canada, the Ohio Academy of Sciences, Newark and the Botanical Society ofAmerica meetings in Davis, California. Although the systernatics and morphology of ferns is well understood, very little Is known about the factors which control their distribution and their physiological ecology. Efforts to understand the role of the various life-history stages, including the free-living garnetophyte, In fern distribution are underway In the laboratory of DR. ROBERT M. lLOYD. These studies Include studies by doctoral candidate ROBERT HAMILTON on an analysis of preferential habitats by two co-occurring species of Athyrium and the responses of these species to the various abiotic factors present, and studies by masters student THOMAS BRENNAN on the role of the gametophyte in the distribution of calciphilic ferhs. Bob Hamilton will be completing his studies this corning year. They have included docurnenta tion of the presence of a spore bank, antheridiogen effects In gametophyte populations, gametophyte distribution and gender in field populations, se quence of frond developrnent during the growing season and photosynthetic measurements and response to various light levels. He has also studied genetic differentiation within his populations using isoenzymes and hopes to finish with restriction analysis of DNA of his plants this coming summer and fall. Bob presented various aspects of his work

at the Botanical Society of America meetings in Davis last summer and again this summer at the meetings in Toronto. Tom’s work will try to dis criminate why certain species ofAsplenium are mostly restricted to limestone habitats while other species are not found there. He will culture gameto phytes of a variety of calciphilic and non-calciphiic species on various concentrations of calcium at various pH levels and determine their responses. The work will be supplemented with field studies to determine more exactly the calcium concentration and pH of the soils sporophytes occur in. Dr. Lloyd continues to develop a volume on the evolutionary biology of ferns and has completed two taxonomic treatments of Acrostichum and Ceratopteris for the Flora North America project, organized by the Missouri Botanical Garden, St. Louis.

RECENT FACULTY AND STUDENT PUBLICATIONS: Braselton, J.P. 1989. Karyotypic analysis of Ligniera verrucosa (Plasmodiophoromycetes). Can. J. Bot. 67: 1216-1218. Braselton, J. P. 1989. Karyotypic analysis of Membranosorus heterarttherae (Plasrnodio phoromycetes). Can. J. Bot. 67: 12 191220. Cohn, N. S. J. P. Mitchell and Z. Zhou. 1989. Ultrastructural immunolocalization of developmental proteins in phloem cell wall regions of Pisum sativum L. shoot tissue. Plant Science 60: 137-143. ,

Feng, B. C. and G. W. Rothwell. 1988. Microspo rangiate cones of Mazocarpon bensonii from the Upper Pennsylvanian of the Appalachian Basin. Review of Paleobotany and Palynol ogy57: 289-297. Good, C. W. and G.W. Rothwell. 1988. A reinterpretation of the Paleozoic fern Norwoodiaartgustum. Review of Palaeobotany and Palynology 56: 199-204. Hamer, J. J. and G. W. Rothwell. 1988. The vegetative structure of Medullosa enclocentrica Baxter (Pterldospermopsida). Can. J. Bot. 66: 375-387.


Botany Newsletter 1989 Hamilton, Robert G. 1988. The significance of spore banks in natural populations of Athyrium pycrtocarport and A. thelypterioides. Amer. Fern J. 78: 96-104. Kauffman, G., J. C. Cavender and H. R Hohi. 1988. Polysphortdylium luridum, a new dictyostelid species with unique spores. Botanica Helvetica 98: 123-13 1.

Larson, L. A. 1988. Curiosity about biology Kendall-Hunt. Lloyd, R. M. 1988. Experimental studies on the probability of selfing by protandrous gametophytes. Amer. Fern J. 78: 1 1 7- 121. Mapes, G. K. and R. H. Mapes (Eds.). 1988. Regional geology arid paleontology of upper Paleozoic Hamilton quamj area in southeastem Kansas. Kansas Geol. Survey Guidebook Series 6. Mapes, G. K. and G. W. Rothwell. 1988. Diversity among Hamilton conifers. j G. Mapes and R. Mapes, thici, pp. 225-244. Mapes, G. K., G. A. Leisman and W. Gillespie. 1988. Plant megafossils from the Hartford Limestone (Virgiian-Upper Pennsylvanian) near Hamilton, Kansas. in G. Mapes and R. Mapes, thid. Mapes, G. K., G. W. Rothwell and M. T. Haworth. 1989. EvolutIon of seed dormancy. Nature 337: 645-646. Mitchell, J. P. N. S. Cohn and Z. Zhou. 1988. Localization of a nuclear protein during development in the pea seedling. Plant Science 58: 253-260. ,

