obligada (parasitoides) o de forma facultativa (hormigas y otros depredadores) (Fig. 10). Tratar de desvelar este sistema complejo de interacciones es de vital importancia para poder conocer en profundidad este maravilloso micromundo. REFERENCIAS Axen, A.H. & Pierce, N.E., 1998. Aggregation as a cost-reducing strategy for lycaenid larvae. Behavioral Ecology, 9 (2): 109-115. Álvarez, M., 2009. Estudio de la comunicación acústica en licénidos (Lepidoptera, Lycaenidae) y formícidos (Hymenoptera, Formicidae) Tesis Doctoral. Departamento de Zoología y Antropología Física (Zoología), Facultad de Ciencias Biológicas, Universidad Complutense de Madrid. Begon, M., Townsend, C.R. & Harper, J.L., 2006. Ecology. From individuals to ecosystems. Fourth edition. Blackwell Publishing Ltd. Pp. 759. Both, C., Melo A.S., Zanini, S. & Hartz, S., 2011. Tadpole co-ocurrence in ponds: when do guilds and time matter? Acta Oecologica 37, 140-145. Chapman, J.W.; Williams, T.; Martínez, A.M.; Cisneros, J.; Caballero, P.; Cave, R.D. & Goulson, D., 2000. Does cannibalism in Spodoptera frugiperda (Lepidoptera, Noctuidae) reduce the risk of predation? Behavioral Ecology and Sociobiology, 48: 321-327. Church, S.C. & Sherratt, T.N., 1996. The selective advantages of cannibalism in a Neotropical mosquito. Behavioral Ecology and Sociobiology, 39: 117-123. Dempster J.P., 1997. The role of larval food resources and adult movement in the population dynamics of the orange-tip butterfly (Anthocharis cardamines) Oecologia, 111: 549-556. Dong, Q. & Polis, G.A., 1992. The dynamics of cannibalistic populations: a foraging perspective. In: Elgar, M.A. & Crespi B.J. (eds.) Cannibalism – ecology and evolution among diverse taxa. Oxford Science. Oxford 13-37. Ehrlich, P.R. & Raven, P.H., 1964. Butterflies and Plants: A study in coevolution. Evolution, 18 (4): 586-608. Fernández-Haeger, J. & Jordano, D., 2009. La mariposa monarca Danaus plexippus(L., 1758) en el Estrecho de Gibraltar (Lepidoptera: Danaidae). SHILAP Revta. lepid., 37 (148): 421-438 Fiedler, K., 1991. Systhematic, evolutionary and ecological implications of myrmecophily within the Lycaenidae (Insecta: Lepidoptera: Papilionoidea). Bonn Zoological Monograph, 31: 1-20. Fiedler, K., 1996. Host-plant relationships of ly-
caenid butterflies: Large-scale patterns, interactions with plant chemistry and mutualism with ants. Entomologia Experimentalis et Applicata, 80 (1): 259-267. Fiedler, K., Krug, E., & Proksch, P., 1993. Complete elimination of hostplant quinolizidine alkaloids by larvae of a polyphagous lycaenid butterfly, Callophrys rubi. Oecologia, 94 (3): 441-445. Gil-T., F., 2004. Nuevos datos sobre la biología de Iolana iolas Ochsenheimer (Lepidoptera, Lycaenidae) y su interacción con himenópteros mirmecófilos, fitófagos y parasitoides (Hymenoptera, Formicidae, Eurytomidae, Ichneumonoidea). Boletín de la Sociedad Entomológica Aragonesa, 34: 139-145. Hardin, G., 1968., The tragedy of the commons. Science, 162: 1243-1248. Holzinger, F., Frick, C. & Wink, M., 1992. Molecular basis for the insentivity of the Monarch (Danaus plexippus) to cardiac glycosides. Federation of the Societies of Biochemistry and Molecular Biology, 314 (3): 477-480. Hutchinson, G.E., 1959. Homage to Santa Rosalia, or why are there so many kinds of animals?. American Naturalist, 93: 145-59. Jordano, D., Rodríguez, J., Thomas, C.D. & Fernández-Haeger, J., 1992 The distribution and density of a lycaenid butterfly in relation to Lasius ants. Oecologia, 91 (3): 439-446. Kakimoto, T., Fujisaki, K., & Miyatake, T., 2003. Egg laying preference, larval dispersion and cannibalism in Helicoverpa armigera (Lepidoptera: Noctuidae). Annals of the Entomological Society of America, 96 (6): 793-798. Kinoshita, M., 1998. Effects of time-dependent intraspecific competition on offspring survival in the butterfly, Anthocharis scolymus (L.) (Lepidoptera: Pieridae). Oecologia, 114: 31-36. Leon-Beck, M. & Coll, M., 2007. Plant and prey consumption cause a similar reductions in cannibalism by an omnivorous bug. Journal of Insect Behavior, 20 (1): 67-76. Mattson, W.J., 1980. Herbivory in relation to plant nitrogen content. Annual Review of Ecology, 11: 119-161. MacArthur, R.H. & Levins, R., 1967. The limiting similarity, convergence, and divergence of coexisting species. American Naturalist, 101: 377-85. Muñoz-Sariot, M.G., 2011. Biología y ecología de los licénidos españoles (Lepidoptera, Lycaenidae). 384 pp. Obregón, R., 2011. Interacciones en un gremio de mariposas (Lepidoptera: Lycaenidae) asociadas 53