Muenchow,G.E.andM.Grebus. 1989. The evolution of dloecy from distyly: reevaluation of the hypothesis of the loss of long-tongued pollinators. Amer. Nat. 133: 149-156. Rothwell, G. W. 1988. Upper Pennsylvanian Steubenvile coalball flora. Ohio J. Sd. 88: 61-64. Rothwell, G. W. 1988. Cordaltales, C. B. Beck (Ed.) Origin and evolution ofgymrtosperms. Columbia Univ. Press. pp. 273-297.

Rothwell, G. W. and G. K. Mapes. 1988. Vegetation of a Paleozoic conifer community. j G. Mapes and R. H. Mapes (Eds.) Regional geology and paleontology ofthe Upper Paleo zoic Hamilton quarry area in southeastern Kansas. Kansas Geol. Survey Guidebook Series6: 213-224. Rothwell, G. W. and S. E. Seheckler. 1988. Biology of ancestral gyrmnosperms fl C. B. Beck (Ed.), Origin and evolution ofgymnosperms. Columbia Univ. Press, pp. 85-134. Rothwell, 0. W. and Ruth A. Stockey. 1989. Fossil Ophioglossales in the Paleocene of western North America. Amer. J. Bot. 78: 637-644. Showalter, A. M. and J. E. Varner. 1989. Plant hydroxyproline-rich glycoproteins. P. K. Stumpf and E. E. Conn (Eds.), The biochem Lsty ofplants, Vol. 15, molecular biology. Academic Press, New York, pp. 485-520. Showalter, A. M., D. Rumeau, J. Zhou, S. G. Worst and J. E. Varner. 1989. Tomato hydroxyproline-rich glycoprotein genes: structure and expression in response to wounding. J. Cell Biochem. 13D: 326. Smith, I. K. and A. L. Lang. 1988. Translocation of sulfate in soybean. Plant Physiol. 86: 798-802. Smith, I. K., T. L. Vierheller and C. A. Thorne. 1988. Assay of glutathione reductase in crude tissue homogenates using DTNB. Anal. Biochem. 175: 408-417. Trivett, Mary L. and G. W. Rothwell. 1988. Diversity among Paleozoic Cordaitales: The vascular architecture of Mesoxylort birwne Baxter. Bot. Gaz. 149: 116-125. Trivett, Mary L. and G. W. Rothwell. 1988. Modeling the growth architecture of fossil plants: A Paleozoic Filicalean fern. Evolutionary Trends in Plants, 2: 25-29.

Ungar, I. A. 1988. Effects of the parental environ ment on the temperature requirements and salinity tolerance of Spergularia marina seeds. Bot. Gaz. 149: 432-436.


Botany Newsletter 1989 Ungar, LA. 1988. A significant seed bank for Spergularta marina (Caryophyllaceae). Ohio J. Sd. 88: 200-202.

Wame, T. R. and R M. Lloyd. 1987. Expression of a clumped-chloroplast mutant In the fern Acrostichum danaelfolium. Bot. Gaz. 148: 120-122. Warne, T. R and R. M. Lloyd. 1987. Gametophytic density and sex expression in Ceratopteris. Can. J. Bot. 65: 362-365.

STUDENTS COMPLETING GRADUATE DEGREES: M. S. : Bryan Hoffman Ph.D. : Barbara Ballard, Kemuel Badger, Marlis Rahman, Zeqi Thou

NOTES ON FACILITIES: This past year a major Impetus has been given by Dr. Smith to evaluating and improving the departmental greenhouse and botanical garden complex. Under the direction of Jim Braselton, plans have been made to remodel the existing garden, next to the Botanical Research Facility and greenhouse complex by improving floral displays, plantings and labeling and walkways. A portion of the south end of the greenhouse will also be remod eled to provide displays of tropical plants and eth nobotanically useful plants. These Improvements are intended to serve not only the teaching and research needs of the department but also to provide a pleasant educational atmosphere for the public and the university. The departmental garden cqmmittee, headed by Dr. Braselton, has been joined this year by JEANANDREWS. Jean has recently moved to Athens from Berkeley, California, where she served as Assistant Museum Scientist and Curatorial Assistant at the University of Califor nia Botanical Garden. While at Berkeley, she initiated a flowering phenology program, computerized their plant records, produced the biennial seed exchange catalog and co-authored three successful grant proposals. She has also worked at the Mildred E. Mathias Botanical Garden at UCLA.

Jean has provided a number of new ideas to the committee and has attended a number of conferences and regional botanical gardens on behalf of the department. She has also revived the Athens chapter of the Ohio Native Plant Society and most recently submitted an extensive proposal recom mending the establishment of an Ohio University Ecological Study Area and Botanical Garden at ‘The Ridges,” land recently acquired by the university from the Athens Mental Health Facility. ALDEN UBRARY of Ohio University was the recipient of a gift of the first of two volumes of Linnaeus’ Genera plantarum, edition 8, attributed to Johann Christian Daniel Schreber, and published in Frankfort, Germany, in April 1789. The donor wasMr. H. C. Torrey of Highland Park, New Jersey, a descendent of Manasseh Cutler. The volume bears Cutler’s signature.


William (Bil) Alverson, University of Wisconsin K. Arumuganathen, Cornell University Jane H. Bock, University of Colorado John L. Caruso, University of Cincinnati Peter S. Curtis, Ohio State University Peter R. Crane, Field Museum, Chicago William Crepet, University of Connecticut Michael Ellis, Ohio State University, Wooster Ray Evert, University of Wisconsin Keith Garbutt, University of West Virginia Leslie G. Hickok, University of Tennessee Llewellya Hillis, Ohio State University Rafael Pont Lezica, Washington University, St. Louis Eileen A. Maher, University of Wisconsin James McCarthy, Harvard University David J. McLaughlin, University of Minnesota John H. Miller, Syracuse University Natasha Raikel, Michigan State University John Rotenberly, Bowling Green State University Jan Salick, New York Botanical Garden Eugene W. Schupp, Estacion Biologica de Donana, Sevilla, Spain Mark Shotwell, Purdue University Allison Snow, Ohio State University Ruth A. Stockey, University of Alberta Desh Pal Verma, Ohio State University



Botany Newsletter 1989

DEPARTMENT OF BOTANY PERSONNEL: FACULVñ Arthur Blickle, Associate Professor Emeritus James E. Braselton, Professor Phifip D. Cantino, Associate Professor James C. Cavender, Professor Norman S. Cohn, Distinguished Professor J. Herbert Grafflus, Associate Professor Laurence A. Larson, Professor Robert M. Lloyd, Professor Gene K. Mapes, Adjunct Assistant Professor John P. Mitchell, Professor Gayle E. Muenchow, Assistant Professor Gar W. Rothwell, Professor Allan M. Showalter, Assistant Professor Ivan K. Smith, Professor and Chair Invin A. Ungar, Professor Monroe T. Vermilion, Associate Professor Emeritus Warren A. Wistendahi, Professor Emeritus POSTDOCIDRALS AND RESEARCH ASSOCIATES: Veronique Delesalle, Theodora Lee Gregg, Doris Powell, Dominique Rumeau STAFF Judith Dowler, Department Secretary Brenda S. Ingraham, Technical Typist Carolyn H. Keiffer, Technical Assistant Elizabeth D. Moore, Technical Assistant Richard Rypma, Greenhouse Curator GRADUATE STUDENTS: Doctoral Students Mones Abu-Asab, Jackie M. Hardy Adams, Kemuel S. Badger, Finley A. Bryan, Jon J. Hamer, Robert G. Hamilton, Chenzhao Jian, LI Bal, James E. Nellessen, Janelle S. Pryor, Mary L. Trivett, Thomas L. Vierheller, Steven J. Wagstaff, Liulai Wu, Jin Zhou.

Master’s Students D. Bryan Bishop, Thomas H. Brennan, Michael T. Holmes, William Jira Katembe, Nandini Ramachan dra, Tonya Selby, Carol A. Thorne, Eduardo M. Vadell.

NEWS OF ALUMNI DIANA WOZNIAK BALDI (BS, 1977). Diana is a chemist In the Quality Assurance Section in Region III of the Environmental Protection Agency. She writes that “My nine years with EPA have been constantly evolving and exciting. Now, I manage nearly 20 contractor staff for superfund. The challenges of toxic waste cleanup are continually expanding.” Diana is currently living on S. Victor Parkway In Annapolis, Maryland.

BARBARA BALLARD (Ph.D., 1988) has had a paper accepted by the Can. J. Bot. on the phenology of mycorrhizal infection in Hordeumjubatum. She has two children, Kitie, 5, and John, 8 months.

DAVID BARNES (BS/BSEd, 1979) is a Senior Engineering Representative in Denver, Colorado, working as a loss control representative for Aetna Insurance Company. He writes that “The job basically involves hazard recognition, analysis, suggesting possible alternative solutions and monttoring Implementation. Over the last three years this has involved asbestos and pollution liabilities and asbestos removal. He has also been involved in air sampling and noise testing for various compa nies to determine workman compensation expo sures and has been educating the workforce with personalized slide shows. Aetna is paying for part of his additional education in loss control management and data processing, programs he is about to complete. Following this, he will pursue programs for becoming an Associate in Risk Management and/or a Certified Safety Professional. “In particu lar, the latter will involve recalling and studying technical material—much of which involves former o.U. study.” Dave says that there is always another step to be taken, even beyond the intellect. He has done quite a bit of enthusiastic backpacking in the Colorado Rockies and New Zealand the past two years. This year he had a fantastic garden using the Square Foot Gardening approach. Burpee has some new corn hybrids out that are both disease resistant (vertidillium) and extra sweet (taking longer for the sugar/starch conversion after har vest). “It was so sweet, no butter or salt was needed!” “


Botany Newsletter 1989 CHRIS CLINGMAN (BS, Field Biology, 1985). Chris is an Interpretive Naturalist for St. Joseph County Parks in South Bend, Indiana. Since receiving his degree he has worked In various aspects of the environmental education field. His first position was as a seasonal naturalist at Brown County State Park in Indiana. He then worked at Woodland Altars Outdoor Education Center, Peebles, Ohio, as a senior naturalist. He then moved to northern Indiana to work at Potato Creek State Park as a naturalist and as a multi-image slide show producer. He writes that “I just started with St. Joseph Thisjob gets me Into everything County Parks from making sorghum syrup to maple syrup to spinning/weaving to voyageur living history programs (we have a 34 ft. voyageur canoe), to the building of frontier settlement, an environmental education center and now possibly a residential outdoor education center.” He occasionally leads school groups on hikes, and “these school groups, whether they be environmental or historical, are our main purpose. But, one thing is for sure. It Is too flat up here! No hills to be found. It drives you crazy after a while.” .


indulged in her passions for traveling, gardening, music and theater, including several trips to Europe with visits to all the musical and theatrical greats in the major European capitals as well as New York City. She currently owns a 75 year old duplex and is renovating the house and gardens (1 1, so far). Because of the renovation, she has become seri ously involved in the preservation and use of herbs and flowers. As she writes, her life, thus far, has been interesting, at times productive and rarely ever calm, sedate or boring!


DONNA G. COUCH (MS. 1971) is a Professional Sales Representative for Winthrop Pharmaceuti cals/Sterling Drug and is living in Cleveland Hts., Ohio. After moving to Cleveland, she was employed by the Cleveland Board of Education as a math and science instructor at the Adult Education Center from 1972 until 1978. While there she wrote a diagnostic math development curriculum for adults and for four years she was chair of the Science Department. During this period she developed a quarterly minicurricula In biology for adults and taught chemistry and physics to adult college bound students. Upon leaving the AEC, she took a position as a youth jobs coordinator and adrninis trator until 1980 when shejolned the pharmaceuti cal company. She writes that “This position provided me an entire new area of knowledge and expertise. During my initial pharmaceutical sales representative stage, I was a novelty in the medical community. Womenjust weren’t outside sales representatives! Over the years I have been involved with sales training, representing Winthrop at national conventions, marketing research and trials, and of course Winthrop’s product training and promotion to physicians, interns and residents as well as health support personnel.” She has

LESLIE G. DYBIEC (BS, 1987) will be completing her M.S. degree in horticulture this August at Ohio State University, Columbus.

PAUL GOLDSTEIN (MS. 1975) is an Associate Professor of Genetics in the Department of Biologi cal Sciences, University of Texas at El Paso. He writes that he has published over 50 papers on the ultrastructure of melotic chromosomes and recently received two copyrights for 3-D graphics computer programs and has applied for a patent on immu noassay development.

DAVID HALL (MS. 1982) writes that “teaching high school isn’t as bad as the media portrays it, but it is demanding. We have finished another year and I am still recovering. On the news front, Lori and I now have two kids, a girl (now almost 5 years old) and a boy (three years old). We bought a house and have two cats. Ifwe only had a dog and another 0.3 children we’d be a statistician’s dream of the ideal family. I have been very active in state biology organizations and have been nominated for miscel laneous awards, so I guess some folks think I know what I am doing. Sure have them fooled!”

H. MICHAEL HARRINGTON (Ph.D. 1978) is a faculty member in the Department of Plant Molecu lar Physiology at the University of HawaII, Honolulu. He was recently a co-organizer of a workshop entitled “Calcium Research on Plants: A Workshop on Methods”, held January 8-1 1, 1989, at the Kona Surf Resort, Kona, HawaII. This workshop evalu-. ated methodologies commonly used to investigate calcium-mediated systems, responses and proc esses in plants. ,


Botany Newsletter 1989 CECILE HENAULT (BS, 1982) is Senior Representa tive for Merck Sharp and Dohme Pharmaceutical Company in Larsen, Wisconsin. She specializes In musculoskeletal diseases and in cholesterol inhibi tion. She spent September 1987- 1988 retraining nursing staff and physicians on cell-mediated encocytosis and lipoprotein structures. She writes that “It has been a challenge and exhilarating as cholesterol is the hot topic of the 80’s.”

LESLIE G. HICKOK (MS. 1971). Les Is Professor of Botany and Director of the Plant Physiology and Genetics Graduate Program at the University of Tennessee, Knoxville. He has recently received a $172,000 grant from the National Science Founda tion to characterize salt tolerant mutants in Cera topteris. He has recently been elected into membership of Science Alliance, a Centers of Excellence Program for the State of Tennessee and returned to Athens this past May to receive an Alumni Achievement Award from the College of Arts and Sciences. CHRIS HOPKA (MS. 1972) has started his own business, DRUMLIN Nurseries, Ltd., In Deerfield, Wisconsin, where he specializes in his own Cape Primrose (Streptocarpus) hybrids. In the DRUMLIN collection are also xerophytes, cymbidium orchids and geraniums.

GARY KAUFFMAN (MS. 1986) published a paper with Jim Cavender and H. R. Hol on a new species, Polysphortdylium luridum. He is teaching plant taxonomy at a small college in North Carolina and will be conducting a horticultural tour of Costa Rica this summer.

DAVID G. LOVELAND (MS. 1981) has resigned from his position as Environmental Advisor to the League of Women Voters and has taken a new position as Executive Director of the National Recycling Coali tion. He recently appeared on the Cable News Network (CNN) discussing the environmental impact of disposable diapers.

SCOTF MARIS (BS, 1982) is an Environmental Manager at Chemical Waste Management, Inc. in Vickery, Ohio. His job is to keep the site in cornplete compliance with all the hazardous waste rules and regulations. Scott is living on a farm with his wife and first child, Emily, who he describes as “the best thing ever.”

BRIAN McCARTHY (MS. 1984) is now employed in a tenure-track position as a forest ecologist at Frostburg State University. He writes that “I am quite excited about the position for a variety of reasons. Firstly, it is a special research appointment with a significantly reduced teaching load (never more than 6 contact hours per semester). Secondly, they have a relatively new graduate program to provide for various specializations in Biology, Resource Management and Conservation Biology. So there is the potential for graduate students. Thirdly, and perhaps most exciting, is that this position holds with it a joint appointment to the newly created International Institute of Renewable Resources (to be based at FSU). I am encouraged and partially subsidized to engage in as much basic and applied tropical research as I can handle or want. I have already set in motion the steps to go to Costa Rica, Argentina and Malawi within the upcoming 18 months. In addition, I hope to teach a 3-week undergraduate field course in Tropical Ecology in Costa Rica. Unfortunately, this position will require some substantial understanding on the part of my wife who has now moved fairly high up in the NJ DEP. She will be teaching anatomy and physiology at Allegheny Community College in Cumberland.”

JAMES E. MICKLE (Ph.D., 1983) is on the botany faculty at North Carolina State University and has received a Guggenheim Fellowship for the summer of 1989 to work with Dr. Manford Barthel (East Berlin) on Pennsylvanian age tree ferns.

STEVEN H. MILLER (BS, Environmental Biology/ Botany, 1977) is living in Solon, Ohio. For the past 12 years he has been working in fire protection and safety engineering. For the past five years he has been with Chubb & Son, Inc. Cleveland Branch office. The position involves evaluating automatic sprinkling systems and other fire protection sys tems; evaluating environmental and industrial hygiene exposures; evaluating product and public ,


Botany Newsletter 1989 to risk management and safety of the large indus trial concerns insured by Chubb. Steven has attained the Certified Safety Professional designa tion recognized by the Board of Certified Safety Professionals of the Americas.

VIRGINIA S. MORAN (BS, 1983; MS. 1986) is a botanist for Sweetwater Environmental Biologists, an environmental consulting company that pre pares biological surveys for environmental impact statements or reports. From January to May, 1987, she worked as a research technician for Dr. Paul Zedler of San Diego State University, on vernal pools, monitoring “a mint of all things (Pogogyrte abrortsiO for demography, pollination biology and phenology. She also traveled to Alaska where she worked as a research technician at the Toolik Lake Arctic Research Station on the edge of the Brooks Range, 1 50 miles south of Prudhoe Bay. She is also the chairperson of the San Diego Chapter of the California Native Plant Society and is writing music for a local nature program for children (Mr. B’s World). She is active in the local environmental movement and “learning my Phaeophyta! here in


EDWIN PERKINS (MS. 1973) is an organic farmer living in New Marshfleld, Ohio. He writes, “We live on a small farm in Athens County. My wife, Amy, is a teacher, writer and church organist. Our kids are Willie, 12, and Rebecca, 10. They both take violin lessons at O.U. Music School. We have started an organic farm raising vegetables and fruits.” Their products are available at the Farmer’s Market in Athens.

KATHLEEN B. PIGG (BS, MS. 1983) Is a new Assis tant Professor of Botany at Arizona State University, Tempe, and is presently on a NATO Postdoctoral appointment at the University ofAlberta, Canada.

BOYD W. POST (BS, 1950) is a Forest Biologist for the Cooperative State Research Service, U.S. De partment ofAgriculture, Washington, D.C. He writes that “The kids are all grown up and the dog died last summer—life has begun! Three years ago we added a greenhouse to the south side of our home and are enjoying collecting fragrant flowering plants. After nineteen years with USDA I still enjoy

what I do very much. The Mclntire-Stennis Cooperative Forestry Research Program occupies most of my time, but I also have responsibilities In agricul tural meteorology/climatology and wildlife/fisheries research. During the coming year I will lead 7 or 8 comprehensive reviews of research instruction, and extension activities in the cooperative state forestry schools and land-grant institutions. I would appre date suggestions of fragrant plants for home greenhouses.” Boyd’s address is 1 107 Pekay St., Vienna, VA 22180.

MARLIS RAHMAN (Ph.D. 1988) completed his degree and has resumed his academic appointment at Universitas Andalas, Indonesia. ,

RUTh A. STOCKEY (MS. 1984) has been promoted to full professor in the Department of Botany, University ofAlberta. She is currently a Visiting Scholar in the Laboratory of Phylogenetic Botany at Chiba University, Chiba, Japan.

FRANK W. TELEWSKI (MS. 1980) is an Assistant Professor in the Laboratory of Tree Ring Research, at the University of Arizona, Tucson. This past year he traveled to Australia’s Great Barrier Reef on a NSF sponsored trip to talk to coral researchers about Arizona’s use of tree rings to study past climates. While there he participated in a planningtraining workshop following the International Coral Reef Symposium in Queensland.

SALLY LEAR TIRRELL (MS. 1969) is a high school science teacher (chemistry and physiology) in Norman, Oklahoma. She writes that she was one of Dr. Ungar% first graduate students over 20 years ago. Since receiving her degree she has been involved in a variety of interesting careers, an interpretive naturalist for the National Audobon Society, consulting for Nature Centers, medical technologist doing medical research as well as teaching. She would love to hear from Jerry Sledge, Mark McClelland, Dave Enterline, Dennis Gavoni, to catch up on the last 19 years. Sally’s current address is 1630 Beverly Hills, Norman, Oklahoma 73072.


Botany Newsletter 1989 SUSAN HARRISON WYLIE (BS, 1977) is a housewife and volunteer for the NaUonal Ski Patrol and lives in Clarkson, Michigan. Following graduation, in the Cleveland area, she worked in marketing, then sales, first for a medical equipment company and then for a pharmaceutical company, and then went into market research. She moved to Clarkson in 1987 with her husband. They have a one year old daughter and Susan is applying her botanical training to gardening.

MARK K. WOURMS has written “a quick update of past and a brief look into the future: I have been giving quite a few talks lately on zoo horticulture. As a member of the Assoc. of Zoological Horticultu ralists, I gave two papers In our annual meeting in Tucson. One was on a “Living Wall” we created at the Central Park Zoo, the other was on using hotanical resources at zoos to teach ecology. They were well received. The highlight of the conference were several trips into the Sonoran Desert, includ ing 3 days near Hermosillo, Mexico. I have been teaching a Zoo Horticulture class at the New York Botanical Garden. I find myself explaining philoso phy as much as horticulture. It is difficult to keep people from falling into the Shopping Mall syndrome

of interior/exterior landscaping. I was part of a symposium on Interior Landscaping in New York. I pushed the 95 participants to push the growers and industry in general to look for more diversity in plants. Additionally, I mentioned that I am worried that plant breeding for ornamentals may lead away from enhancement and too far into architectural form and function. I think some of the hybrid bromeliads for example are now so colorful, perfect, that they no longer seem real, the nature has been lost. They are now as architectural as a marble column. One reason I have little prep time is that I am going to spend three weeks in December in Thailand. I will be traveling with an ecologist friend from Harvard. We will bird, botanize and just see the country.

NECROLOGY: JOHN WARD MILLER (BS, Agriculture, 1950) on August 9, 1987, Dublin, Ohio. John was retired from the International Harvester Co. DON A. FREDERICK (BS, 1953), Marietta, Ohio, on December 7, 1987.

Botany Faculty



(Left to rqht) Phillip Cantino, Norman Cohn, Robert Lloyd, Gar Rothwell, Irwin Ungar, Gayle Muenchow, Warren Wistenclahi, James Cande r, Lairy Larson, James Braseltoii John Mitchell, Herbert Graffiu s, Ivan Smith (Missing: Allan Showalter)

1989 PBIO Newsletter